Gray Francolin
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The grey francolin (''Ortygornis pondicerianus''), also known as "manu moa" or "chicken bird", is a species of francolin found in the plains and drier parts of the
Indian subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
and
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. This species was formerly also called the grey partridge, not to be confused with the European grey partridge. They are mainly ground-living birds and are found in open cultivated lands as well as scrub forest and their local name of ''teetar'' is based on their calls, a loud and repeated ''Ka-tee-tar...tee-tar'' which is produced by one or more birds. The term ''teetar'' can also refer to other partridges and quails. During the breeding season calling males attract challengers, and decoys were used to trap these birds especially for
fighting Combat ( French for ''fight'') is a purposeful violent conflict meant to physically harm or kill the opposition. Combat may be armed (using weapons) or unarmed ( not using weapons). Combat is sometimes resorted to as a method of self-defense, or ...
.


Taxonomy

The grey francolin was formally described in 1789 by the German naturalist
Johann Friedrich Gmelin , fields = , workplaces = University of GöttingenUniversity of Tübingen , alma_mater = University of Tübingen , doctoral_advisor = Philipp Friedrich GmelinFerdinand Christoph Oetinger , academic_advisors = , doctora ...
in his revised and expanded edition of
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
's ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
''. He placed it with all the grouse like birds in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Tetrao ''Tetrao'' is a genus of birds in the grouse subfamily known as capercaillies. They are some of the largest living grouse. Taxonomy The genus ''Tetrao'' was introduced in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his ...
'' and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Tetrao pondicerianus''. Gmelin based his description on "Le perdix de Pondichéry" that had been described in 1782 by the French naturalist
Pierre Sonnerat Pierre Sonnerat (18 August 1748 – 31 March 1814) was a French natural history, naturalist, colonial administrator, writer and List of explorers, explorer. He described numerous species of plants and animals on his travels and is honoured in th ...
in his ''Voyage aux Indes orientales et a la Chine''. The grey francolin was formerly placed in the genus '' Francolinus''. Based on a phylogenetic study published in 2019 the grey francolin, together with the
crested francolin The crested francolin (''Ortygornis sephaena'') is a species of bird in the family Phasianidae. It is found in southern Africa. One of its subspecies, ''Ortygornis sephaena rovuma'', is sometimes considered a separate species, Kirk's francolin. ...
and
swamp francolin The swamp francolin (''Ortygornis gularis''), also called swamp partridge, is a francolin species native to the foothills of the Himalayas in northern India and Nepal. It is considered extinct in the Ganges-Brahmaputra delta in Bangladesh. It is ...
, were moved to the resurrected genus ''Ortygornis'' that had been introduced in 1852 by the German naturalist
Ludwig Reichenbach Heinrich Gottlieb Ludwig Reichenbach (8 January 1793 – 17 March 1879) was a German botanist and ornithologist. It was he who first requested Leopold Blaschka to make a set of glass marine invertebrate models for scientific education and museu ...
. The genus name combines the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''ortux'' meaning "quail" with ''ornis'' meaning "bird". The specific epithet ''pondicerianus'' is from the toponym Pondicherry, a town in southeast India. Three
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''O. p. mecranensis'' ( Zarudny & Härms, 1913) – south Iran and south Pakistan * ''O. p. interpositus'' ( Hartert, E, 1917) – east Pakistan, north India and Nepal * ''O. p. pondicerianus'' ( Gmelin, JF, 1789) – south India and Sri Lanka


Description

This bird is a medium-sized francolin, with males averaging and females averaging . The males weigh whereas the weight of the females is . The francolin is barred throughout and the face is pale with a thin black border to the pale throat. The only similar species is the
painted francolin The painted francolin or painted partridge (''Francolinus pictus'') is a species of francolin found in grassy areas in central and southern India and in the lowlands of southeastern Sri Lanka. They are easily detected by their loud calls especia ...
, which has a rufous vent. The male can have up to two spurs on the legs while females usually lack them. Subspecies ''mecranensis'' is palest and found in arid North-Western India, Eastern Pakistan and Southern Iran. Subspecies ''interpositus'' is darker and intermediate found in northern India. The nominate race in the southern peninsula of India has populations with a darker rufous throat,
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
and is richer brown. They are weak fliers and fly short distances, escaping into undergrowth after a few spurts of flight. In flight it shows a chestnut tail and dark primaries. The race in Sri Lanka is sometimes given the name ''ceylonensis'' or considered as belonging to the nominate.


