The coyote (''Canis latrans''), also known as the American jackal, prairie wolf, or brush wolf, is a
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. It is smaller than its close relative, the
gray wolf
The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...
, and slightly smaller than the closely related
eastern wolf
The eastern wolf (''Canis lycaon'' or ''Canis lupus lycaon''), also known as the timber wolf, Algonquin wolf and eastern timber wolf, is a canine of debated taxonomy native to the Great Lakes region and southeastern Canada. It is considered eith ...
and
red wolf
The red wolf (''Canis rufus'') is a Canis, canine native to the southeastern United States. Its size is intermediate between the coyote (''Canis latrans'') and Wolf, gray wolf (''Canis lupus'').
The red wolf's taxonomic classification as being ...
. It fills much of the same
ecological niche
In ecology, a niche is the match of a species to a specific environmental condition.
Three variants of ecological niche are described by
It describes how an organism or population responds to the distribution of Resource (biology), resources an ...
as the
golden jackal
The golden jackal (''Canis aureus''), also called the common jackal, is a wolf-like canid that is native to Eurasia. The golden jackal's coat varies in color from a pale creamy yellow in summer to a dark tawny beige in winter. It is smaller a ...
does in
Eurasia
Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
; however, the coyote is generally larger.
The coyote is listed as
least concern
A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the
International Union for Conservation of Nature
The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
, due to its wide distribution and abundance throughout North America. The species is versatile, able to adapt to and expand into environments modified by humans; urban coyotes are common in many cities. The coyote was sighted in eastern
Panama
Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
(across the
Panama Canal
The Panama Canal () is an artificial waterway in Panama that connects the Caribbean Sea with the Pacific Ocean. It cuts across the narrowest point of the Isthmus of Panama, and is a Channel (geography), conduit for maritime trade between th ...
from their home range) for the first time in 2013.
The coyote has 19 recognized
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
. The average male weighs and the average female . Their fur color is predominantly light gray and red or
fulvous
Fulvous is a colour, sometimes described as dull orange, brownish-yellow or tawny; it can also be likened to a variation of buff, beige or butterscotch. As an adjective it is used in the names of many species of birds, and occasionally other ...
interspersed with black and white, though it varies somewhat with geography. It is highly flexible in social organization, living either in a family unit or in loosely knit packs of unrelated individuals. Primarily
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
, its diet consists mainly of
deer
A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
,
rabbit
Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s,
hare
Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
s,
rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s,
bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s,
reptile
Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s,
amphibian
Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s,
fish
A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
, and
invertebrate
Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s, though it may also eat fruits and vegetables on occasion. Its characteristic vocalization is a
howl
Howl most often refers to:
* Howling, an animal vocalization in many canine species
* "Howl" (poem), a 1956 poem by Allen Ginsberg
Howl or The Howl may also refer to:
Film
* '' The Howl'', a 1970 Italian film
* ''Howl'' (2010 film), a 2010 Am ...
made by solitary individuals.
Humans
Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
are the coyote's greatest threat, followed by
cougar
The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...
s and gray wolves. Despite predation by gray wolves, coyotes sometimes mate with them, and with eastern, or red wolves, producing "
coywolf
A coywolf is a canid hybrid descended from coyotes (''Canis latrans''), Eastern wolf, eastern wolves (''Canis lycaon''), Gray wolf, gray wolves (''Canis lupus''), and dogs (''Canis familiaris''). All of these species are members of the genus ''C ...
" hybrids. In the northeastern regions of North America, the
eastern coyote
An eastern coyote in the snow near the Virginia.html" ;"title="West Virginia–Virginia">West Virginia–Virginia state line
The eastern coyote is a wild North American canine hybrid with both coyote and wolf parentage. The hybridization likely ...
(a larger subspecies, though still smaller than wolves) is the result of various historical and recent matings with various types of wolves. Genetic studies show that most North American wolves contain some level of coyote DNA.
The coyote is a prominent character in Native American folklore, mainly in
Aridoamerica
Aridoamerica is a cultural and ecological region spanning Northern Mexico and the Southwestern United States, defined by the presence of the drought-resistant, culturally significant staple food, the tepary bean ('' Phaseolus acutifolius'').P ...
, usually depicted as a
trickster
In mythology and the study of folklore and religion, a trickster is a character in a story (god, goddess, spirit, human or anthropomorphisation) who exhibits a great degree of intellect or secret knowledge and uses it to play tricks or otherw ...
that alternately assumes the form of an actual coyote or a man. As with other trickster figures, the coyote uses deception and humor to rebel against social conventions. The animal was especially respected in
Mesoamerica
Mesoamerica is a historical region and cultural area that begins in the southern part of North America and extends to the Pacific coast of Central America, thus comprising the lands of central and southern Mexico, all of Belize, Guatemala, El S ...
n
cosmology
Cosmology () is a branch of physics and metaphysics dealing with the nature of the universe, the cosmos. The term ''cosmology'' was first used in English in 1656 in Thomas Blount's ''Glossographia'', with the meaning of "a speaking of the wo ...
as a symbol of military might. After the
European colonization of the Americas
During the Age of Discovery, a large scale colonization of the Americas, involving a number of European countries, took place primarily between the late 15th century and the early 19th century. The Norse explored and colonized areas of Europe a ...
