botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...

, a fruit is the

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

-bearing structure in

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s that is formed from the

ovary The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. ...

after

flowering A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...

. Fruits are the means by which flowering plants (also known as

angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s) disseminate their

s. Edible fruits in particular have long propagated using the movements of humans and animals in a

symbiotic relationship Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...

that is the means for

seed dispersal In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...

for the one group and

nutrition Nutrition is the biochemical and physiological process by which an organism uses food to support its life. It provides organisms with nutrients, which can be metabolized to create energy and chemical structures. Failure to obtain sufficient n ...

for the other; in fact, humans and many animals have become dependent on fruits as a source of

food Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is inge ...

. Consequently, fruits account for a substantial fraction of the world's

agricultural Agriculture or farming is the practice of cultivating Plant, plants and livestock. Agriculture was the key development in the rise of Sedentism, sedentary human civilization, whereby farming of Domestication, domesticated species created food ...

output, and some (such as the

apple An apple is an edible fruit produced by an apple tree (''Malus domestica''). Apple fruit tree, trees are agriculture, cultivated worldwide and are the most widely grown species in the genus ''Malus''. The tree originated in Central Asia, wh ...

and the

pomegranate The pomegranate (''Punica granatum'') is a fruit-bearing deciduous shrub in the family Lythraceae, subfamily Punicoideae, that grows between tall. The pomegranate was originally described throughout the Mediterranean Basin, Mediterranean re ...

) have acquired extensive cultural and symbolic meanings. In common language usage, "fruit" normally means the seed-associated fleshy structures (or produce) of plants that typically are sweet or sour and edible in the raw state, such as

banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", distinguis ...

grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years ago, ...

lemon The lemon (''Citrus limon'') is a species of small evergreen trees in the flowering plant family Rutaceae, native to Asia, primarily Northeast India (Assam), Northern Myanmar or China. The tree's ellipsoidal yellow fruit is used for culin ...

oranges An orange is a fruit of various citrus species in the family Rutaceae (see list of plants known as orange); it primarily refers to ''Citrus'' × ''sinensis'', which is also called sweet orange, to distinguish it from the related ''Citrus × ...

, and

strawberries The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...

. In botanical usage, the term "fruit" also includes many structures that are not commonly called "fruits" in everyday language, such as nuts,

bean A bean is the seed of several plants in the family Fabaceae, which are used as vegetables for human or animal food. They can be cooked in many different ways, including boiling, frying, and baking, and are used in many traditional dishes th ...

pods,

corn Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...

kernels Kernel may refer to: Computing * Kernel (operating system), the central component of most operating systems * Kernel (image processing), a matrix used for image convolution * Compute kernel, in GPGPU programming * Kernel method, in machine learnin ...

tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...

es, and

wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...

grains.

Botanical vs. culinary

Many common language terms used for fruit and seeds differ from botanical classifications. For example, in botany, a ''fruit'' is a ripened

carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

that contains seeds; e.g., an apple, pomegranate, tomato or a pumpkin. A ''

nut Nut often refers to: * Nut (fruit), fruit composed of a hard shell and a seed, or a collective noun for dry and edible fruits or seeds * Nut (hardware), fastener used with a bolt Nut or Nuts may also refer to: Arts, entertainment, and media Com ...

'' is a type of fruit (and not a seed), and a ''seed'' is a ripened

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

. In culinary language, a ''fruit'' is the sweet- or not sweet- (even sour-) tasting produce of a specific plant (e.g., a peach, pear or lemon); ''nuts'' are hard, oily, non-sweet plant produce in shells (

hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus ''Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according t ...

acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera ''Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and borne ...

). ''

Vegetables Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the flowers, fruits, stems, ...

'', so called, typically are

savory Savory or Savoury may refer to: Common usage * Herbs of the genus ''Satureja'', particularly: ** Summer savory (''Satureja hortensis''), an annual herb, used to flavor food ** Winter savory (''Satureja montana''), a perennial herb, also used to ...

or non-sweet produce (

zucchini The zucchini (; plural: zucchini or zucchinis), courgette (; plural: courgettes) or baby marrow (''Cucurbita pepo'') is a summer squash, a vining herbaceous plant whose fruit are harvested when their immature seeds and epicarp (rind) are stil ...

, lettuce, broccoli, and tomato); but some may be sweet-tasting (sweet potato). Examples of botanically classified fruit that typically are called vegetables include:

cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated Vine#Horticultural climbing plants, creeping vine plant in the Cucurbitaceae family that bears usually cylindrical Fruit, fruits, which are used as culinary vegetables.

pumpkin A pumpkin is a vernacular term for mature winter squash of species and varieties in the genus ''Cucurbita'' that has culinary and cultural significance but no agreed upon botanical or scientific meaning. The term ''pumpkin'' is sometimes use ...

, and

squash Squash may refer to: Sports * Squash (sport), the high-speed racquet sport also known as squash racquets * Squash (professional wrestling), an extremely one-sided match in professional wrestling * Squash tennis, a game similar to squash but pla ...

( all are cucurbits);

peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible Seed, seeds. It is widely grown in the tropics and subtropics, important to both small ...

s, and

pea The pea is most commonly the small spherical seed or the seed-pod of the flowering plant species ''Pisum sativum''. Each pod contains several peas, which can be green or yellow. Botanically, pea pods are fruit, since they contain seeds and d ...

s ( all legumes);

eggplant Eggplant ( US, Canada), aubergine ( UK, Ireland) or brinjal (Indian subcontinent, Singapore, Malaysia, South Africa) is a plant species in the nightshade family Solanaceae. ''Solanum melongena'' is grown worldwide for its edible fruit. Mos ...

, bell pepper (or sweet pepper), and tomato (see image). The spices

chili pepper Chili peppers (also chile, chile pepper, chilli pepper, or chilli), from Nahuatl '' chīlli'' (), are varieties of the berry-fruit of plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for ...

and

allspice Allspice, also known as Jamaica pepper, myrtle pepper, pimenta, or pimento, is the dried unripe berry (botany), berry of ''Pimenta dioica'', a Canopy (forest), midcanopy tree native to the Greater Antilles, southern Mexico, and Central America, ...

are fruits, botanically speaking. In contrast,

rhubarb Rhubarb is the fleshy, edible stalks ( petioles) of species and hybrids (culinary rhubarb) of ''Rheum'' in the family Polygonaceae, which are cooked and used for food. The whole plant – a herbaceous perennial growing from short, thick rhizo ...

is often called a fruit when used in making

pies A pie is a baked dish which is usually made of a pastry dough casing that contains a filling of various sweet or savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts (pecan pie), brown sugar (sugar pie), sweeten ...

