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The haddock (''Melanogrammus aeglefinus'') is a saltwater ray-finned fish from the family Gadidae, the true cods. It is the only species in the monotypic genus ''Melanogrammus''. It is found in the North Atlantic Ocean and associated seas where it is an important species for fisheries, especially in northern Europe where it is marketed fresh, frozen and
smoked Smoking is the process of flavoring, browning, cooking, or preserving food by exposing it to smoke from burning or smoldering material, most often wood. Meat, fish, and ''lapsang souchong'' tea are often smoked. In Europe, alder is the tradi ...
; smoked varieties include the Finnan haddie and the
Arbroath smokie The Arbroath smokie is a type of smoked haddock, and is a speciality of the town of Arbroath in Angus, Scotland. History The Arbroath smokie is said to have originated in the small fishing village of Auchmithie, three miles northeast of A ...
.


Description

The haddock has the elongated, tapering body shape typical of members of the cod family. It has a relatively small mouth which does not extend to below the eye; with the lower profile of the face being straight and the upper profile slightly rounded, this gives its snout a characteristic wedge-shaped profile. The upper jaw projects beyond the lower more so than in the Atlantic cod. There is a rather small barbel on the chin. There are three dorsal fins, the first being triangular in shape and these dorsal fins have 14 to 17 fin rays in the first, 20 to 24 in the second, and 19 to 22 in the third. There are also two
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
s and in these there are 21 to 25 fin rays in the first and 20 to 24 fin rays in the second. The anal and dorsal fins are all separated from each other. The pelvic fins are small with an elongated first fin ray. The upper side of the haddock's body varies in colour from dark grey brown to nearly black while the lower part of the body is dull silvery white. It has a distinctive black
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
contrasting with the whitish background colour and which curves slightly over the pectoral fins. It also has a distinctive oval black blotch or ‘thumbprint’, sometimes called the "Devil's thumbprint", which sits between the lateral line and the pectoral fin, a feature which leads to the name of the genus ''Melanogrammus'' which derives from Greek ''"melanos"'' meaning "black" and ''"gramma"'' meaning letter or signal. The dorsal, pectoral, and caudal fins are dark grey in colour while the anal fins are pale matching the colour of the silvery sides, with black speckles at their bases. The pelvic fins are white with a variable amount of black spots. Occasionally there are differently coloured variants recorded which may be barred, golden on the back or lack the dark shoulder blotch. The longest haddock recorded was in length and weighed . However, haddock are rarely over in length and the vast majority of haddocks caught in the United Kingdom measure between . In eastern Canada waters, haddock range in size from in length and in weight.


Distribution

The haddock has populations on either side of the north Atlantic but it is more abundant in the eastern Atlantic than it is on the North American side. In the north-east Atlantic it occurs from the
Bay of Biscay The Bay of Biscay (), known in Spain as the Gulf of Biscay ( es, Golfo de Vizcaya, eu, Bizkaiko Golkoa), and in France and some border regions as the Gulf of Gascony (french: Golfe de Gascogne, oc, Golf de Gasconha, br, Pleg-mor Gwaskogn), ...
north to
Spitzbergen Svalbard ( , ), also known as Spitsbergen, or Spitzbergen, is a Norwegian archipelago in the Arctic Ocean. North of mainland Europe, it is about midway between the northern coast of Norway and the North Pole. The islands of the group range ...
; however, it is most abundant north of the English Channel. It also occurs around
Novaya Zemlya Novaya Zemlya (, also , ; rus, Но́вая Земля́, p=ˈnovəjə zʲɪmˈlʲa, ) is an archipelago in northern Russia. It is situated in the Arctic Ocean, in the extreme northeast of Europe, with Cape Flissingsky, on the northern island, ...
and the Barents Sea in the Arctic. The largest stocks are in the North Sea, off the Faroe Islands, off Iceland and the coast of Norway but these are discrete populations with little interchange between them. Off North America, the haddock is found from western Greenland south to Cape Hatteras, but the main commercially fished stock occurs from Cape Cod and the Grand Banks.