, it was seen in
Anglo-America
Anglo-America most often refers to a region in the Americas in which English is the main language and British culture and the British Empire have had significant historical, ethnic, linguistic, and cultural impact."Anglo-America", vol. 1, Mic ...
n culture as a cowardly and untrustworthy animal. Unlike wolves, which have seen their public image improve, attitudes towards the coyote remain largely negative.
Description
Coyote males average in weight, while females average , though size varies geographically. Northern subspecies, which average , tend to grow larger than the southern subspecies of Mexico, which average . Total length ranges on average from ; comprising a tail length of , with females being shorter in both body length and height. The largest coyote on record was a male killed near
Afton, Wyoming
Afton is a town in Lincoln County, Wyoming, United States. The population was 2,172 at the 2020 census. Afton is home to the world's largest arch made of elk antlers. Spanning across the four lanes of U.S. Highway 89, the arch, completed in ...
, on November19, 1937, which measured from nose to tail, and weighed . Scent glands are located at the upper side of the base of the tail and are a bluish-black color.
The color and texture of the coyote's fur vary somewhat geographically. The hair's predominant color is light gray and red or
fulvous
Fulvous is a colour, sometimes described as dull orange, brownish-yellow or tawny; it can also be likened to a variation of buff, beige or butterscotch. As an adjective it is used in the names of many species of birds, and occasionally other ...
, interspersed around the body with black and white. Coyotes living at high elevations tend to have more black and gray shades than their
desert
A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
-dwelling counterparts, which are more fulvous or whitish-gray. The coyote's fur consists of short, soft underfur and long, coarse guard hairs. The fur of northern subspecies is longer and denser than in southern forms, with the fur of some Mexican and Central American forms being almost hispid (bristly). Generally, adult coyotes (including coywolf hybrids) have a sable coat color, dark neonatal coat color, bushy tail with an active supracaudal gland, and a white facial mask.
Albinism
Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and reddish pink or blue eyes. Individuals with the condition are referred to as albinos.
Varied use and interpretation of ...
is extremely rare in coyotes. Out of a total of 750,000 coyotes killed by federal and cooperative hunters between March 1938 and June 1945, only two were albinos.
The coyote is typically smaller than the gray wolf, but has longer ears and a relatively larger
braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, brain-pan, or brainbox, is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calv ...
, as well as a thinner frame, face, and muzzle. The scent glands are smaller than the gray wolf's, but are the same color. Its fur color variation is much less varied than that of a wolf. The coyote also carries its tail downwards when running or walking, rather than horizontally as the wolf does.
Coyote tracks can be distinguished from those of dogs by their more elongated, less rounded shape. Unlike dogs, the upper canines of coyotes extend past the mental foramina.
Taxonomy and evolution
History
At the time of the European colonization of the Americas, coyotes were largely confined to open plains and arid regions of the western half of the continent. In early post-Columbian historical records, determining whether the writer is describing coyotes or wolves is often difficult. One record from 1750 in
Kaskaskia, Illinois
Kaskaskia is a village in Randolph County, Illinois on the Mississippi River. Having been inhabited by indigenous peoples, the village was settled by France as part of the Illinois Country and was named for the Kaskaskia people. Its population p ...
, written by a local priest, noted that the "wolves" encountered there were smaller and less daring than European wolves. Another account from the early 1800s in Edwards County, Illinois mentioned wolves howling at night, though these were likely coyotes. This species was encountered several times during the
Lewis and Clark Expedition
The Lewis and Clark Expedition, also known as the Corps of Discovery Expedition, was the United States expedition to cross the newly acquired western portion of the country after the Louisiana Purchase. The Corps of Discovery was a select gro ...
(1804–1806), though it was already well known to European traders on the upper
Missouri
Missouri (''see #Etymology and pronunciation, pronunciation'') is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it border ...
.
Meriwether Lewis
Meriwether Lewis (August 18, 1774 – October 11, 1809) was an American explorer, soldier, politician, and public administrator, best known for his role as the leader of the Lewis and Clark Expedition, also known as the Corps of Discovery, with ...
