The Fagales are an

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s, including some of the best-known

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

s. The order name is derived from

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Fagus'',

beech Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...

es. They belong among the rosid group of

dicotyledon The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, t ...

s. The families and genera currently included are as follows: * Betulaceae – birch family (''

Alnus Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few spec ...

'', '' Betula'', ''

Carpinus Hornbeams are hardwood trees in the flowering plant genus ''Carpinus'' in the birch family Betulaceae. The 30–40 species occur across much of the temperate regions of the Northern Hemisphere. Origin of names The common English name ''hornbeam' ...

'', '' Corylus'', '' Ostrya'', ''

Ostryopsis ''Ostryopsis'' is a small genus of deciduous shrubs belonging to the birch family Betulaceae. The species have no common English name, though hazel-hornbeam has been suggested, reflecting their similarities to the closely related hazels and h ...

'') *

Casuarinaceae The Casuarinaceae are a family of dicotyledonous flowering plants placed in the order Fagales, consisting of four genera and 91 species of trees and shrubs native to eastern Africa, Australia, Southeast Asia, Malesia, Papuasia, and the Pacifi ...

– she-oak family (''

Allocasuarina ''Allocasuarina'' is a genus of trees in the flowering plant family Casuarinaceae. They are endemic to Australia, occurring primarily in the south. Like the closely related genus ''Casuarina'', they are commonly called sheoaks or she-oaks. Wi ...

'', ''

Casuarina ''Casuarina'' is a genus of 17 tree species in the family Casuarinaceae, native to Australia, the Indian subcontinent, southeast Asia, islands of the western Pacific Ocean, and eastern Africa. It was once treated as the sole genus in the fami ...

'', '' Ceuthostoma'', '' Gymnostoma'') *

Fagaceae The Fagaceae are a family of flowering plants that includes beeches, chestnuts and oaks, and comprises eight genera with about 927 species. Fagaceae in temperate regions are mostly deciduous, whereas in the tropics, many species occur as evergre ...

– beech family (''

Castanea The chestnuts are the deciduous trees and shrubs in the genus ''Castanea'', in the beech family Fagaceae. They are native to temperate regions of the Northern Hemisphere. The name also refers to the edible nut (fruit), nuts they produce. Th ...

'', '' Castanopsis'', ''

Chrysolepis ''Chrysolepis'' is a small genus of plants in the family Fagaceae, endemic to the western United States. Its two species have the common name chinquapin. The genus occurs from western Washington south to the Transverse Ranges in Southern Califor ...

'', ''

Colombobalanus ''Trigonobalanus'' is a genus of three species of evergreen trees in the family Fagaceae, related to oaks, beeches and chestnuts. The species are widely scattered, with one in northern South America and two in southeast Asia; some botanists t ...

'', '' Fagus'', ''

Lithocarpus ''Lithocarpus'' is a genus in the beech family, Fagaceae. Trees in this genus are commonly known as the stone oaks and differ from ''Quercus'' primarily because they produce insect-pollinated flowers on erect spikes and the female flowers have ...

'', '' Notholithocarpus'', ''

Quercus An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably '' ...

'') * Juglandaceae – walnut family (''

Alfaroa ''Alfaroa'' is a genus of evergreen trees in the Juglandaceae family of the Fagales, growing in montane and submontane tropical rain forests in Central America. The wood is characterized by solid pith, pink heartwood, and vessels with scalarifor ...

'', '' Carya'', ''

Cyclocarya ''Cyclocarya'' is a genus of flowering plants in the family Juglandaceae, comprising a single species ''Cyclocarya paliurus'' (qing qian liu or wheel wingnut), formerly treated in the genus ''Pterocarya'' as ''Pterocarya paliurus''. It is native ...

'', ''

Engelhardia ''Engelhardia'' is a genus of seven species of trees in the family Juglandaceae, native to southeast Asia from northern India east to Taiwan, Indonesia and the Philippines. The genus name is commonly misspelled "''Engelhardtia''", a "correctio ...

'', '' Juglans'', ''

Oreomunnea ''Oreomunnea'' is a genus of two species of flowering plants in the family Juglandaceae, native to southern Mexico and Central America, where they occur in montane rainforest.Manning, W. E. (1979). The Classification within the Juglandaceae. ''An ...

'', ''

Platycarya ''Platycarya'' is a genus of flowering plants in the family Juglandaceae native to eastern Asia in China, Korea, and Japan.Flora of China''Platycarya''/ref>Flora of China''Platycarya strobilacea''/ref> It is a deciduous tree growing to 15 m tall ...

'', '' Pterocarya'', ''

Rhoiptelea ''Rhoiptelea'' is a monotypic genus of flowering plants in the family Juglandaceae. It contains a single species, ''Rhoiptelea chiliantha'', commonly known as the horsetail tree. This genus was previously recognized in its own family, Rhoipteleac ...

'') * Myricaceae – bayberry family ('' Canacomyrica'', '' Comptonia'', '' Myrica'') * Nothofagaceae – southern beech family ('' Nothofagus'') *

Ticodendraceae ''Ticodendron incognitum'' is the only species of ''Ticodendron'', and the only member of the family Ticodendraceae. It is most closely related to the family Betulaceae. It was discovered only in 1989 in Costa Rica, having been overlooked previ ...

– ticodendron family (''

Ticodendron ''Ticodendron incognitum'' is the only species of ''Ticodendron'', and the only member of the family Ticodendraceae. It is most closely related to the family Betulaceae. It was discovered only in 1989 in Costa Rica, having been overlooked previo ...

'') The older Cronquist system only included four families (Betulaceae, Corylaceae, Fagaceae, Ticodendraceae; Corylaceae now being included within Betulaceae); this arrangement is followed by, for example, the World Checklist of selected plant families. The other families were split into three different orders, placed among the

Hamamelidae Hamamelididae is an obsolete botanical name at the rank of subclass. Because some hamamelidid members bear aments (''i.e.'', catkins), this subclass has been formerly known as ''Amentiferae''. Based on molecular phylogeny works, Hamamelididae app ...

. The Casuarinales comprised the single family Casuarinaceae, the Juglandales comprised the Juglandaceae and Rhoipteleaceae, and the Myricales comprised the remaining forms (plus '' Balanops''). The change is due to studies suggesting the Myricales, so defined, are

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

to the other two groups.

Characteristics

Most Fagales are wind pollinated and are

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...

with unisexual flowers.

Evolutionary history

The oldest member of the order is the flower '' Soepadmoa cupulata'' preserved in the late

Turonian The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by t ...

Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...

New Jersey amber New Jersey Amber, sometimes called Raritan amber, is amber found in the Raritan and Magothy Formations of the Central Atlantic (Eastern) coast of the United States. It is dated to the Late Cretaceous, Turonian age, based on pollen analysis of th ...

, which is a

mosaic A mosaic is a pattern or image made of small regular or irregular pieces of colored stone, glass or ceramic, held in place by plaster/mortar, and covering a surface. Mosaics are often used as floor and wall decoration, and were particularly pop ...

with characteristics characteristic of both ''Nothofagus'' and other Fagales, suggesting that the ancestor of all Fagales was ''Nothofagus''-like.

Systematics

Modern

molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

suggest the following relationships:

References

External links

Missouri Botanical Gardens - Fagales
{{Taxonbar, from=Q21881 Angiosperm orders