Habitat and biology

The haddock is a demersal species which occurs at depths from , although it is most frequently recorded at . It is found over substrates made up of rock, sand, gravel or shells and it prefers temperatures of between . Off Iceland and in the Barents Sea, haddock undergo extensive migrations, but in the north western Atlantic its movements are more restricted, consisting of movements to and from their spawning areas. They reach sexual maturity at 4 years old in males and 5 years old in females, except for the population in the North Sea which matures at ages of 2 years in males and 3 years in females. The overall sex ratio is roughly 1:1, but in shallower areas, females predominate, while the males show a preference for waters further offshore. The fecundity of the females varies with size: a fish of length bears 55,000 eggs while a fish at has 1,841,000 eggs. Spawning takes place from depths of around . In the northwestern Atlantic spawning lasts from January to July, although it does not occur simultaneously in all areas, and in the northeastern Atlantic the spawning season runs from February to June, peaking in March and April. The eggs are pelagic with a diameter of , and they take one to three weeks to hatch. Following metamorphosis, the past larval fish remain pelagic until they attain a length of around , when they settle to a demersal habit. Their growth rate shows considerable regional variation and fish at one year old can measure , at 2 years old , up to at 13 years old. Their lifespan is around 14 years. The most important spawning grounds are in the waters off the central coast of Norway, off the southwest of Iceland, and over the Georges Bank. The fish which spawn in inshore waters are normally smaller and younger fish than those which occur in offshore areas. The younger fish have a spawning season which is less than half of that of the larger and older stock offshore. Once hatched the larvae do not appear to travel far from their spawning grounds, however some larvae spawning off the west coast of Scotland are transported into the North Sea through the Fair Isle-
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
Gap or to the northeast of Shetland. In their larval stages, haddock mainly feed on the immature stages of
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
s and the pelagic post-larvae up to in length prey on krill, Larvaceans, decapod larvae, copepods and on small fish. Once they have reached the settled, demersal, post-larval stage,
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
invertebrates become increasingly important although they still feed on pelagic organisms such as krill, however the benthic invertebrates form an increasing part of their diet as they grow. Adults prey on fish such as sand eels, '' Trisopterus esmarkii'', American plaice, gobies, European sprat, and
Atlantic herring Atlantic herring (''Clupea harengus'') is a herring in the family (biology), family Clupeidae. It is one of the most abundant fish species in the world. Atlantic herrings can be found on both sides of the Atlantic Ocean, congregating in large ...
, as well as capelin, silver hake,
American eel The American eel (''Anguilla rostrata'') is a facultative catadromous fish found on the eastern coast of North America. Freshwater eels are fish belonging to the elopomorph superorder, a group of phylogenetically ancient teleosts. The America ...
s and argentines. When a number of fish taken at the same time have their stomach contents sampled, the majority of stomachs contain similar prey, this suggests that haddocks feed in shoals. Shellfish,
sea urchins Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) of ...
, brittlestars and worms are also important prey, especially in the winter. Juvenile haddock are an important prey for larger demersal fish, including other gadoids, while seals prey on the larger fish. The recorded growth rates of haddock underwent significant change over the 30 to 40 years up to 2011. Growth has been more rapid in recent years, with haddock attaining adult size much earlier than was noted 30–40 years ago. However, the degree to which these larger, younger fish contribute to reproductive success of the population is unknown. The growth rates of haddock, however, have slowed in recent years. There is some evidence which indicates that these slower growth rates may be the result of an exceptionally large year class in 2003. The haddock stock periodically has higher than normal productivity; for example in 1962 and 1967, and to a lesser extent, 1974 and 1999. These result in a more southerly distribution of the fish and have a strong effect on the biomass of the spawning stock, but because of high fishing mortality, these revivals do not have any lasting effect on the population. In general, there was above average recruitment from the 1960s up to the early 1980s, similar to recruitment for Atlantic cod and whiting, this has been called the ''gadoid outburst''. There was strong recruitment in 1999 but since then, the recruitment rate has been very low.