, but the edible produce of rhubarb is actually the leaf stalk or petiole of the plant. Edible

gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνό� ...

seeds are often given fruit names, e.g.,

ginkgo ''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus within ...

nuts and

pine nut Pine nuts, also called piñón (), pinoli (), pignoli or chilgoza (), are the edible seeds of pines (family Pinaceae, genus ''Pinus''). According to the Food and Agriculture Organization, only 29 species provide edible nuts, while 20 are trade ...

s. Botanically, a

cereal A cereal is any Poaceae, grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, Cereal germ, germ, and bran. Cereal Grain, grain crops are grown in greater quantit ...

grain, such as

rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...

, or

is a kind of fruit (termed a

caryopsis In botany, a caryopsis (plural caryopses) is a type of simple fruit—one that is monocarpellate (formed from a single carpel) and indehiscent (not opening at maturity) and resembles an achene, except that in a caryopsis the pericarp is fused ...

). However, the fruit wall is thin and fused to the seed coat, so almost all the edible grain-fruit is actually a seed.

Structure

Pomegranate fruit - whole and piece with arils

The outer layer, often edible, of most fruits is called the ''pericarp''. Typically formed from the ovary, it surrounds the seeds; in some species, however, other structural tissues contribute to or form the edible portion. The pericarp may be described in three layers from outer to inner, i.e., the ''epicarp'', ''mesocarp'' and ''endocarp''. Fruit that bears a prominent pointed terminal projection is said to be ''beaked''.

Development

A fruit results from the fertilizing and maturing of one or more flowers. The

gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

, which contains the '' stigma-style-ovary'' system, is centered in the flower-head, and it forms all or part of the fruit —(see graphic: 'the parts of a flower'). Inside the ovary(ies) are one or more

s. Here begins a complex sequence called ''

double fertilization Double fertilization is a complex fertilization mechanism of flowering plants (angiosperms). This process involves the joining of a female gametophyte ( megagametophyte, also called the embryo sac) with two male gametes (sperm). It begins when ...

'': a female

gametophyte A gametophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has on ...

produces an egg cell for the purpose of fertilization. (A female gametophyte is called a

megagametophyte A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the ...

, and also called the

embryo sac In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...

.) After

, the ovules will become seeds. Ovules are fertilized in a process that starts with

pollination Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds ...

, which is the movement of pollen from the stamens to the stigma-style-ovary system within the flower-head, (see graphic). After pollination, a

pollen tube A pollen tube is a tubular structure produced by the male gametophyte of seed plants when it germinates. Pollen tube elongation is an integral stage in the plant life cycle. The pollen tube acts as a conduit to transport the male gamete cells fro ...

grows from the (deposited) pollen through the stigma down the style into the ovary to the ovule. Two sperm are transferred from the pollen to a megagametophyte. Within the megagametophyte one sperm unites with the egg, forming a

zygote A zygote (, ) is a eukaryotic cell formed by a fertilization event between two gametes. The zygote's genome is a combination of the DNA in each gamete, and contains all of the genetic information of a new individual organism. In multicellula ...

, while the second sperm enters the central cell forming the endosperm mother cell, which completes the double fertilization process. Later the zygote will give rise to the embryo of the seed, and the endosperm mother cell will give rise to

endosperm The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the embryo and ...

, a nutritive tissue used by the embryo. As the ovules develop into seeds, the ovary begins to ripen and the ovary wall, the ''pericarp'', may become fleshy (as in berries or

drupe In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

s), or it may form a hard outer covering (as in nuts). In some multiseeded fruits, the extent to which a fleshy structure develops is proportional to the number of fertilized ovules. The pericarp typically is differentiated into two or three distinct layers; these are called the ''exocarp'' (outer layer, also called epicarp), ''mesocarp'' (middle layer), and ''endocarp'' (inner layer) —(see image of apple-section). In some fruits the

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s and/or

the style ''De Stijl'' (; ), Dutch for "The Style", also known as Neoplasticism, was a Dutch art movement founded in 1917 in Leiden. De Stijl consisted of artists and architects. In a more narrow sense, the term ''De Stijl'' is used to refer to a body ...

of the flower fall away as the fleshy fruit ripens. However, for simple fruits derived from an ''

inferior ovary In the flowering plants, an ovary is a part of the female reproductive organ of the flower or gynoecium. Specifically, it is the part of the pistil which holds the ovule(s) and is located above or below or at the point of connection with the bas ...

'' —i.e., one that lies ''below'' the attachment of other floral parts, (see graphic re 'insertion point')— there are parts (including petals, sepals, and stamens) that fuse with the ovary and ripen with it. For such a case, when floral parts other than the ovary form a significant part of the fruit that develops, it is called an

accessory fruit An accessory fruit is a fruit in which some of the flesh is derived not from the floral ovary but from some adjacent tissue exterior to the carpel.Esau, K. 1977. ''Anatomy of seed plants''. John Wiley and Sons, New York. Accessory fruits are us ...

. Examples of accessory fruits include apple, rose hip, strawberry, pineapple; see below, and "Table of fleshy fruit examples". Because several parts of the flower besides the ovary may contribute to the structure of a fruit, it is important to study flower structure to understand how a particular fruit forms. There are three general modes of fruit development: *

Apocarpous Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists of (one or m ...

fruits develop from a ''single flower'' (while having one or more separate, unfused, carpels); they are the simple fruits. *

Syncarpous Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

fruits develop from a ''single

'' (having two or more carpels fused together). * Multiple fruits form from many flowers —i.e., an inflorescence of flowers. File:Nectarine Fruit Development.jpg, The development sequence of a typical

, the

nectarine The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-fuz ...

(''Prunus persica'') over a 7.5 month period, from bud formation in early winter to fruit

ripening Ripening is a process in fruits that causes them to become more palatable. In general, fruit becomes sweeter, less green, and softer as it ripens. Even though the acidity of fruit increases as it ripens, the higher acidity level does not make the ...

in midsummer (see image page for further information) File:Mature flower diagram.svg, The parts of a flower, showing the stigma-style-ovary system. File:Pome apples text.jpg, An apple is a simple fleshy fruit. Key parts are the epicarp, or exocarp, or outer skin, (not labelled); and the mezocarp and endocarp (labelled). image:Ovary position.svg, upright 1.5, Insertion point: There are three positions of insertion of the ovary at the base of a flower: I superior; II half-inferior; III inferior. The 'insertion point' is where the

androecium The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...

parts ''(a)'', the petals ''(p)'', and the sepals ''(s)'' all converge and attach to the receptacle ''(r)''. (Ovary= gynoecium ''(g)''.) File:Noni fruit dev.jpg, upright 1.5, In the

noni ''Morinda citrifolia'' is a fruit-bearing tree in the coffee family, Rubiaceae. Its native range extends across Southeast Asia and Australasia, and was spread across the Pacific by Polynesian sailors. The species is now cultivated throughout th ...