, writing on 5 May 1805, in northeastern
Montana
Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
, described the coyote in these terms:
The coyote was first scientifically described by
naturalist
Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Thomas Say
Thomas Say (June 27, 1787 – October 10, 1834) was an American entomologist, conchologist, and Herpetology, herpetologist. His studies of insects and shells, numerous contributions to scientific journals, and scientific expeditions to Florida, Ge ...
in September 1819, on the site of Lewis and Clark's Council Bluffs, up the Missouri River from the mouth of the Platte during a government-sponsored expedition with Major Stephen Long. He had the first edition of the Lewis and Clark journals in hand, which contained Biddle's edited version of Lewis's observations dated 5 May 1805. His account was published in 1823. Say was the first person to document the difference between a "''prairie wolf''" (coyote) and on the next page of his journal a wolf which he named ''Canis nubilus'' ( Great Plains wolf). Say described the coyote as:
Naming and etymology
The first published usage of the word "coyote" (which is a Spanish borrowing of its
Nahuatl
Nahuatl ( ; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahuas, most of whom live mainly in Central Mexico and have smaller popul ...
name ''coyōtl'' ) comes from the
historian
A historian is a person who studies and writes about the past and is regarded as an authority on it. Historians are concerned with the continuous, methodical narrative and research of past events as relating to the human species; as well as the ...
Francisco Javier Clavijero's ''Historia de México'' in 1780. The first time it was used in English occurred in William Bullock's ''Six months' residence and travels in Mexico'' (1824), where it is variously transcribed as ''cayjotte'' and ''cocyotie''. The word's spelling was standardized as "coyote" by the 1880s.
The English pronunciation is heard both as a two-syllable word (with the final "e" silent) and as three-syllables (with the final "e" pronounced), with a tendency for the three-syllable pronunciation in eastern states and near the Mexican border, and outside the United States, with two syllables in western and central states.
Alternative English names for the coyote include "prairie wolf", "brush wolf", "cased wolf", "little wolf" and "American jackal". Its
binomial name
In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Canis latrans'' translates to "barking dog", a reference to the many vocalizations they produce.
Evolution
Fossil record
Xiaoming Wang and Richard H. Tedford, one of the foremost authorities on carnivore evolution, proposed that the genus ''Canis'' was the descendant of the coyote-like '' Eucyon davisi'' and its remains first appeared in the
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
6million years ago (Mya) in the southwestern US and Mexico. By the
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Canis lepophagus'' appeared in the same region and by the
early Pleistocene
The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
(1Mya) ''C.latrans'' (the coyote) was in existence. They proposed that the progression from ''Eucyon davisi'' to ''C.lepophagus'' to the coyote was linear evolution.
''C.latrans'' and ''C.aureus'' are closely related to '' C.edwardii'', a species that appeared earliest spanning the mid-
Blancan
The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years BP, a period of .late Pliocene
Late or LATE may refer to:
Everyday usage
* Tardy, or late, not being on time
* Late (or the late) may refer to a person who is dead
Music
* Late (The 77s album), ''Late'' (The 77s album), 2000
* Late (Alvin Batiste album), 1993
* Late!, a pseudo ...
) to the close of the
Irvingtonian
The Irvingtonian North American Land Mammal Age on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), spanning from 1.8 million – 250,000 years Before Present, BP.
(late Pleistocene), and coyote remains indistinguishable from ''C. latrans'' were contemporaneous with ''C.edwardii'' in North America. Johnston describes ''C.lepophagus'' as having a more slender skull and skeleton than the modern coyote. Ronald Nowak found that the early populations had small, delicate, narrowly proportioned skulls that resemble small coyotes and appear to be ancestral to ''C. latrans''.
''C. lepophagus'' was similar in weight to modern coyotes, but had shorter limb bones that indicate a less cursorial lifestyle. The coyote represents a more primitive form of ''Canis'' than the gray wolf, as shown by its relatively small size and its comparatively narrow skull and jaws, which lack the grasping power necessary to hold the large prey in which wolves specialize. This is further corroborated by the coyote's
sagittal crest
A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are excepti ...
, which is low or totally flattened, thus indicating a weaker bite than the wolves. The coyote, unlike the wolf, is not a specialized carnivore, as shown by the larger chewing surfaces on the
molars
The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, '' ...
, reflecting the species' relative dependence on vegetable matter. In these respects, the coyote resembles the fox-like progenitors of the genus more so than the wolf.
The oldest fossils that fall within the range of the modern coyote date to 0.74–0.85 Ma (million years) in Hamilton Cave, West Virginia; 0.73 Ma in Irvington, California; 0.35–0.48 Ma in Porcupine Cave, Colorado, and in Cumberland Cave, Pennsylvania. Modern coyotes arose 1,000 years after the
Quaternary extinction event
The Late Pleistocene to the beginning of the Holocene saw the extinction of the majority of the world's megafauna, typically defined as animal species having body masses over , which resulted in a collapse in faunal density and diversity acro ...