Parasites

Cod and related species are plagued by parasites. For example, the
cod worm ''Lernaeocera branchialis'', sometimes called cod worm, is a parasite of marine fish, found mainly in the North Atlantic. It is a marine copepod which starts life as a small pelagic crustacean larva. It is among the largest of copepods, ranging i ...
, ''Lernaeocera branchialis'', starts life as a
copepod Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (inhabiting sea waters), some are benthos, benthic (living on the ocean floor) ...
, a small, free-swimming crustacean larva. The first host used by cod worm is a
flatfish A flatfish is a member of the Ray-finned fish, ray-finned demersal fish order (biology), order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the ...
or
lumpsucker The Cyclopteridae are a family of marine fishes, commonly known as lumpsuckers or lumpfish, in the order Scorpaeniformes. They are found in the cold waters of the Arctic, North Atlantic, and North Pacific oceans. The greatest number of species ar ...
, which they capture with grasping hooks at the front of their bodies. They penetrate the lumpsucker with a thin
filament The word filament, which is descended from Latin ''filum'' meaning " thread", is used in English for a variety of thread-like structures, including: Astronomy * Galaxy filament, the largest known cosmic structures in the universe * Solar filament ...
which they use to suck its blood. The nourished cod worms then mate on the lumpsucker.Matthews B (1998
''An Introduction to Parasitology''
Page 73–74. Cambridge University Press. .
The female worm, with her now fertilized eggs, then finds a cod, or a cod-like fish such as a haddock or whiting. There, the worm clings to the gills while it metamorphoses into a plump, sinusoidal, wormlike body, with a coiled mass of egg strings at the rear. The front part of the worm's body penetrates the body of the cod until it enters the rear bulb of the host's heart. There, firmly rooted in the cod's circulatory system, the front part of the parasite develops like the branches of a tree, reaching into the main artery. In this way, the worm extracts nutrients from the cod's blood, remaining safely tucked beneath the cod's gill cover until it releases a new generation of offspring into the water.


Taxonomy and etymology

The haddock was first formally described as ''Gadus aeglefinus'' in 1758 by Carolus Linnaeus in the 10th edition of volume one of his ''
Systema naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' with a type locality given as "European seas". In 1862 Theodore Nicholas Gill created the genus ''Melanogrammus'' with ''M. aeglefinus'' as its only species. The generic name ''Melanogrammus'' means "black line", a reference to the black lateral line of this species. The specific name is a latinisation of the vernacular names “Egrefin” and “Eglefin”, used in France and England.


Fisheries

Haddock is fished year-round using gear such as Danish seine nets,
trawlers Trawler may refer to: Boats * Fishing trawler, used for commercial fishing * Naval trawler, a converted trawler, or a boat built in that style, used for naval purposes ** Trawlers of the Royal Navy * Recreational trawler, a pleasure boat built tra ...
, long lines and gill nets and is often caught in mixed species fishery with other
groundfish Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They occ ...
species such as
cod Cod is the common name for the demersal fish genus '' Gadus'', belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and one species that belongs to genus ''Gadus'' is commonly not call ...
and whiting. The main fishing grounds in the eastern Atlantic are in the Barents Sea, around Iceland, around the Faeroe Islands, in the North Sea, Celtic Sea, and in the English Channel. Landings in the eastern Atlantic have fluctuated around 200–350 thousand tonnes in the period 1980–2017. During the 1980s, the largest portion of the catch was taken at Rockall but from about 2000, the majority of the catch is caught in the Barents Sea. All the stocks in eastern Atlantic are assessed by
ICES ICES (formerly known as the Institute for Clinical Evaluative Sciences) is an independent, non-profit corporation that applies the study of health informatics for health services research and population-wide health outcomes research in Ontario ...
which publish a recommendations on an annual basis for Total Allowable Catch. In the western Atlantic the eastern Georges Bank haddock stock is jointly assessed on an annual basis by Canada and the United States and the stock is collaboratively managed through the Canada-United States Transboundary Management Guidance Committee, this committee was established in 2000. The commercial catch of haddock in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
was approximately 40–60 thousand tonnes per year between 1920 and 1960, this declined sharply in the late 1960s to between 5 and 30 thousand tonnes per year. Despite a few good year classes post 1970, landings have not returned to historical levels. Haddock currently resides on the Greenpeace seafood red due to concerns regarding the impact of bottom trawls on the marine environment. In contrast, Monterey Bay Aquarium considers haddock a "good alternative". Many haddock fisheries have been certified as sustainable by the Marine Stewardship Council. All seven stocks assessed in the eastern Atlantic are currently considered by ICES to be harvested sustainably. The haddock populations in the western Atlantic (offshore grounds of Georges Bank off New England and Nova Scotia) are also considered to be harvested sustainably.