, flowers are produced in time-sequence along the stem. It is possible to see a progression of flowering, fruit development, and fruit ripening.

Classification of fruits

Consistent with the three modes of fruit development plant scientists have classified fruits into three main groups: simple fruits, aggregate fruits, and multiple (or composite) fruits. The groupings reflect how the ovary and other flower organs are arranged and how the fruits develop, but they are not evolutionarily relevant as diverse plant

taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

may be in the same group. While the section of a

fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...

that produces

spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...

s is called a ''fruiting'' body, fungi are members of the

fungi kingdom A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from th ...

and not of the

plant kingdom Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude ...

Simple fruits

Simple fruits are the result of the ripening-to-fruit of a simple or compound ovary in a ''single flower'' with a ''single

pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

''. In contrast, a single flower with numerous pistils typically produces an

aggregate fruit A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...

; and the merging of several flowers, or a 'multiple' of flowers, results in a 'multiple' fruit. A simple fruit is further classified as to whether it is dry or fleshy. To distribute their seeds, dry fruits may split open and discharge their seeds to the winds, which is called dehiscence. Or the distribution process may rely upon the decay and degradation of the fruit to expose the seeds; or it may rely upon the eating of fruit and excreting of seeds by

frugivores A frugivore is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance and ...

—both are called indehiscence. Fleshy fruits do not split open, but they also are indehiscent and they may also rely on frugivores for distribution of their seeds. Typically, the entire outer layer of the ovary wall ripens into a potentially edible

pericarp Fruit anatomy is the plant anatomy of the internal structure of fruit. Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits. Aggre ...

. Types of dry simple fruits, (with examples) include: *

achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not ope ...

– most commonly seen in aggregate fruits (e.g.,

strawberry The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid species of the genus '' Fragaria'', collectively known as the strawberries, which are cultivated worldwide for their fruit. The fruit is widely ap ...

, see below). * capsule – (

Brazil nut The Brazil nut (''Bertholletia excelsa'') is a South American tree in the family Lecythidaceae, and it is also the name of the tree's commercially harvested edible seeds. It is one of the largest and longest-lived trees in the Amazon rainforest. ...

: botanically, it is not a nut). *

– (cereal grains, including

oats The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural, unlike other cereals and pseudocereals). While oats are suitable for human con ...

barley Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley pr ...

). * cypsela – an achene-like fruit derived from the individual florets in a

capitulum capitulum (plural capitula) may refer to: *the Latin word for chapter ** an index or list of chapters at the head of a gospel manuscript ** a short reading in the Liturgy of the Hours *** derived from which, it is the Latin for the assembly known ...

: (

dandelion ''Taraxacum'' () is a large genus of flowering plants in the family Asteraceae, which consists of species commonly known as dandelions. The scientific and hobby study of the genus is known as taraxacology. The genus is native to Eurasia and Nor ...

). * fibrous drupe – (

coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...

walnut A walnut is the edible seed of a drupe of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. Although culinarily considered a "nut" and used as such, it is not a true ...

: botanically, neither is a true nut.). * follicle – follicles are formed from a single carpel, and opens by one suture: (

milkweed ''Asclepias'' is a genus of herbaceous, perennial, flowering plants known as milkweeds, named for their latex, a milky substance containing cardiac glycosides termed cardenolides, exuded where cells are damaged. Most species are toxic to humans ...

); also commonly seen in aggregate fruits: (

magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendro ...

peony The peony or paeony is a flowering plant in the genus ''Paeonia'' , the only genus in the family Paeoniaceae . Peonies are native to Asia, Europe and Western North America. Scientists differ on the number of species that can be distinguished, ...

). *

legume A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock f ...

– (

: botanically, the peanut is the seed of a legume, not a nut). *

loment A loment (or lomentum) is a type of dehiscent legume fruit that breaks apart at constrictions occurring between segments, so that each segment contains one seed. It is a type of schizocarp. Tick trefoil (''Desmodium'') and sweet vetch (''Hedysarum ...

– a type of

indehiscent Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...

legume: ( sweet vetch or wild potato). *

– ( beechnut,

(of the

oak An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ''L ...

): botanically, these are true nuts). *

samara Samara ( rus, Сама́ра, p=sɐˈmarə), known from 1935 to 1991 as Kuybyshev (; ), is the largest city and administrative centre of Samara Oblast. The city is located at the confluence of the Volga and the Samara (Volga), Samara rivers, with ...

– (

ash Ash or ashes are the solid remnants of fires. Specifically, ''ash'' refers to all non-aqueous, non- gaseous residues that remain after something burns. In analytical chemistry, to analyse the mineral and metal content of chemical samples, ash ...

elm Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North ...

, maple key). *

schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...

, see below – ( carrot seed). *

silique A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit i ...

– (

radish The radish (''Raphanus raphanistrum'' subsp. ''sativus'') is an Eating, edible root vegetable of the family Brassicaceae that was domesticated in Asia prior to Roman Empire, Roman times. Radishes are grown and consumed throughout the world, be ...

seed). *

silicle A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit i ...

– (

shepherd's purse ''Capsella bursa-pastoris'', known as shepherd's purse because of its triangular flat fruits, which are purse-like, is a small annual and ruderal flowering plant in the mustard family (Brassicaceae). It is native to eastern Europe and Asia minor, ...

). * utricle – (

beet The beetroot is the taproot portion of a beet plant, usually known in North America as beets while the vegetable is referred to as beetroot in British English, and also known as the table beet, garden beet, red beet, dinner beet or golden beet ...

, ''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distribu ...

''). Fruits in which part or all of the ''pericarp'' (fruit wall) is fleshy at maturity are termed ''fleshy simple fruits''. Types of fleshy simple fruits, (with examples) include: *

berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...

– The berry is the most common type of fleshy fruit. The entire outer layer of the ovary wall ripens into a potentially edible "pericarp", (see below). *

stone fruit In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or ''pyrena'') of hardened endocarp with a seed (''kernel'') ...

