Gouramy
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Gouramis, or gouramies , are a group of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
anabantiform fishes that comprise the family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeasterly towards Korea. The name "gourami", of
Indonesian Indonesian is anything of, from, or related to Indonesia, an archipelagic country in Southeast Asia. It may refer to: * Indonesians, citizens of Indonesia ** Native Indonesians, diverse groups of local inhabitants of the archipelago ** Indonesian ...
origin, is also used for fish of the families Helostomatidae and Anabantidae. Many gouramis have an elongated, feeler-like ray at the front of each of their
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
s. All living species show parental care until fry are free swimming: some are mouthbrooders, like the Krabi mouth-brooding betta (''Betta Simplex''), and others, like the Siamese fighting fish (''Betta splendens''), build bubble nests. Currently, about 133 species are recognised, placed in four subfamilies and about 15
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
. The name Polyacanthidae has also been used for this family. Some fish now classified as gouramis were previously placed in family Anabantidae. The subfamily Belontiinae was recently demoted from the family Belontiidae. As
labyrinth fish The Anabantoidei are a suborder of anabantiform ray-finned freshwater fish distinguished by their possession of a lung-like labyrinth organ, which enables them to breathe air. The fish in the Anabantoidei suborder are known as anabantoids or ...
es, gouramis have a
lung The lungs are the primary organs of the respiratory system in humans and most other animals, including some snails and a small number of fish. In mammals and most other vertebrates, two lungs are located near the backbone on either side of t ...
-like labyrinth organ that allows them to gulp air and use atmospheric oxygen. This organ is a vital adaptation for fish that often inhabit warm, shallow, oxygen-poor water.