As food

Haddock is very popular as a food fish. It is sold fresh or preserved by smoking, freezing, drying, or to a small extent canning. Haddock, along with Atlantic cod and plaice, is one of the most popular fish used in British
fish and chips Fish and chips is a popular hot dish consisting of fried fish in crispy batter, served with chips. The dish originated in England, where these two components had been introduced from separate immigrant cultures; it is not known who created t ...
. When fresh, the flesh of haddock is clean and white and its cooking is often similar to that of cod. A fresh haddock fillet will be firm and translucent and hold together well but less fresh fillets will become nearly opaque. Young, fresh haddock and cod fillets are often sold as scrod in Boston, Massachusetts; this refers to the size of the fish which have a variety of sizes, i.e., scrod, markets, and cows. Haddock is the predominant fish of choice in Scotland in a fish supper. It is also the main ingredient of Norwegian fishballs (''fiskeboller''). Unlike cod, haddock is not an appropriate fish for salting and preservation is more commonly effected by
drying Drying is a mass transfer process consisting of the removal of water or another solvent by evaporation from a solid, semi-solid or liquid. This process is often used as a final production step before selling or packaging products. To be considered ...
and
smoking Smoking is a practice in which a substance is burned and the resulting smoke is typically breathed in to be tasted and absorbed into the bloodstream. Most commonly, the substance used is the dried leaves of the tobacco plant, which have bee ...
. The smoking of haddock was highly refined in Grimsby.
Traditional Grimsby smoked fish Traditional Grimsby smoked fish are regionally processed fish food products from the British fishing town of Grimsby, England. Grimsby has long been associated with the sea fishing industry, which once gave the town much of its wealth. At its peak ...
(mainly haddock, but sometimes cod) is produced in the traditional smokehouses in Grimsby, which are mostly family-run businesses that have developed their skills over many generations.Grimsby Traditional Fish Smokers Group
Grimsby fish market sources its haddock from the North East Atlantic, principally Iceland, Norway and the Faroe Islands. These fishing grounds are sustainably managedIcelandic Request on the Evaluation of Icelandic Cod and Haddock Management Plan
and have not seen the large scale depreciation in fish stocks seen in EU waters.European Commission, Communication on Fishing Opportunities for 2009. May 2008
/ref> One popular form of haddock is Finnan haddie which is named after the fishing village of Finnan or Findon in Scotland, where the fish was originally cold-smoked over smouldering peat. Finnan haddie is often poached in milk and served for
breakfast Breakfast is the first meal of the day usually eaten in the morning. The word in English refers to breaking the fasting period of the previous night.Anderson, Heather Arndt (2013)''Breakfast: A History'' AltaMira Press. Various "typical" or "t ...
.Full recipe for Finnan Haddie from Scottish chef John Quigley
The town of
Arbroath Arbroath () or Aberbrothock ( gd, Obar Bhrothaig ) is a former royal burgh and the largest town in the council area of Angus, Scotland, with a population of 23,902. It lies on the North Sea coast some ENE of Dundee and SSW of Aberdeen. The ...
on the east coast of Scotland produces the
Arbroath smokie The Arbroath smokie is a type of smoked haddock, and is a speciality of the town of Arbroath in Angus, Scotland. History The Arbroath smokie is said to have originated in the small fishing village of Auchmithie, three miles northeast of A ...
. This is a hot-smoked haddock which requires no further cooking before eating. Smoked haddock is naturally an off-white colour and it is frequently
dye A dye is a colored substance that chemically bonds to the substrate to which it is being applied. This distinguishes dyes from pigments which do not chemically bind to the material they color. Dye is generally applied in an aqueous solution an ...
d yellow, as are other smoked fish. Smoked haddock is the essential ingredient in the Anglo-Indian dish
kedgeree Kedgeree (or occasionally ) is a dish consisting of cooked, flaked fish (traditionally smoked haddock), boiled rice, parsley, hard-boiled eggs, curry powder, butter or cream, and occasionally sultanas. The dish can be eaten hot or cold. Other ...
, and also in the Scottish dish Cullen skink, a chowder-like soup.


References


Other references

* Alan Davidson, ''North Atlantic Seafood'', 1979, .3
Haddock
''NOAA FishWatch''. Retrieved 5 November 2012. * {{Taxonbar, from=Q202406, from2=Q17379593 Gadidae Commercial fish Fish of the Arctic Ocean Fish of the North Sea Fish of Europe Fish described in 1758 Taxa named by Carl Linnaeus