– The definitive characteristic of a drupe is the hard, "lignified" stone (sometimes called the "pit"). It is derived from the ovary wall of the flower:

apricot An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus ''Prunus''. Usually, an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are also ...

cherry A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet ''Prunus avium'' and the sour ''Prunus cerasus''. The nam ...

olive The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' ...

peach The peach (''Prunus persica'') is a deciduous tree first domesticated and cultivated in Zhejiang province of Eastern China. It bears edible juicy fruits with various characteristics, most called peaches and others (the glossy-skinned, non-fu ...

plum A plum is a fruit of some species in ''Prunus'' subg. ''Prunus'.'' Dried plums are called prunes. History Plums may have been one of the first fruits domesticated by humans. Three of the most abundantly cultivated species are not found i ...

mango A mango is an edible stone fruit produced by the tropical tree ''Mangifera indica''. It is believed to have originated in the region between northwestern Myanmar, Bangladesh, and northeastern India. ''M. indica'' has been cultivated in South a ...

. *

pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince. Etymology The word ''pome'' entered English in the late 14th century, and re ...

– The

fruits:

pear Pears are fruits produced and consumed around the world, growing on a tree and harvested in the Northern Hemisphere in late summer into October. The pear tree and shrub are a species of genus ''Pyrus'' , in the family Rosaceae, bearing the p ...

rosehip The rose hip or rosehip, also called rose haw and rose hep, is the accessory fruit of the various species of rose plant. It is typically red to orange, but ranges from dark purple to black in some species. Rose hips begin to form after pollinat ...

saskatoon berry ''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to North America. Description It is a deciduous shrub or s ...

, et al., are a

syncarpous Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

(fused) fleshy fruit, a simple fruit, developing from a half-inferior ovary, see graphic re 'Insertion point'. Pomes are of the family

Rosaceae Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus ''Rosa''. Among the most species-rich genera are ''Alchemilla'' (270), ''Sorbus ...

Berries

Berries are a type of simple fleshy fruit that issue from a single ovary. (The ovary itself may be compound, with several carpels.) The botanical term "true berry" includes grapes, currants, cucumbers, eggplants (aubergines), tomatoes, chili peppers, and bananas, but excludes certain fruits that are called "-berry" by culinary custom or by common usage of the term —such as strawberries and raspberries. Berries may be formed from one or more carpels (i.e., from the simple or compound ovary) from the same, single flower. Seeds typically are embedded in the fleshy interior of the ovary. Examples here and in the table below: *

– In culinary terms, the tomato is regarded as a vegetable, but it is botanically classified as a fruit and a berry. *

– The fruit has been described as a "leathery berry". In cultivated varieties, the seeds are diminished nearly to non-existence. *

pepo PEPO Lappeenranta (abbreviated PEPO) is a football club from Lappeenranta in Finland. The club was formed in 1958 and their home ground is at the Kimpinen Sports Centre. The men's first team currently plays in the Ykkönen (Second Division). ...

– Berries with skin that is hardened:

cucurbits The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most important to humans are: *''Cucurbita'' – squash, pumpkin, zucchini, some gourds *''Lagena ...

, including gourds, squash, melons. *

hesperidium A hesperidium (plural hesperidia) is a modified berry with a tough, leathery rind. Etymology Carl Linnaeus gave the name Hesperideæ to an order containing the genus ''Citrus'', in allusion to the golden apples of the Hesperides. Development ...

– Berries with a rind and a juicy interior: most

citrus ''Citrus'' is a genus of flowering plant, flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as Orange (fruit), oranges, Lemon, lemons, grapefruits, pomelos, and lim ...

fruit. *

cranberry Cranberries are a group of evergreen dwarf shrubs or trailing vines in the subgenus ''Oxycoccus'' of the genus ''Vaccinium''. In Britain, cranberry may refer to the native species ''Vaccinium oxycoccos'', while in North America, cranberry ...

gooseberry Gooseberry ( or (American and northern British) or (southern British)) is a common name for many species of ''Ribes'' (which also includes currants), as well as a large number of plants of similar appearance. The berries of those in the genu ...

redcurrant The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions. Description ''Ribes r ...

. The ''strawberry'', regardless of its appearance, is classified as a dry, not a fleshy fruit. Botanically, it is ''not'' a

; it is an aggregate-accessory fruit, the latter term meaning the fleshy part is derived not from the plant's ovaries but from the receptacle that holds the

ovaries The ovary is an organ in the female reproductive system that produces an ovum. When released, this travels down the fallopian tube into the uterus, where it may become fertilized by a sperm. There is an ovary () found on each side of the body. T ...

.Esau, K. (1977). ''Anatomy of seed plants''. John Wiley and Sons, New York. Numerous dry

s are attached to the outside of the fruit-flesh, (see image); they appear to be seeds but each is actually an ovary of a flower, with a seed inside.

Schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...

s are dry fruits, though some appear to be fleshy. They originate from syncarpous ovaries but do not actually

dehisce Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...

; rather, they split into segments with one or more seeds. They include a number of different forms from a wide range of families, including

carrot The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in color, though purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild carrot, ''Daucus carota'', nat ...

parsnip The parsnip (''Pastinaca sativa'') is a root vegetable closely related to carrot and parsley, all belonging to the flowering plant family Apiaceae. It is a biennial plant usually grown as an annual. Its long taproot has cream-colored skin an ...

parsley Parsley, or garden parsley (''Petroselinum crispum'') is a species of flowering plant in the family Apiaceae that is native to the central and eastern Mediterranean region (Sardinia, Lebanon, Israel, Cyprus, Turkey, southern Italy, Greece, Por ...

cumin Cumin ( or , or Article title
) (''Cuminum cyminum'') is a

Aggregate fruits

Longitudinal section of raspberry flower

An aggregate fruit is also called an aggregation, or ''

etaerio A raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupe">raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggre ...

''; it develops from a single flower that presents numerous simple

s (see graphic of raspberry pistils). Each pistil contains one

; together they form a fruitlet. The ultimate (fruiting) development of the aggregation of pistils is called an aggregate fruit, etaerio fruit, or simply an etaerio. Different types of aggregate fruits can produce different etaerios, such as achenes, drupelets, follicles, and berries. For example, the Ranunculaceae species, including ''

Clematis ''Clematis'' is a genus of about 300 species within the buttercup family, Ranunculaceae. Their garden hybrids have been popular among gardeners, beginning with ''Clematis'' × ''jackmanii'', a garden standby since 1862; more hybrid cultivars ...

'' and ''

Ranunculus ''Ranunculus'' is a large genus of about almost 1700 to more than 1800 species of flowering plants in the family Ranunculaceae. Members of the genus are known as buttercups, spearworts and water crowfoots. The genus is distributed in Europe ...

'', produces an etaerio of

s; ''

Rubus ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with over 1,350 species. Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of these ...

