''Gigantspinosaurus'' () is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of herbivorous
ornithischian
Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek st ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Middle to
Late Jurassic
The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987.
In European lithostratigraphy, the name ...
. It was a
stegosaur
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, ...
found in
China
China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
.
Discovery
The first fossil was found in 1985 by
Ouyang Hui
Ouyang () is a Chinese surname. It is the most common two-character Chinese compound surname, being the only two-character name of the 400 most common Chinese surnames, according to a 2013 study, and is one of the few two-character surnames that h ...
at
Pengtang near
Jinquan and was reported upon in 1986 by
Gao Ruiqi and colleagues, mistaking it for a specimen of ''
Tuojiangosaurus
''Tuojiangosaurus'' (meaning "Tuo River lizard") is a genus of herbivorous stegosaurian dinosaur from the Late Jurassic Period, recovered from the Upper Shaximiao Formation of what is now Sichuan Province in China.
Description
''Tuojiangosauru ...
''. The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Gigantspinosaurus sichuanensis'', was described and named by Ouyang in 1992 in an abstract of a lecture.
The generic name is derived from
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''gigas'' or ''giganteus'', "enormous", and ''spina'', "spine", in reference to the gigantic shoulder spines. The
specific name refers to Sichuan.
The name was generally considered a ''
nomen nudum
In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
'' in the West,
until in 2006 it was disclosed that the abstract contained a sufficient description.
[ Despite its uncertain nomenclatural status, images of ''Gigantspinosaurus'' had appeared in several sources. Public awareness of this animal was increased in early 2006 when Tracy Ford,] considering it a validly established taxon, published a short article on reconstructing it. Ford suggested that earlier reconstructions of ''Gigantspinosaurus'' attached the shoulder spines upside-down, and his new reconstruction shows the spine extending somewhat upwards, ending higher than the top of the animal's back. Susannah Maidment Susannah Catherine Rose Maidment is a British palaeontologist at the Natural History Museum, London. She is internationally recognised for her research on ornithischian dinosaur evolution, and was awarded the 2016 Hodson Award of the Palaeontologic ...
and Wei Guangbiao in 2006 concluded that ''G. sichuanensis'' was a valid taxon in their review of Late Jurassic Chinese stegosaurs, but did not redescribe it because at that time it was under study by Zigong Dinosaur Museum
The Zigong Dinosaur Museum () is located near the city of Zigong, Sichuan, China, in the township of Dashanpu.
History
The museum sits on top of a large concentration of a diverse dinosaur assemblage from the Shaximiao Formation. The museu ...
staff. In fact, a Chinese redescription by Peng Guangzhao
Peng may refer to:
* Peng (surname) (彭), a Chinese name
* Peng (state) (大彭), a state during the late Shang dynasty
* Peng (mythology) (鵬), a legendary Chinese creature
* ''Peng!'', 1992 album by Stereolab
* ''PENG!'', a 2005 comic
* P.Eng. ...
and colleagues in 2005 would predate Maidment's publication.
The holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, ZDM 0019, was found in layers of the Upper Shaximiao Formation of Zigong
Zigong (, ), formed by the merger of the two former towns of Ziliujing (Tzuliuching, literally "self-flow well") and Gongjing (Kungching, literally "offering well"), is a prefecture-level city in Sichuan, southwestern China.
Demographics
Accord ...
(Sichuan province
Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
), which date to the Oxfordian. It consists of a partial skeleton of a probably subadult individual missing the skull (though the lower jaws are present), hind feet, and the tail end. Apart from skeletal elements also plates, spines and scutes have been found. At the left shoulder an impression of the skin had been preserved. The specimen is part of the collection of the Zigong Dinosaur Museum and has as a mounted restored skeleton been on display since 1996. In 2005 Peng e.a. reported a second specimen, ZDM 0156, a pelvis found at Chenjia near Fuquan.
Description
''Gigantspinosaurus'' was described by Peng and colleagues as a "medium-sized stegosaur". It was estimated by Gregory S. Paul
Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dino ...
in 2010 to have been about long and in weight. ''Gigantspinosaurus'' has a distinctive appearance with relatively small dorsal plates and greatly enlarged shoulder spines, ''spinae parascapulares'', twice the length of the shoulder blades on which they rested via large flat bases. The plates on the neck are small and triangular. The head must have been relatively large with thirty teeth in each lower jaw. The hips are very broad and the low neural spines of the four sacral vertebrae and the first tail vertebra have been fused into a single plate. It also had robust forelimbs.
The skin impressions were described by Xing Lida and colleagues in 2008. They cover a surface of and show rosettes with a central pentagonal or hexagonal scale surrounded by thirteen to fourteen ridged smaller square, pentagonal or hexagonal scales with a diameter of .
Classification
A study by Maidment indicated that ''Gigantspinosaurus'' is the most basal known member of the Stegosauria
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, ...
. , A 2018 redescription by Hao ''et al'' clarified aspects of the anatomy and found that it was an intermediate basal stegosaur, sharing basal traits with huayangosaurids as well as somewhat more advanced traits with other stegosaurs. Nevertheless, this analysis found that it was not the most basal stegosaur, as the huayangosaurids ''Chungkingosaurus
''Chungkingosaurus'', meaning "Chongqing Lizard", is a genus of herbivorous dinosaur from the Late Jurassic Upper Shaximiao Formation in what is now China. It is a member of the Stegosauria.
Description
According to Dong e.a. the ''Chungkingo ...
'' and ''Huayangosaurus
''Huayangosaurus'' is a genus of stegosaurian dinosaur from the Middle Jurassic of China. The name derives from "Huayang" (華陽), an alternate name for Sichuan (the province where it was discovered), and "saurus", meaning "lizard". It lived ...
'' were considered more basal.
See also
* Timeline of stegosaur research
This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period ...
References
{{Taxonbar, from=Q134984
Stegosaurs
Late Jurassic dinosaurs of Asia
Fossil taxa described in 1992
Paleontology in Sichuan
Ornithischian genera