. Compared to their modern
Holocene
The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
counterparts, Pleistocene coyotes (''C.l. orcutti'') were larger and more robust, likely in response to larger
competitors
Competition is a rivalry where two or more parties strive for a common goal which cannot be shared: where one's gain is the other's loss (an example of which is a zero-sum game). Competition can arise between entities such as organisms, indi ...
and prey. Pleistocene coyotes were likely more specialized carnivores than their descendants, as their teeth were more adapted to shearing meat, showing fewer grinding surfaces suited for processing vegetation. Their reduction in size occurred within 1,000 years of the Quaternary extinction event, when their large prey died out. Furthermore, Pleistocene coyotes were unable to exploit the big-game hunting niche left vacant after the extinction of the
dire wolf
The dire wolf (''Aenocyon dirus'' ) is an Extinction, extinct species of Caninae, canine which was native to the Americas during the Late Pleistocene and Early Holocene epochs (125,000–10,000 years ago). The species was named in 1858, four y ...
(''Aenocyondirus''), as it was rapidly filled by gray wolves, which likely actively killed off the large coyotes, with
natural selection
Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
favoring the modern gracile morph. Coyotes are usually 5-7 feet long.
DNA evidence
In 1993, a study proposed that the wolves of North America display skull traits more similar to the coyote than wolves from Eurasia. In 2010, a study found that the coyote was a basal member of the clade that included the Tibetan wolf, the
domestic dog
The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers ...
Eurasian wolf
The Eurasian wolf (''Canis lupus lupus''), also known as the common wolf,Mech, L. David (1981), ''The Wolf: The Ecology and Behaviour of an Endangered Species'', University of Minnesota Press, p. 354, is a subspecies of grey wolf native to Euro ...
, with the Tibetan wolf diverging early from wolves and domestic dogs.
In 2016, a whole-genome DNA study proposed, based on the assumptions made, that all of the North American wolves and coyotes diverged from a common ancestor about 51,000 years ago. However, the proposed timing of the wolf / coyote divergence conflicts with the discovery of a coyote-like specimen in strata dated to 1 Mya. The study also indicated that all North American wolves have a significant amount of coyote ancestry and all coyotes some degree of wolf ancestry, and that the red wolf and eastern wolf are highly admixed with different proportions of gray wolf and coyote ancestry.
Genetic studies relating to wolves or dogs have inferred phylogenetic relationships based on the only reference genome available, that of the Boxer dog. In 2017, the first reference genome of the wolf ''Canis lupus lupus'' was mapped to aid future research. In 2018, a study looked at the genomic structure and admixture of North American wolves, wolf-like canids, and coyotes using specimens from across their entire range that mapped the largest dataset of nuclear genome sequences against the wolf reference genome.
The study supports the findings of previous studies that North American gray wolves and wolf-like canids were the result of complex gray wolf and coyote mixing. A polar wolf from Greenland and a coyote from Mexico represented the purest specimens. The coyotes from Alaska, California, Alabama, and Quebec show almost no wolf ancestry. Coyotes from Missouri, Illinois, and Florida exhibit 5–10% wolf ancestry. There was 40% wolf to 60% coyote ancestry in red wolves, 60% wolf to 40% coyote in Eastern timber wolves, and 75% wolf to 25% coyote in the Great Lakes wolves. There was 10% coyote ancestry in Mexican wolves and the Atlantic Coast wolves, 5% in Pacific Coast and Yellowstone wolves, and less than 3% in Canadian archipelago wolves. If a third canid had been involved in the admixture of the North American wolf-like canids, then its genetic signature would have been found in coyotes and wolves, which it has not.
In 2018,
whole genome sequencing
Whole genome sequencing (WGS), also known as full genome sequencing or just genome sequencing, is the process of determining the entirety of the DNA sequence of an organism's genome at a single time. This entails sequencing all of an organism's ...
was used to compare members of the genus ''Canis''. The study indicates that the common ancestor of the coyote and gray
wolf
The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been recognized, includin ...
has genetically admixed with a ghost population of an extinct, unidentified canid. The "ghost" canid was genetically close to the dhole, and had evolved after the divergence of the
African wild dog
The African wild dog (''Lycaon pictus''), also called painted dog and Cape hunting dog, is a wild canine native to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Lycaon'', which is disti ...
from the other canid species. The basal position of the coyote compared to the wolf is proposed to be due to the coyote retaining more of the mitochondrial genome from the unknown extinct canid.
Subspecies
, 19 subspecies are recognized.
Geographic variation in coyotes is not great; however, taken as a whole, the eastern subspecies ( and ) are large, dark-colored animals, with a gradual paling in color and reduction in size westward and northward (, , , and ), a brightening of 'ochraceous' tones – deep orange or brown – towards the Pacific coast (, ), a reduction in size in
Aridoamerica
Aridoamerica is a cultural and ecological region spanning Northern Mexico and the Southwestern United States, defined by the presence of the drought-resistant, culturally significant staple food, the tepary bean ('' Phaseolus acutifolius'').P ...