'' species, including raspberry: an etaerio of

drupelet In botany, a drupe (or stone fruit) is an indehiscent fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pit'', ''stone'', or '' pyrena'') of hardened endocarp with a seed (''kernel'') ...

s; ''

Calotropis ''Calotropis'' is a genus of flowering plants in the family Apocynaceae, first described as a genus in 1810. It is native to southern Asia and North Africa. They are commonly known as milkweeds because of the latex they produce. ''Calotropis'' ...

'' species: an etaerio of follicles fruit; ''

Annona ''Annona'' (from Taíno ''annon'') is a genus of flowering plants in the pawpaw/sugar apple family, Annonaceae. It is the second largest genus in the family after ''Guatteria'', containing approximately 166

'' species: an etaerio of

berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...

. Some other broadly recognized species and their etaerios (or aggregations) are: *

Teasel ''Dipsacus'' is a genus of flowering plant in the family Caprifoliaceae. The members of this genus are known as teasel, teazel or teazle. The genus includes about 15 species of tall herbaceous biennial plants (rarely short-lived perennial plants ...

; fruit is an aggregation of cypselas. *

Tuliptree ''Liriodendron'' () is a genus of two species of characteristically large trees, deciduous over most of their populations, in the magnolia family (Magnoliaceae). These trees are widely known by the common name tulip tree or tuliptree for their ...

; fruit is an aggregation of samaras. *

Magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendro ...

and

; fruit is an aggregation of follicles. *

American sweet gum American sweetgum (''Liquidambar styraciflua''), also known as American storax, hazel pine, bilsted, redgum, satin-walnut, star-leaved gum, alligatorwood, or simply sweetgum, is a deciduous tree in the genus ''Liquidambar'' native to warm temper ...

; fruit is an aggregation of capsules. *

Sycamore Sycamore is a name which has been applied to several types of trees, but with somewhat similar leaf forms. The name derives from the ancient Greek ' (''sūkomoros'') meaning "fig-mulberry". Species of trees known as sycamore: * ''Acer pseudoplata ...

; fruit is an aggregation of achenes. The ''

raspberry The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with w ...

''; its pistils are called ''drupelets'' because each pistil is like a small

attached to the receptacle. In some

bramble A bramble is any rough, tangled, prickly shrub, usually in the genus ''Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits inclu ...

fruits such as

blackberry The blackberry is an edible fruit produced by many species in the genus ''Rubus'' in the family Rosaceae, hybrids among these species within the subgenus ''Rubus'', and hybrids between the subgenera ''Rubus'' and ''Idaeobatus''. The taxonomy of ...

the receptacle, an accessory part, elongates and then develops as part of the fruit, making the blackberry an '' aggregate-accessory'' fruit. The

is also an aggregate-accessory fruit, of which the seeds are contained in the

s. Notably in all these examples, the fruit develops from a single flower, with numerous pistils.

Multiple fruits

A multiple fruit is formed from a cluster of flowers, (a 'multiple' of flowers) —also called an ''

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...

''. Each ('smallish') flower produces a single fruitlet, which, as all develop, all merge into one mass of fruit. Examples include

pineapple The pineapple (''Ananas comosus'') is a tropical plant with an edible fruit; it is the most economically significant plant in the family Bromeliaceae. The pineapple is indigenous to South America, where it has been cultivated for many centuri ...

fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...

mulberry ''Morus'', a genus of flowering plants in the family Moraceae, consists of diverse species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 identif ...

Osage orange ''Maclura pomifera'', commonly known as the Osage orange ( ), is a small deciduous tree or large shrub, native to the south-central United States. It typically grows about tall. The distinctive fruit, a multiple fruit, is roughly spherical, b ...

breadfruit Breadfruit (''Artocarpus altilis'') is a species of flowering tree in the mulberry and jackfruit family (Moraceae) believed to be a domesticated descendant of ''Artocarpus camansi'' originating in New Guinea, the Maluku Islands, and the Philippi ...

. An inflorescence (a cluster) of white flowers, called a head, is produced first. After

fertilization Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Proce ...

, each flower in the cluster develops into a drupe; as the drupes expand, they develop as a '' connate'' organ, merging into a multiple fleshy fruit called a ''syncarp''. Progressive stages of multiple flowering and fruit development can be observed on a single branch of the Indian mulberry, or ''

'', (see image). During the sequence of development, a progression of second, third, and more inflorescences are initiated in turn at the head of the branch or stem.

Accessory fruit forms

Fruits may incorporate tissues derived from other floral parts besides the ovary, including the receptacle, hypanthium, petals, or sepals. Accessory fruits occur in all three classes of fruit development—simple, aggregate, and multiple. Accessory fruits are frequently designated by the hyphenated term showing both characters. For example, a pineapple is a multiple-accessory fruit. a blackberry is an aggregate-accessory fruit, and an apple is a simple-accessory fruit.

Table of fleshy fruit examples

Seedless fruits

Seedlessness is an important feature of some fruits of commerce. Commercial

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

s of

s and

s are examples of

seedless fruit A seedless fruit is a fruit developed to possess no mature seeds. Since eating seedless fruits is generally easier and more convenient, they are considered commercially valuable. Most commercially produced seedless fruits have been developed from ...

s. Some cultivars of

fruits (especially

grapefruit The grapefruit (''Citrus'' × ''paradisi'') is a subtropical citrus tree known for its relatively large, sour to semi-sweet, somewhat bitter fruit. The interior flesh is segmented and varies in color from pale yellow to dark pink. Grapefruit is ...

mandarin orange The mandarin orange (''Citrus reticulata''), also known as the mandarin or mandarine, is a small citrus tree fruit. Treated as a distinct species of orange, it is usually eaten plain or in fruit salads. Tangerines are a group of orange-coloured ...

s, navel

), satsumas,

table grape Table grapes are grapes intended for consumption while fresh, as opposed to grapes grown for wine production, juice production, or for drying into raisins. ''Vitis vinifera'' table grapes can be in the form of either seeded or non-seeded varieta ...

s, and of

watermelon Watermelon (''Citrullus lanatus'') is a flowering plant species of the Cucurbitaceae family and the name of its edible fruit. A scrambling and trailing vine-like plant, it is a highly cultivated fruit worldwide, with more than 1,000 varieti ...

s are valued for their seedlessness. In some species, seedlessness is the result of ''

parthenocarpy In botany and horticulture, parthenocarpy is the natural or artificially induced production of fruit without fertilisation of ovules, which makes the fruit seedless. Stenospermocarpy may also produce apparently seedless fruit, but the seeds are ac ...

