Heterometrus Glaucus
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''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
southeastern Asia, including Indonesia, Brunei, Malaysia,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, Philippines, Singapore, Cambodia,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, Thailand, Vietnam, India (
Nicobar Islands The Nicobar Islands are an archipelagic island chain in the eastern Indian Ocean. They are located in Southeast Asia, northwest of Aceh on Sumatra, and separated from Thailand to the east by the Andaman Sea. Located southeast of the Indian s ...
,
Andaman Islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
), and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
( Hainan). It is notable for containing some of the largest living species of scorpions.


Taxonomy

The genus was introduced by
C.G. Ehrenberg Christian Gottfried Ehrenberg (19 April 1795 – 27 June 1876) was a German Natural history, naturalist, zoologist, comparative anatomist, geologist, and microscopy, microscopist. Ehrenberg was an Evangelicalism, evangelist and was considered to ...
(in Hemprich & Ehrenberg, 1828), originally as a
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of the genus ''
Buthus ''Buthus'' is a genus of scorpion belonging and being eponymous to the family Buthidae. It is distributed widely across northern Africa, including Morocco, Mauritania, Algeria, Tunisia, Libya, Egypt, Senegal, Guinea-Bissau, Nigeria, Sudan, Somali ...
''. It was elevated to genus rank by F. Karsch in 1879. H.W.C. Couzijn (1978, 1981) subdivided the genus into several
subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
, but F. Kovařík (2004)
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
ized these subgenera with the nominal genus. In 2020 the genus was reviewed by L. Prendini & S. F. Loria, three of the former subgenera were revalidated and elevated to genera and one valid subgenus was elevated to genus rank, species were transferred to appropriate genera, resulting in 28 new combinations.


Diversity

The content of this genus may vary, depending on the authority. Eight species are known, many of which are quite similar in appearance: *'' Heterometrus glaucus'' (Thorell, 1876) *''
Heterometrus laevigatus ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Thorell, 1876) *''
Heterometrus laoticus ''Heterometrus laoticus'' or Vietnam forest scorpion, is a scorpion species found in peaty areas of Vietnam and Laos. They can reach lengths of . They are a communal species, but cannibalism has been known to occur, and if caught, they can be ex ...
'' Couzijn, 1981 *'' Heterometrus longimanus'' (Herbst, 1800) *''
Heterometrus petersii ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Thorell, 1876) *''
Heterometrus silenus ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Simon, 1884) *''
Heterometrus spinifer ''Heterometrus spinifer'', the Asian forest scorpion, giant blue scorpion, or giant forest scorpion, is a species of scorpions belonging to the family Scorpionidae. Description ''H. spinifer'' can reach a length around . The body is shiny blac ...
'' (Ehrenberg, 1828) *''
Heterometrus thorellii ''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, ...
'' (Pocock, 1897)


General characteristics

Members of ''Heterometrus'' are generally large-sized scorpions (100–200 mm or about 4-8  in total length). Coloration is dark in most species, often uniformly brown or black, sometimes with a greenish shine, with brighter-colored telson, walking legs, and/or pedipalp pincers in some species. The scorpions are heavily built with especially powerful and globose pedipalp pionkes, broad mesosomal tergites and a proportionally slender and thin metasoma. The telson is proportionally small and the stinger is often shorter than the vesicle. The
cephalothorax The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
and mesosoma are largely devoid of carinae and granulation and the median eyes are situated in a small, lenticular depression on the cephalothorax. Some species are parthenogenic. Orthobothriotaxy type C. Pedipalp femur with three trichobothria and pedipalp patella consists with 19 trichobothria. Pedipalp chela with 26 trichobothria. Retrolateral pedal spurs are absent. Stridulatory organ is located on the opposing surfaces of pedipalp coxa and first leg.


Toxicity

As in other genera of the Scorpionidae, the symptoms from ''Heterometrus'' envenomations are rather mild and no human fatalities are known. The sting causes local pain, inflammation, oedema, swelling, and redness of the skin, lasting for hours to a few days. Plant extracts known in the traditional Thai medicine as natural scorpion venom antidotes are effective as symptomatic treatment of ''H. laoticus'' stings. The protein heteroscorpine-1 was found the major component of the venom in ''H. laoticus''.


Habitat for the scorpion

Species of ''Heterometrus'' live in vegetated, often forested,
humid Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation, dew, or fog to be present. Humidity depen ...
regions with
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
to tropical climates. As most scorpions, they are predominantly
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
and hide in
burrow An Eastern chipmunk at the entrance of its burrow A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s, below logs, and in leaf litter.


In captivity

Due to their impressive size, low toxicity, and docile behavior, species of ''Heterometrus'' are popular pet scorpions. Unlike many other scorpions, they can be kept in pairs or small groups.


References


External links

* Keeping instructions for ''H. spinifer'' and ''H. laoticus''
Exotic Pets
* Images and descriptions of various species of ''Heterometrus''
Mes Scorpions
{{Taxonbar, from=Q138995 Scorpionidae Scorpion genera Taxa named by Christian Gottfried Ehrenberg