(, ) and a general trend towards dark reddish colors and short muzzles in Mexican and Central American populations.
Hybridization
Coyotes occasionally mate with
domestic dog
The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers ...
s, sometimes producing crosses colloquially known as "
coydog
A coydog is a canid hybrid resulting from a mating between a male coyote and a female dog. Hybrids of both sexes are fertile and can be successfully bred through four generations. Similarly, a dogote is a hybrid with a dog father and a coyote mothe ...
s". Such matings are rare in the wild, as the mating cycles of dogs and coyotes do not coincide, and coyotes are usually antagonistic towards dogs. Hybridization usually only occurs when coyotes are expanding into areas where conspecifics are few, and dogs are the only alternatives. Even then, pup survival rates are lower than normal, as dogs do not form pair bonds with coyotes, thus making the rearing of pups more difficult. In captivity, F1 hybrids (first generation) tend to be more mischievous and less manageable as pups than dogs, and are less trustworthy on maturity than wolf-dog hybrids.
Hybrids vary in appearance, but generally retain the coyote's usual characteristics. F1 hybrids tend to be intermediate in form between dogs and coyotes, while F2 hybrids (second generation) are more varied. Both F1 and F2 hybrids resemble their coyote parents in terms of shyness and intrasexual aggression. Hybrids are fertile and can be successfully bred through four generations.Melanistic coyotes owe their black pelts to a mutation that first arose in domestic dogs. A population of non-albino white coyotes in Newfoundland owe their coloration to a
melanocortin 1 receptor
The melanocortin 1 receptor (MC1R), also known as melanocyte-stimulating hormone receptor (MSHR), melanin-activating peptide receptor, or melanotropin receptor, is a G protein–coupled receptor that binds to a class of pituitary peptide hormon ...
mutation inherited from Golden Retrievers.
Coyotes have hybridized with wolves to varying degrees, particularly in eastern
North America
North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. The so-called "
eastern coyote
An eastern coyote in the snow near the Virginia.html" ;"title="West Virginia–Virginia">West Virginia–Virginia state line
The eastern coyote is a wild North American canine hybrid with both coyote and wolf parentage. The hybridization likely ...
" of northeastern North America probably originated in the aftermath of the extermination of gray and eastern wolves in the northeast, thus allowing coyotes to colonize former wolf ranges and mix with the remnant wolf populations. This hybrid is smaller than either the gray or eastern wolf, and holds smaller territories, but is in turn larger and holds more extensive home ranges than the typical western coyote. , the eastern coyote's genetic makeup is fairly uniform, with minimal influence from eastern wolves or western coyotes.
Adult eastern coyotes are larger than western coyotes, with female eastern coyotes weighing 21% more than male western coyotes. Physical differences become more apparent by the age of 35 days, with eastern coyote pups having longer legs than their western counterparts. Differences in dental development also occurs, with
tooth eruption
Tooth eruption is a process in tooth development in which the teeth enter the mouth and become visible. It is currently believed that the periodontal ligament plays an important role in tooth eruption. The first human teeth to appear, the decidu ...
being later, and in a different order in the eastern coyote. Aside from its size, the eastern coyote is physically similar to the western coyote. The four color phases range from dark brown to blond or reddish blond, though the most common phase is gray-brown, with reddish legs, ears, and flanks.
No significant differences exist between eastern and western coyotes in aggression and fighting, though eastern coyotes tend to fight less, and are more playful. Unlike western coyote pups, in which fighting precedes play behavior, fighting among eastern coyote pups occurs after the onset of play. Eastern coyotes tend to reach
sexual maturity
Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...
at two years of age, much later than in western coyotes.
Eastern and red wolves are also products of varying degrees of wolf-coyote hybridization. The eastern wolf probably was a result of a wolf-coyote admixture, combined with extensive
backcrossing
Backcrossing is a crossing of a hybrid with one of its parents or an individual genetically similar to its parent, to achieve offspring with a genetic identity closer to that of the parent. It is used in horticulture, animal breeding, and produc ...
with parent gray wolf populations. The red wolf may have originated during a time of declining wolf populations in the
Southeastern Woodlands
Indigenous peoples of the Southeastern Woodlands, Southeastern cultures, or Southeast Indians are an Ethnography, ethnographic classification for Native Americans in the United States, Native Americans who have traditionally inhabited the area now ...
, forcing a wolf-coyote hybridization, as well as backcrossing with local parent coyote populations to the extent that about 75–80% of the modern red wolf's genome is of coyote derivation.
Behavior
Social and reproductive behaviors
Like the Eurasian golden jackal, the coyote is gregarious, but not as dependent on conspecifics as more social canid species like wolves are. This is likely because the coyote is not a specialized hunter of large prey as the latter species is. The basic social unit of a coyote pack is a family containing a reproductive female. However, unrelated coyotes may join forces for companionship, or to bring down prey too large to attack on their own. Such "nonfamily" packs are only temporary, and may consist of bachelor males, nonreproductive females and subadult young. Families are formed in midwinter, when females enter
estrus
The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...