'', where fruits set without fertilization. Parthenocarpic fruit-set may (or may not) require pollination, but most seedless citrus fruits require a stimulus from pollination to produce fruit. Seedless bananas and grapes are

triploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei ( eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contain ...

s, and seedlessness results from the abortion of the

embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male spe ...

nic plant that is produced by fertilization, a phenomenon known as ''

stenospermocarpy Stenospermocarpy is the biological mechanism that produces parthenocarpy (seedlessness) in some fruits, notably many table grapes. In stenospermocarpic fruits, normal pollination and fertilization are still required to ensure that the fruit 'sets', ...

'', which requires normal pollination and fertilization.

Seed dissemination

Variations in fruit structures largely depend on the modes of dispersal applied to their seeds. Dispersal is achieved by wind or water, by

explosive dehiscence Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part; structures that op ...

, and by interactions with animals. Some fruits present their outer skins or shells coated with spikes or hooked burrs; these evolved either to deter would-be foragers from feeding on them, or to serve to attach themselves to the hair, feathers, legs, or clothing of animals, thereby using them as dispersal agents. These plants are termed

zoochorous In Spermatophyte plants, seed dispersal is the movement, spread or transport of seeds away from the parent plant. Plants have limited mobility and rely upon a variety of dispersal vectors to transport their seeds, including both abiotic vectors, ...

; common examples include

cocklebur ''Xanthium'' (cocklebur) is a genus of flowering plants in the tribe Heliantheae within the family Asteraceae, native to the Americas and eastern Asia and some parts of south Asia . Description Cockleburs are coarse, herbaceous annual plants ...

unicorn plant ''Proboscidea'' is a genus of flowering plant in the family Martyniaceae, some of whose species are known as devil's claw, devil's horn, ram's horn, or unicorn plant. The plants produce long, hooked seed pods. The hooks catch on the feet of anim ...

, and beggarticks (or Spanish needle). By developments of mutual evolution the fleshy produce of fruits typically appeals to hungry animals, such that the seeds contained within are taken in, carried away and later deposited (i.e.,

defecated Defecation (or defaecation) follows digestion, and is a necessary process by which organisms eliminate a solid, semisolid, or liquid waste material known as feces from the digestive tract via the anus. The act has a variety of names ranging f ...

) at a distance from the parent plant. Likewise, the nutritious, oily kernels of nuts typically motivate birds and

squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. Squ ...

s to

hoard A hoard or "wealth deposit" is an archaeological term for a collection of valuable objects or artifacts, sometimes purposely buried in the ground, in which case it is sometimes also known as a cache. This would usually be with the intention of ...

them, burying them in soil to retrieve later during the winter of scarcity; thereby, uneaten seeds are sown effectively under natural conditions to

germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...

and grow a new plant some distance away from the parent. Other fruits have evolved flattened and elongated wings or helicopter-like blades, e.g.,

maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since http ...

, and

tuliptree ''Liriodendron'' () is a genus of two species of characteristically large trees, deciduous over most of their populations, in the magnolia family (Magnoliaceae). These trees are widely known by the common name tulip tree or tuliptree for their ...

. This mechanism increases dispersal distance away from the parent via wind. Other wind-dispersed fruit have tiny "

parachutes A parachute is a device used to slow the motion of an object through an atmosphere by creating drag or, in a ram-air parachute, aerodynamic lift. A major application is to support people, for recreation or as a safety device for aviators, who ...

", e.g.,

, salsify.

Coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...

fruits can float thousands of miles in the ocean, thereby spreading their seeds. Other fruits that can disperse via water are

nipa palm ''Nypa fruticans'', commonly known as the nipa palm (or simply nipa, from ms, nipah) or mangrove palm, is a species of palm native to the coastlines and estuarine habitats of the Indian and Pacific Oceans. It is the only palm considered adapte ...

and

screw pine ''Pandanus'' is a genus of monocots with some 750 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. The greatest number of species are found in Madagascar and Malaysia. Common nam ...

. Some fruits have evolved propulsive mechanisms that fling seeds substantial distances —(perhaps up to 100 m in the case of the

sandbox tree ''Hura crepitans'', the sandbox tree, also known as possumwood and jabillo, is an evergreen tree of the spurge family (Euphorbiaceae), native to tropical regions of North and South America including the Amazon rainforest. It is also present in p ...

)— via

or other such mechanisms, (see

impatiens ''Impatiens'' is a genus of more than 1,000 species of flowering plants, widely distributed throughout the Northern Hemisphere and the tropics. Together with the genus ''Hydrocera'' (one species), ''Impatiens'' make up the family Balsaminaceae. ...

and

squirting cucumber ''Ecballium'' is a genus of flowering plants in the family Cucurbitaceae containing a single species, ''Ecballium elaterium'', also called the squirting cucumber or exploding cucumber (not the same plant as ''Cyclanthera brachystachya''). Its u ...

Food uses

cornucopia In classical antiquity, the cornucopia (), from Latin ''cornu'' (horn) and ''copia'' (abundance), also called the horn of plenty, was a symbol of abundance and nourishment, commonly a large horn-shaped container overflowing with produce, flowers ...

of fruits —fleshy (simple) fruits from apples to berries to watermelon; dry (simple) fruits including beans and rice, coconuts and carrots; aggregate fruits including strawberries, raspberries, blackberries, pawpaw; multiple fruits such as pineapple, fig, mulberries; (see above re all)— are commercially valuable as human food. They are eaten both fresh and as jams, marmalade and other

fruit preserves Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the method ...

. They are used extensively in manufactured and processed foods (cakes, cookies, baked goods, flavorings, ice cream, yogurt, canned vegetables, frozen vegetables and meals) and beverages such as fruit juices and

alcoholic beverages An alcoholic beverage (also called an alcoholic drink, adult beverage, or a drink) is a drink that contains ethanol, a type of alcohol that acts as a drug and is produced by fermentation of grains, fruits, or other sources of sugar. The cons ...

(brandy,

fruit beer Fruit beer is beer made with fruit added as an adjunct or flavouring. Fruit beer is a beer made from malt-scented strawberries, plums, raspberries, and cherries. It's an additive- or flavoring-containing form. Initially manufactured in Belgium, f ...