Distribution and habitat

The grey francolin is normally found foraging on bare or low grass covered ground in scrub and open country, and is rarely found above an altitude of 500 m above sea level in India, and 1200 m in Pakistan. The distribution is south of the foothills of the
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
s westwards to the
Indus Valley The Indus ( ) is a transboundary river of Asia and a trans-Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in Western Tibet, flows northwest through the disputed region of Kashmir, ...
and eastwards to
Bengal Bengal ( ; bn, বাংলা/বঙ্গ, translit=Bānglā/Bôngô, ) is a geopolitical, cultural and historical region in South Asia, specifically in the eastern part of the Indian subcontinent at the apex of the Bay of Bengal, predom ...
. It is also found in north-western Sri Lanka. Introduced populations are found in the
Andaman Andaman may refer to: * Andaman Islands * Andaman Sea * ''Andaman'' (1998 film), a Kannada-language film * ''Andaman'' (2016 film), a Tamil-language film * ''Andaman'' (2021 film), a Hindi-language film See also * Andaman and Nicobar Islands ...
and Chagos Islands. They have been introduced to
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
in the United States of America and
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
, along with several other species of francolin.


Behaviour and ecology

The loud calls of the birds are commonly heard early in the mornings. Pairs of birds will sometimes engage in a duet. The female call is a ''tee...tee...tee'' repeated and sometimes a ''kila..kila..kila'' and the challenge call ''kateela..kateela..kateela'' is a duet. They are usually seen in small groups. The main breeding season is April to September and the nest is a hidden scrape on the ground. The nest may sometimes be made above ground level in a niche in a wall or rock. The clutch is six to eight eggs, but larger clutches, potentially reflecting intraspecific brood parasitism, have been noted. Food includes seeds, grains as well as insects, particularly termites and beetles (especially
Tenebrionidae Darkling beetle is the common name for members of the beetle family Tenebrionidae. The number of species in the Tenebrionidae is estimated at more than 20,000 and the family is cosmopolitan in distribution. Taxonomy ''Tenebrio'' is the Latin gen ...
and Carabidae). They may occasionally take larger prey such as snakes. They roost in groups in low thorny trees. Several species of feather mites, helminth and blood parasites have been described from the species.


Status

They are hunted in much of their range using low nets and easily caught using calling decoy birds.


In culture

The species has long been domesticated in areas of northern Indian subcontinent where it is used for fighting. The domesticated birds can be large at around 500-600g, compared to 250g for wild birds. They are usually carefully reared by hand and become as tame and confiding as a pet dog. Several authors have described the running of the birds as being particularly graceful:
John Lockwood Kipling John Lockwood Kipling (6 July 1837 – 26 January 1911) was an English art teacher, illustrator and museum curator who spent most of his career in India. He was the father of the author Rudyard Kipling. Life and career Lockwood Kipling was b ...
,
Rudyard Kipling Joseph Rudyard Kipling ( ; 30 December 1865 – 18 January 1936)''The Times'', (London) 18 January 1936, p. 12. was an English novelist, short-story writer, poet, and journalist. He was born in British India, which inspired much of his work. ...
's father, wrote of this and other partridges such as the chukar partridge:


References


Further reading

* * Johnson, J Mangalaraj (1968) Grey Partridge abandoning nest on removal of grass cover over its path to nest. Indian Forester 94:780. * Davis, G (1939) On Indian Grey and Black Partridges (''Francolinus pondicerianus'' and ''Francolinus francolinus''). The Avicultural Magazine, 5 4(5):148-151. * Gabriel, A (1970) Some observations on the Ceylon Grey Partridge. Loris 12(1):60-62. * *Sharma, IK (1983) The Grey Partridge (''Francolinus pondicerianus'') in the Rajasthan desert. Annals Arid Zone. 22(2), 117–120. *Soni, VC (1978) Intersexuality in the Gray Partridge. Game Bird Breeders Avicult. Zool. Conserv. Gaz. 27(7), 12–13. *Hartert, E (1917) Notes on game-birds. VII. The forms of ''Francolinus pondicerianus''. Novit. Zool. 24, 287–288. * Purwar, RS (1975) Anatomical, neurohistological and histochemical observations on the tongue of Francolinus pondicerianus (grey partridge or safed teeter). Acta Anat. 93(4):526-33. * Purwar, RS (1976) Neuro-histochemical observations on the pancreas of Francolinus pondicerianus (grey partridge or safed teeter) as revealed by the cholinesterase technique. Z. Mikrosk. Anat. Forsch. 90(6):1009-16.


External links


Aviculture


{{Taxonbar, from=Q939650 grey francolin Birds of the Middle East Birds of South Asia grey francolin grey francolin Taxobox binomials not recognized by IUCN