. Pair bonding can occur 2–3 months before actual copulation takes place.
The
copulatory tie
Canine reproduction is the process of sexual reproduction in domestic dogs, wolves, coyotes and other canine species.
Canine sexual anatomy and development Male reproductive system
Erectile tissue
As with all mammals, a dog's penis is made up ...
can last 5–45 minutes. A female entering estrus attracts males by scent marking and howling with increasing frequency. A single female in heat can attract up to seven reproductive males, which can follow her for as long as a month. Although some squabbling may occur among the males, once the female has selected a mate and copulates, the rejected males do not intervene, and move on once they detect other estrous females. Unlike the wolf, which has been known to practice both
monogamous
Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
and bigamous matings, the coyote is strictly monogamous, even in areas with high coyote densities and abundant food.
Females that fail to mate sometimes assist their sisters or mothers in raising their pups, or join their siblings until the next time they can mate. The newly mated pair then establishes a territory and either constructs their own den or cleans out abandoned
badger
Badgers are medium-sized short-legged omnivores in the superfamily Musteloidea. Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by their squat bodies and adaptions for fossorial activity rather than by the ...
,
marmot
Marmots are large ground squirrels in the genus ''Marmota'', with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, w ...
, or
skunk
Skunks are mammals in the family Mephitidae. They are known for their ability to spray a liquid with a strong, unpleasant scent from their anal glands. Different species of skunk vary in appearance from black-and-white to brown, cream or gi ...
earths. During the pregnancy, the male frequently hunts alone and brings back food for the female. The female may line the den with dried grass or with fur pulled from her belly. The
gestation period
In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once i ...
is 63 days, with an average litter size of six, though the number fluctuates depending on coyote population density and the abundance of food.
Coyote pups are born in dens, hollow trees, or under ledges, and weigh at birth. They are
altricial
Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
, and are completely dependent on
milk
Milk is a white liquid food produced by the mammary glands of lactating mammals. It is the primary source of nutrition for young mammals (including breastfeeding, breastfed human infants) before they are able to digestion, digest solid food. ...
for their first 10 days. The
incisor
Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s erupt at about 12 days, the canines at 16, and the second
premolar
The premolars, also called premolar Tooth (human), teeth, or bicuspids, are transitional teeth located between the Canine tooth, canine and Molar (tooth), molar teeth. In humans, there are two premolars per dental terminology#Quadrant, quadrant in ...
s at 21. Their eyes open after 10 days, by which point the pups become increasingly more mobile, walking by 20 days, and running at the age of six weeks. The parents begin supplementing the pup's diet with regurgitated solid food after 12–15 days. By the age of four to six weeks, when their
milk teeth
Milk Teeth (often stylised as MILK TEETH) were a British punk rock band from Stroud, Gloucestershire, which formed in May 2013. The band officially announced their breakup on Facebook and Twitter on 4 September 2020.
History Formation and earl ...
are fully functional, the pups are given small food items such as mice, rabbits, or pieces of
ungulate
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
carcasses, with
lactation
Lactation describes the secretion of milk from the mammary glands and the period of time that a mother lactates to feed her young. The process naturally occurs with all sexually mature female mammals, although it may predate mammals. The process ...
steadily decreasing after two months.
Unlike wolf pups, coyote pups begin seriously fighting (as opposed to play fighting) prior to engaging in play behavior. A common play behavior includes the coyote "hip-slam". By three weeks of age, coyote pups bite each other with less inhibition than wolf pups. By the age of four to five weeks, pups have established dominance hierarchies, and are by then more likely to play rather than fight. The male plays an active role in feeding, grooming, and guarding the pups, but abandons them if the female goes missing before the pups are completely
weaned
Weaning is the process of gradually introducing an infant human or other mammal to what will be its adult diet while withdrawing the supply of its mother's milk. In the UK, weaning primarily refers to the introduction of solid foods at 6 mont ...
. The den is abandoned by June to July, and the pups follow their parents in patrolling their territory and hunting. Pups may leave their families in August, though can remain for much longer. The pups attain adult dimensions at eight months and gain adult weight a month later.
Territorial and sheltering behaviors
Individual feeding territories vary in size from , with the general concentration of coyotes in a given area depending on food abundance, adequate denning sites, and competition with conspecifics and other predators. The coyote generally does not defend its territory outside of the denning season, and is much less aggressive towards intruders than the wolf is, typically chasing and sparring with them, but rarely killing them. Conflicts between coyotes can arise during times of food shortage. Coyotes mark their territories by raised-leg urination and ground-scratching.