Subfamilies and genera

The family Osphronemidae is divided into the following subfamilies and genera: * family Osphronemidae
van der Hoeven Van der Hoeven (Vanderhoeven) or Van der Hoeve is a surname of noble Dutch family. People with the name include: * Abraham Pruijs van der Hoeven (1829–1907), Dutch colonial governor of Aceh * Agnes van Ardenne-van der Hoeven (b. 1950), Dutch pol ...
, 1832
** Subfamily Belontiinae
Liem Many ethnic Chinese people have lived in Indonesia for many centuries. Over time, especially under social and political pressure during the New Order era, most Chinese Indonesians have adopted names that better match the local language. History ...
, 1962
*** '' Belontia''
Myers Myers as a surname has several possible origins, e.g. Old French ("physician"), Old English ("mayor"), and Old Norse ("marsh"). People *Abram F. Myers (born 1889), chair of the Federal Trade Commission and later general counsel and board chai ...
, 1923
** Subfamily Osphroneminae van der Hoeven, 1832 *** ''
Osphronemus ''Osphronemus'' is a genus of large gouramis, the only genus within the subfamily Osphroneminae. These fish are known as the giant gouramis and are native to rivers, lakes, pools, swamps and floodplains in Southeast Asia, with ''O. exodon'' fro ...
'' Lacepède, 1801 ** Subfamily
Luciocephalinae Luciocephalinae is a subfamily of the gourami family Osphronemidae. The members of this subfamily differ from the other groups within the gourami family by having a reduced number of rays supporting the branchiostegal membrane, five rather than ...
Bleeker Bleeker is a Dutch occupational surname. Bleeker is an old spelling of ''(linnen)bleker'' ("linen bleacher").Luciocephalus ''Luciocephalus'' is a genus of gouramies native to Southeast Asia. Both are extremely specialized niche predators native to parts of Thailand, Malaysia, Indonesia and Brunei Darussalam; in addition, both species - like a number of other osphrone ...
'' Bleeker, 1851 *** ''
Sphaerichthys ''Sphaerichthys'' is a genus of gouramies native to Southeast Asia known as chocolate gourami. Species There are currently four recognized species in this genus: * '' Sphaerichthys acrostoma'' Vierke, 1979 (Giant chocolate gourami) * '' Sphaeri ...
''
Canestrini Giovanni Canestrini (26 December 1835 – 14 February 1900) was an Italian naturalist and biologist and translator who was a native of Revò. Career He initially studied in Gorizia and Meran, then furthered his education in natural sciences at ...
, 1860
*** ''
Ctenops The frail gourami (''Ctenops nobilis'') is a species of gourami native to northeastern India and Bangladesh found in lakes, ponds, rivers, and streams with plentiful vegetation overgrowth. This species grows to a length of . It is only seldom f ...
'' McClelland, 1845 *** ''
Parasphaerichthys ''Parasphaerichthys'' is a genus of gouramies known only from streams and freshwater pools in the Irrawaddy basin of Myanmar. They are the small gouramies that, depending on the exact species, reach up to in length. Species There are currently ...
'' Prashad & Mukerji, 1929 *** ''
Trichopodus ''Trichopodus'' (formerly included in ''Trichogaster'') is a genus of tropical freshwater labyrinth fish of the gourami family found in Southeast Asia. Gouramis of the genus ''Trichopodus'' are closely related to those of ''Trichogaster'' (former ...
'' Lacepède, 1801 ** Subfamily
Macropodusinae The Macropodusinae are a subfamily of freshwater anabantiform fishes in the gourami family Osphronemidae, which includes the paradisefish, fighting fish and licorice gouramis. Like all members of the family, these are air breathing fishes ...
Hoedeman, 1948 *** '' Betta'' Bleeker, 1850 *** ''
Parosphromenus ''Parosphromenus'' is a genus of gouramies native to freshwater in Southeast Asia. All species are highly specialized peat swamp inhabitants native to southeast Asia, and the males are usually brightly colored in breeding dress; however, their ne ...
'' Bleeker, 1877 *** ''
Macropodus ''Macropodus'' is a genus of small to medium-sized labyrinth fish native to freshwater habitats in eastern Asia. Most species are restricted to southern China (including Hong Kong and Taiwan) and Vietnam, but ''M. opercularis'' occurs as far nort ...
'' Lacepède, 1801 *** ''
Malpulutta The ornate paradisefish (''Malpulutta kretseri'') or spotted gourami, is a species of gourami Endemism, endemic to Sri Lanka. It is the only recognized species in its genus. It inhabits shallow, slow-flowing streams in forested areas shaded with ...
''
Deraniyagala Deraniyagala is a town in the Kegalle district in the Sabaragamuwa Province of Sri Lanka. Deraniyaga city area consists major government administrative offices, textile industries and transportation hub and the district hospital which has seve ...
, 1937
*** ''
Pseudosphromenus ''Pseudosphromenus'' is a genus of medium-small gouramies native to south and southeast Asia. Species There are currently 2 recognized species in this genus: * '' Pseudosphromenus cupanus'' (G. Cuvier, 1831) (Spiketail paradisefish) * '' Pseudos ...
'' Bleeker, 1879 *** ''
Trichopsis ''Trichopsis'' is a genus of gouramies native to Southeast Asia. Species There are currently three recognized species in this genus: * ''Trichopsis pumila'' ( J. P. Arnold, 1936) (Pygmy gourami) * ''Trichopsis schalleri'' Ladiges, 1962 (Threest ...
'' Canestrini, 1860 ** Subafmily
Trichogastrinae ''Trichogaster'' is a genus of gouramis native to South Asia from Pakistan to Myanmar. It is the only genus in the monotypic subfamily Trichogastrinae as set out in the 5th Edition of ''Fishes of the World'', although that book states that there ...
Bleeker, 1879 *** '' Trichogaster''
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Schneider Schneider may refer to: Hospital * Schneider Children's Medical Center of Israel People * Schneider (surname) Companies and organizations * G. Schneider & Sohn, a Bavarian brewery company * Schneider Rundfunkwerke AG, the former owner of th ...
, 1801


As food

Giant gouramis, ''Osphronemus goramy'', or Kaloi in Malay language, are eaten in some parts of the world. In Maritime Southeast Asian countries, they are often deep-fried and served in sweet-sour sauce, chili sauce, and other spices. The paradise fish, ''Macropodus opercularis'', and other members of that genus are the target of a cannery industry in China, the products of which are available in Asian supermarkets around the world. Gouramis are particularly found in
Sundanese cuisine Sundanese cuisine is the cuisine of the Sundanese people of Western Java, and Banten, Indonesia. It is one of the most popular foods in Indonesia. Sundanese food is characterised by its freshness; the famous lalab eaten with sambal and also k ...
. In Malaysia, Indonesia, Thailand, Philippines, Brunei, gouramis are readily fished at streams, brooks, canal, rivers and many more large water area systems.


