genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of very large
ray-finned fish
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species.
The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
belonging to the
family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Chondrosteidae
Chondrosteidae is a family (biology), family of extinct marine actinopterygian fishes in the order Acipenseriformes. Three genera are known from the Early Jurassic of Europe, ''Chondrosteus'', ''Gyrosteus'', and ''Strongylosteus''. Included sp ...
. It comprises the type species,
''Gyrosteus mirabilis'', which lived during the early
Toarcian
The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian.
The Toarcian ...
(Late
Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-J ...
) in what is now northern
Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. A possible second species, ''"Gyrosteus" subdeltoideus'', is known from
otoliths
An otolith ( grc-gre, ὠτο-, ' ear + , ', a stone), also called statoconium or otoconium or statolith, is a calcium carbonate structure in the saccule or utricle of the inner ear, specifically in the vestibular system of vertebrates. The sa ...
.
Fossil remains of ''G. mirabilis'' have been recovered from the
Whitby Mudstone Formation
The Whitby Mudstone is a Toarcian (Early Jurassic; ''Falciferum''-''Bifrons'' in regional chronostratigraphy) geological formation in Yorkshire and Worcestershire, England.Ahrensburg erratics assemblage in
Schleswig-Holstein
Schleswig-Holstein (; da, Slesvig-Holsten; nds, Sleswig-Holsteen; frr, Slaswik-Holstiinj) is the northernmost of the 16 states of Germany, comprising most of the historical duchy of Holstein and the southern part of the former Duchy of Sch ...
, northern Germany. It was mentioned but not formally described in subsequent publications and was left as a '' nomen nudum'' for more than 25 years. Then in 1889 it was featured and formally described by
Arthur Smith Woodward
Sir Arthur Smith Woodward, FRS (23 May 1864 – 2 September 1944) was an English palaeontologist, known as a world expert in fossil fish. He also described the Piltdown Man fossils, which were later determined to be fraudulent. He is not relate ...
. ''Gyrosteus'' was thought to be exclusive of the “British faunal province” and separated from the “Germanic faunal province” until the discovery of a
hyomandibula
The hyomandibula, commonly referred to as hyomandibular one( la, os hyomandibulare, from el, hyoeides, "upsilon-shaped" (υ), and Latin: mandibula, "jawbone") is a set of bones that is found in the hyoid region in most fishes. It usually plays ...
in the baltic realm, mostly populated by Germanic fauna, which possibly implicates that Baltic region represented an interdigitating zone between both regions.
The members of the
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Gyrosteus'' were massive fishes, with a maximum calculated standard length of to , and with a reported hyomandibula reaching .