Like wolves, coyotes use a den, usually the deserted holes of other species, when gestating and rearing young, though they may occasionally give birth under sagebrushes in the open. Coyote dens can be located in
canyon
A canyon (; archaic British English spelling: ''cañon''), gorge or chasm, is a deep cleft between escarpments or cliffs resulting from weathering and the erosive activity of a river over geologic time scales. Rivers have a natural tendency t ...
bank
A bank is a financial institution that accepts Deposit account, deposits from the public and creates a demand deposit while simultaneously making loans. Lending activities can be directly performed by the bank or indirectly through capital m ...
s, rock bluffs, or level ground. Some dens have been found under abandoned homestead shacks, grain bins, drainage pipes, railroad tracks, hollow logs, thickets, and thistles. The den is continuously dug and cleaned out by the female until the pups are born. Should the den be disturbed or infested with fleas, the pups are moved into another den. A coyote den can have several entrances and passages branching out from the main chamber. A single den can be used year after year.
Hunting and feeding behaviors
While the popular consensus is that
olfaction
The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste.
In humans, ...
is very important for hunting, two studies that experimentally investigated the role of olfactory, auditory, and visual cues found that visual cues are the most important ones for hunting in red foxes and coyotes.
When hunting large prey, the coyote often works in pairs or small groups. Success in killing large
ungulate
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
s depends on factors such as snow depth and crust density. Younger animals usually avoid participating in such hunts, with the breeding pair typically doing most of the work. The coyote pursues large prey, typically hamstringing the animal, and subsequently then harassing it until the prey falls. Like other canids, the coyote caches excess food. Coyotes catch mouse-sized rodents by pouncing, whereas
ground squirrel
Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s are chased. Although coyotes can live in large groups, small prey is typically caught singly.
Coyotes have been observed to kill
porcupine
Porcupines are large rodents with coats of sharp Spine (zoology), spines, or quills, that protect them against predation. The term covers two Family (biology), families of animals: the Old World porcupines of the family Hystricidae, and the New ...
s in pairs, using their paws to flip the rodents on their backs, then attacking the soft underbelly. Only old and experienced coyotes can successfully prey on porcupines, with many predation attempts by young coyotes resulting in them being injured by their prey's quills. Coyotes sometimes
urinate
Urination is the release of urine from the bladder through the urethra in placental mammals, or through the cloaca in other vertebrates. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, ure ...
on their food, possibly to claim ownership over it.Wells, Michael C., and Marc Bekoff. An observational study of scent-marking in coyotes, Canis latrans " (1981). Recent evidence demonstrates that at least some coyotes have become more nocturnal in hunting, presumably to avoid humans.
Coyotes may occasionally form mutualistic hunting relationships with American badgers, assisting each other in digging up rodent prey. The relationship between the two species may occasionally border on apparent "friendship", as some coyotes have been observed laying their heads on their badger companions or licking their faces without protest. The amicable interactions between coyotes and badgers were known to pre-Columbian civilizations, as shown on a jar found in Mexico dated to 1250–1300 CE depicting the relationship between the two.
Food scraps, pet food, and animal feces may attract a coyote to a trash can.
Communication
Body language
Being both a gregarious and solitary animal, the variability of the coyote's visual and vocal repertoire is intermediate between that of the solitary foxes and the highly social wolf. The aggressive behavior of the coyote bears more similarities to that of foxes than it does that of wolves and dogs. An aggressive coyote arches its back and lowers its tail. Unlike dogs, which solicit playful behavior by performing a "play-bow" followed by a "play-leap", play in coyotes consists of a bow, followed by side-to-side head flexions and a series of "spins" and "dives". Although coyotes will sometimes bite their playmates' scruff as dogs do, they typically approach low, and make upward-directed bites.
Pups fight each other regardless of sex, while among adults, aggression is typically reserved for members of the same sex. Combatants approach each other waving their tails and snarling with their jaws open, though fights are typically silent. Males tend to fight in a vertical stance, while females fight on all four paws. Fights among females tend to be more serious than ones among males, as females seize their opponents' forelegs, throat, and shoulders.
Vocalizations
The coyote has been described as "the most vocal of all ildNorth American mammals". Its loudness and range of vocalizations was the cause for its binomial name ''Canis latrans'', meaning "barking dog". At least 11 different vocalizations are known in adult coyotes. These sounds are divided into three categories: agonistic and alarm, greeting, and contact. Vocalizations of the first category include woofs, growls, huffs, barks, bark howls, yelps, and high-frequency whines. Woofs are used as low-intensity threats or alarms and are usually heard near den sites, prompting the pups to immediately retreat into their burrows.
Growls are used as threats at short distances but have also been heard among pups playing and copulating males. Huffs are high-intensity threat vocalizations produced by rapid expiration of air. Barks can be classed as both long-distance threat vocalizations and alarm calls. Bark howls may serve similar functions. Yelps are emitted as a sign of submission, while high-frequency whines are produced by dominant animals acknowledging the submission of subordinates. Greeting vocalizations include low-frequency whines, 'wow-oo-wows', and group yip howls. Low-frequency whines are emitted by submissive animals and are usually accompanied by tail wagging and muzzle nibbling.