, wine). Spices like vanilla, black pepper, paprika, and

are derived from berries. Olive fruit is pressed for

olive oil Olive oil is a liquid fat obtained from olives (the fruit of ''Olea europaea''; family Oleaceae), a traditional tree crop of the Mediterranean Basin, produced by pressing whole olives and extracting the oil. It is commonly used in cooking: f ...

and similar processing is applied to other oil bearing fruits/vegetables. Fruits are also used for socializing and gift-giving in the form of

fruit basket A gift basket or fruit basket is typically a gift delivered to the recipient at their home or workplace. A variety of gift baskets exist: some contain fruit; while others might contain dry or canned foods such as tea, Cracker (food), crackers an ...

s and

fruit bouquet A gift basket or fruit basket is typically a gift delivered to the recipient at their home or workplace. A variety of gift baskets exist: some contain fruit; while others might contain dry or canned foods such as tea, Cracker (food), crackers an ...

s. Typically, many botanical fruits —"vegetables" in culinary ''parlance''— (including tomato, green beans, leaf greens, bell pepper, cucumber, eggplant, okra, pumpkin, squash, zucchini) are bought and sold daily in fresh produce markets and

greengrocer A greengrocer is a person who owns or operates a shop selling primarily fruit and vegetables. The term may also be used to refer to a shop selling primarily produce. It is used predominantly in the United Kingdom and Australia. In the United ...

ies and carried back to kitchens, at home or restaurant, for preparation of meals.

Storage

All fruits benefit from proper post harvest care, and in many fruits, the plant hormone

ethylene Ethylene (IUPAC name: ethene) is a hydrocarbon which has the formula or . It is a colourless, flammable gas with a faint "sweet and musky" odour when pure. It is the simplest alkene (a hydrocarbon with carbon-carbon double bonds). Ethylene i ...

causes

. Therefore, maintaining most fruits in an efficient

cold chain A cold chain is a low temperature-controlled supply chain network. An unbroken cold chain is an uninterrupted series of refrigerated production, storage and distribution activities, along with associated equipment and logistics, which maintain qu ...

is optimal for post harvest storage, with the aim of extending and ensuring shelf life.Why Cold Chain for Fruits:

Nutritional value

Various culinary fruits provide significant amounts of

fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...

and water, and many are generally high in

vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits and vegetables, also sold as a dietary supplement and as a topical 'serum' ingredient to treat melasma (dark pigment spots) an ...

. An overview of numerous studies showed that fruits (e.g., whole apples or whole oranges) are satisfying (filling) by simply eating and chewing them. The

dietary fiber Dietary fiber (in British English fibre) or roughage is the portion of plant-derived food that cannot be completely broken down by human digestive enzymes. Dietary fibers are diverse in chemical composition, and can be grouped generally by the ...

consumed in eating fruit promotes

satiety Satiety ( ) is a state or condition of fullness gratified beyond the point of satisfaction, the opposite of hunger. It is a state which induces meal termination.Hetherington, M.Sensory-specific satiety and its importance in meal termination ''Neuro ...

, and may help to control body weight and aid reduction of blood

cholesterol Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol (or modified steroid), a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell mem ...

, a

risk factor In epidemiology, a risk factor or determinant is a variable associated with an increased risk of disease or infection. Due to a lack of harmonization across disciplines, determinant, in its more widely accepted scientific meaning, is often use ...

for

cardiovascular diseases Cardiovascular disease (CVD) is a class of diseases that involve the heart or blood vessels. CVD includes coronary artery diseases (CAD) such as angina and myocardial infarction (commonly known as a heart attack). Other CVDs include stroke, ...

. Fruit consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. Regular consumption of fruit is generally associated with reduced risks of several diseases and functional declines associated with aging.

Food safety

For

food safety Food safety (or food hygiene) is used as a scientific method/discipline describing handling, preparation, and storage of food in ways that prevent food-borne illness. The occurrence of two or more cases of a similar illness resulting from t ...

, the

CDC The Centers for Disease Control and Prevention (CDC) is the National public health institutes, national public health agency of the United States. It is a Federal agencies of the United States, United States federal agency, under the United S ...

recommends proper fruit handling and preparation to reduce the risk of

food contamination Food contamination refers to the presence of harmful chemicals and microorganisms in food, which can cause consumer illness. This article addresses the chemical contamination of foods, as opposed to microbiological contamination, which can be found ...

and

foodborne illness Foodborne illness (also foodborne disease and food poisoning) is any illness resulting from the spoilage of contaminated food by pathogenic bacteria, viruses, or parasites that contaminate food, as well as prions (the agents of mad cow disease) ...

. Fresh fruits and vegetables should be carefully selected; at the store, they should not be damaged or bruised; and precut pieces should be refrigerated or surrounded by ice. All fruits and vegetables should be rinsed before eating. This recommendation also applies to produce with rinds or skins that are not eaten. It should be done just before preparing or eating to avoid premature spoilage. Fruits and vegetables should be kept separate from raw foods like meat, poultry, and seafood, as well as from utensils that have come in contact with raw foods. Fruits and vegetables that are not going to be cooked should be thrown away if they have touched raw meat, poultry, seafood, or eggs. All cut, peeled, or cooked fruits and vegetables should be refrigerated within two hours. After a certain time, harmful bacteria may grow on them and increase the risk of foodborne illness.

Allergies

Fruit allergies make up about 10 percent of all food related allergies.

Nonfood uses

Because fruits have been such a major part of the human diet, various cultures have developed many different uses for fruits they do not depend on for food. For example: PorcelainBerry

Bayberry ''Myrica'' is a genus of about 35–50 species of small trees and shrubs in the family Myricaceae, order Fagales. The genus has a wide distribution, including Africa, Asia, Europe, North America and South America, and missing only from Aust ...

fruits provide a wax often used to make candles; * Many dry fruits are used as decorations or in dried flower arrangements (e.g.,

annual honesty ''Lunaria annua'', called honesty or annual honesty in English, is a species of flowering plant in the cabbage and mustard family Brassicaceae. It is native to the Balkans and south west Asia, and cultivated throughout the temperate world. Desc ...

cotoneaster ''Cotoneaster'' is a genus of flowering plants in the rose family, Rosaceae, native to the Palaearctic region (temperate Asia, Europe, north Africa), with a strong concentration of diversity in the genus in the mountains of southwestern China an ...

lotus Lotus may refer to: Plants *Lotus (plant), various botanical taxa commonly known as lotus, particularly: ** ''Lotus'' (genus), a genus of terrestrial plants in the family Fabaceae **Lotus flower, a symbolically important aquatic Asian plant also ...