In the aquarium

Numerous gourami species, such as the dwarf gourami,
pearl gourami The pearl gourami (''Trichopodus leerii'') is a species of gourami native to Southeast Asia. Description It reaches about TL. Its body is a brownish-silver colour, covered in a pearl-like pattern with a distinct black line running from the f ...
, are popular aquarium fish widely kept throughout the world. They are sought after due to their bright colours and relative intelligence, being able to recognise their owners and “greeting” them, having a desire to explore the plants and rocks placed across their aquarium, and displaying extensive paternal care with the males protecting the eggs until they hatch, and building a foam raft to keep them afloat. As labyrinth fish, they will often swim near the top of the tank in order to breathe air. As with other tropical freshwater fish, an
aquarium heater An aquarium heater is a device used in the fishkeeping hobby to warm the temperature of water in aquariums. Most tropical Freshwater aquarium, freshwater and marine aquariums are maintained at temperatures that range from 22 to 30 °C (71-86&n ...
is often used. Gouramis will eat either prepared or live
foods Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is ingest ...
. Some species can grow quite large and are unsuitable for the general hobbyist. Big Gouramis may become territorial with fish that are colourful and a considerable size to them, however that generally depends on the individual’s temperament as some gourami will be more tolerant of tankmates than others. Gouramis may nip at other fish, and males should never be kept together as they will become aggressive.


Compatibility

Generally regarded as peaceful, gouramis are still capable of harassing or killing smaller or long-finned fish. Depending on the species, adult and juvenile males have been known to spar with one another. Aggression can also occur as a result of overcrowding. Gouramis have been housed with many species, such as
danio ''Danio'' is a genus of small freshwater fish in the family Cyprinidae found in South and Southeast Asia, commonly kept in aquaria. They are generally characterised by a pattern of horizontal stripes, rows of spots or vertical bars. Some species ...
s, mollies, silver dollars, Neon tetras, and plecostomus catfish. Compatibility depends on the species of gourami and the fish it is housed with. Some species (e.g., ''
Macropodus ''Macropodus'' is a genus of small to medium-sized labyrinth fish native to freshwater habitats in eastern Asia. Most species are restricted to southern China (including Hong Kong and Taiwan) and Vietnam, but ''M. opercularis'' occurs as far nort ...
'' or '' Belontia'') are highly aggressive or predatory and may harass or kill smaller or less aggressive fish; whereas, others (''
Parosphromenus ''Parosphromenus'' is a genus of gouramies native to freshwater in Southeast Asia. All species are highly specialized peat swamp inhabitants native to southeast Asia, and the males are usually brightly colored in breeding dress; however, their ne ...
'' and ''
Sphaerichthys ''Sphaerichthys'' is a genus of gouramies native to Southeast Asia known as chocolate gourami. Species There are currently four recognized species in this genus: * '' Sphaerichthys acrostoma'' Vierke, 1979 (Giant chocolate gourami) * '' Sphaeri ...
'', for instance) are very shy or have specific water requirements and thus will be outcompeted by typical community fish.


Gallery

Image:Male_female_dwarf_gourami.jpg, Female and male dwarf gouramis (''Trichogaster lalius'') showing sexual dimorphism Image:Makropode_Quang_Binh-Makropode_060114_1.jpg, ''
Macropodus erythropterus The red-backed paradise fish (''Macropodus erythropterus'') is a species of gourami endemic to streams in Vietnam. This species grows to a standard length of , and it is sometimes seen in the aquarium trade. Range and habitat The red-backed p ...
'' Image:Makropode_Makropodus_opercularis_060114_1.jpg, Paradise fish (''Macropodus opercularis'') Image:giant.gourami.arp.jpg, Giant gourami (''Osphronemus goramy'') Image:Trichogaster trichopterus 7.jpg,
Three spot gourami The three spot gourami (''Trichopodus trichopterus''), also known as the opaline gourami, blue gourami, and gold gourami, is a species of fish native to southeastern Asia, but also introduced elsewhere. This gourami gets its name from the two sp ...
(''Trichopodus trichopterus'') Image:Gold_gourami.jpg ,
Gold gourami The three spot gourami (''Trichopodus trichopterus''), also known as the opaline gourami, blue gourami, and gold gourami, is a species of fish native to southeastern Asia, but also introduced elsewhere. This gourami gets its name from the two sp ...

color morph of ''Trichopodus trichopterus''


See also

The name "gourami" is used of several other related fish that are now placed in different families: * Kissing gourami (the sole species in the family Helostomatidae) *
Climbing gourami The Anabantidae are a family of ray-finned fish within the order Anabantiformes commonly called the climbing gouramies or climbing perches. The family includes about 34 species. As labyrinth fishes, they possess a labyrinth organ, a structure i ...
s (family Anabantidae)


References

* * {{Taxonbar, from=Q726399 * Anabantiformes