, and

Ornamental tree Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

s and shrubs are often cultivated for their colorful fruits, including

beautyberry ''Callicarpa'' (beautyberry) is a genus of shrubs and small trees in the family Lamiaceae.Heywood, V.H., Brummitt, R.K., Culham, A. & Seberg, O. 2007: Flowering Plant Families of the World. Royal Botanic Gardens, Kew. They are native to east a ...

holly ''Ilex'' (), or holly, is a genus of over 570 species of flowering plants in the family Aquifoliaceae, and the only living genus in that family. ''Ilex'' has the most species of any woody dioecious angiosperm genus. The species are evergreen o ...

pyracantha ''Pyracantha'' (from Greek "fire" and "thorn", hence firethorn) is a genus of large, thorny evergreen shrubs in the family Rosaceae, with common names firethorn or pyracantha. They are native to an area extending from Southwest Europe east to ...

skimmia ''Skimmia'' is a genus of four species of evergreen shrubs and small trees in the rue family Rutaceae, all native to warm temperate regions of Asia. The leaves are clustered at the ends of the shoots, simple, lanceolate, 6–21 cm long and ...

, and

viburnum ''Viburnum'' is a genus of about 150–175 species of flowering plants in the moschatel family Adoxaceae. Its current classification is based on molecular phylogeny. It was previously included in the honeysuckle family Caprifoliaceae. The membe ...

. * Fruits of

opium poppy ''Papaver somniferum'', commonly known as the opium poppy or breadseed poppy, is a species of flowering plant in the family Papaveraceae. It is the species of plant from which both opium and poppy seeds are derived and is also a valuable ornamen ...

are the source of

opium Opium (or poppy tears, scientific name: ''Lachryma papaveris'') is dried latex obtained from the seed capsules of the opium poppy ''Papaver somniferum''. Approximately 12 percent of opium is made up of the analgesic alkaloid morphine, which i ...

, which contains the drugs

codeine Codeine is an opiate and prodrug of morphine mainly used to treat pain, coughing, and diarrhea. It is also commonly used as a recreational drug. It is found naturally in the sap of the opium poppy, ''Papaver somniferum''. It is typically use ...

and

morphine Morphine is a strong opiate that is found naturally in opium, a dark brown resin in poppies (''Papaver somniferum''). It is mainly used as a analgesic, pain medication, and is also commonly used recreational drug, recreationally, or to make ...

, as well as the biologically inactive chemical theabaine from which the drug

oxycodone Oxycodone, sold under various brand names such as Roxicodone and OxyContin (which is the extended release form), is a strong, semi-synthetic opioid used medically for treatment of moderate to severe pain. It is highly addictive and a commonly ...

is synthesized. *

fruits are used to repel

cockroach Cockroaches (or roaches) are a paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are well-known as ...

es. * Many fruits provide

natural dye Natural dyes are dyes or colorants derived from plants, invertebrates, or minerals. The majority of natural dyes are vegetable dyes from plant sources—roots, berries, bark, leaves, and wood—and other biological sources such as fungi. Archaeol ...

s (e.g.,

sumac Sumac ( or ), also spelled sumach, is any of about 35 species of flowering plants in the genus ''Rhus'' and related genera in the cashew family (Anacardiaceae). Sumacs grow in subtropical and temperate regions throughout the world, including Eas ...

, and

). * Dried

gourd Gourds include the fruits of some flowering plant species in the family Cucurbitaceae, particularly ''Cucurbita'' and ''Lagenaria''. The term refers to a number of species and subspecies, many with hard shells, and some without. One of the earli ...

s are used as bird houses, cups, decorations, dishes, musical instruments, and water jugs. *

Pumpkin A pumpkin is a vernacular term for mature winter squash of species and varieties in the genus ''Cucurbita'' that has culinary and cultural significance but no agreed upon botanical or scientific meaning. The term ''pumpkin'' is sometimes use ...

s are carved into

Jack-o'-lantern A jack-o'-lantern (or jack o'lantern) is a carved lantern, most commonly made from a pumpkin or a root vegetable such as a rutabaga or turnip. Jack-o'-lanterns are associated with the Halloween holiday. Its name comes from the reported phenomen ...

s for

Halloween Halloween or Hallowe'en (less commonly known as Allhalloween, All Hallows' Eve, or All Saints' Eve) is a celebration observed in many countries on 31 October, the eve of the Western Christian feast of All Saints' Day. It begins the observanc ...

. * The fibrous core of the mature and dry

Luffa ''Luffa'' is a genus of tropical and subtropical vines in the cucumber family (Cucurbitaceae). In everyday non-technical usage, the luffa, also spelled loofah, usually refers to the fruits of the species ''Luffa aegyptiaca'' and ''Luffa acutang ...

fruit is used as a sponge. * The spiny fruit of

burdock ''Arctium'' is a genus of biennial plants commonly known as burdock, family Asteraceae. Native to Europe and Asia, several species have been widely introduced worldwide. Burdock's clinging properties, in addition to providing an excellent mecha ...

inspired the invention of

Velcro Velcro, officially known as Velcro IP Holdings LLC and trading as Velcro Companies, is a British privately held company, founded by Swiss electrical engineer George de Mestral in the 1950s. It is the original manufacturer of hook-and-loop fasten ...

. *

Coir Coir (), also called coconut fibre, is a natural fibre extracted from the outer husk of coconut and used in products such as floor mats, doormats, brushes, and mattresses. Coir is the fibrous material found between the hard, internal shell ...

fiber from

shells is used for brushes, doormats, floor tiles, insulation, mattresses, sacking, and as a growing medium for container plants. The shell of the coconut fruit is used to make bird houses, bowls, cups, musical instruments, and souvenir heads. * The hard and colorful grain fruits of

Job's tears Job's tears (''Coix lacryma-jobi)'', also known as Adlay or Adlay millet, is a tall grain-bearing perennial tropical plant of the family Poaceae (grass family). It is native to Southeast Asia and introduced to Northern China and India in remote ...

are used as decorative beads for jewelry, garments, and ritual objects. * Fruit is often a subject of

still life A still life (plural: still lifes) is a work of art depicting mostly wikt:inanimate, inanimate subject matter, typically commonplace objects which are either natural (food, flowers, dead animals, plants, rocks, shells, etc.) or artificiality, m ...

paintings.

References

External links

Images of fruit development from flowers
at bioimages.Vanderbilt.edu

at bioimages.Vanderbilt.edu

from California Rare Fruit Growers, Inc.
Photo ID of Fruits
by Capt. Pawanexh Kohli * {{Authority control Fruit Pollination Types of food

Botanical vs. culinary

Structure

Development

Classification of fruits

Simple fruits

Berries

Aggregate fruits

Multiple fruits

Accessory fruit forms

Table of fleshy fruit examples

Seedless fruits

Seed dissemination

Food uses

Storage

Nutritional value

Food safety

Allergies

Nonfood uses

See also

References

Further reading

External links