The sound known as 'wow-oo-wow' has been described as a "greeting song". The group yip howl is emitted when two or more pack members reunite and may be the final act of a complex greeting ceremony. Contact calls include lone howls and group howls, as well as the previously mentioned group yip howls. The lone howl is the most iconic sound of the coyote and may serve the purpose of announcing the presence of a lone individual separated from its pack. Group howls are used as both substitute group yip howls and as responses to either lone howls, group howls, or group yip howls.
Ecology
Habitat
Prior to the near extermination of wolves and cougars, the coyote was most numerous in
grassland
A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s inhabited by
bison
A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised.
Of the two surviving species, the American ...
,
pronghorn
The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...
, elk, and other deer, doing particularly well in short-grass areas with
prairie dog
Prairie dogs (genus ''Cynomys'') are herbivorous burrowing Marmotini, ground squirrels native to the grasslands of North America. There are five recognized species of prairie dog: black-tailed prairie dog, black-tailed, white-tailed prairie dog ...
s, though it was just as much at home in semiarid areas with
sagebrush
Sagebrush is the common name of several woody and herbaceous species of plants in the genus ''Artemisia (plant), Artemisia''. The best-known sagebrush is the shrub ''Artemisia tridentata''. Sagebrush is native to the western half of North Amer ...
cactus
A cactus (: cacti, cactuses, or less commonly, cactus) is a member of the plant family Cactaceae (), a family of the order Caryophyllales comprising about 127 genera with some 1,750 known species. The word ''cactus'' derives, through Latin, ...
,
kangaroo rat
Kangaroo rats, small mostly nocturnal rodents of genus ''Dipodomys'', are native to arid areas of western North America. The common name derives from their bipedal form. They hop in a manner similar to the much larger kangaroo, but developed thi ...
s, and
rattlesnake
Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
s. As long as it was not in direct competition with the wolf, the coyote ranged from the
Sonoran Desert
The Sonoran Desert () is a hot desert and ecoregion in North America that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the Southwestern United States (in Arizona and California). It ...
to the alpine regions of adjoining mountains or the plains and mountainous areas of
Alberta
Alberta is a Provinces and territories of Canada, province in Canada. It is a part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to its west, Saskatchewan to its east, t ...
. With the extermination of the wolf, the coyote's range expanded to encompass broken forests from the tropics of
Guatemala
Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
and the northern slope of
Alaska
Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
.
Coyotes walk around per day, often along trails such as logging roads and paths; they may use iced-over rivers as travel routes in winter. They are often
crepuscular
In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal (active during dawn), vespertine (biology), vespertine/vespertinal (active during dusk), or both. This is distinguished from diurnalit ...
, being more active around evening and the beginning of the night than during the day. However, in urban areas coyotes are known to be more nocturnal, likely to avoid encounters with humans. Like many canids, coyotes are competent swimmers, reported to be able to travel at least across water.
Diet
The coyote is ecologically the North American equivalent of the Eurasian
golden jackal
The golden jackal (''Canis aureus''), also called the common jackal, is a wolf-like canid that is native to Eurasia. The golden jackal's coat varies in color from a pale creamy yellow in summer to a dark tawny beige in winter. It is smaller a ...
. Likewise, the coyote is highly versatile in its choice of food, but is primarily
carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
, with 90% of its diet consisting of meat. Prey species include
bison
A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised.
Of the two surviving species, the American ...
(largely as
carrion
Carrion (), also known as a carcass, is the decaying flesh of dead animals.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
),
white-tailed deer
The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
,
mule deer
The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer.
Unlike the related whit ...
,
moose
The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
bighorn sheep
The bighorn sheep (''Ovis canadensis'') is a species of Ovis, sheep native to North America. It is named for its large Horn (anatomy), horns. A pair of horns may weigh up to ; the sheep typically weigh up to . Recent genetic testing indicates th ...
,
pronghorn
The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...
,
rabbit
Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s,
hare
Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
s,
rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s,
bird
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s (especially
galliformes
Galliformes is an order (biology), order of heavy-bodied ground-feeding birds that includes turkey (bird), turkeys, chickens, Old World quail, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems ...
water bird
A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from seabi ...
amphibian
Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s (except
toad
Toad (also known as a hoptoad) is a common name for certain frogs, especially of the family Bufonidae, that are characterized by dry, leathery skin, short legs, and large bumps covering the parotoid glands.
In popular culture (folk taxonomy ...
s),
lizard
Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s,
snake
Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s,
turtle
Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s and
tortoise
Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...
s,
fish
A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
,
crustacean
Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s, and
insect
Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s. Coyotes may be picky over the prey they target, as animals such as
shrew
Shrews ( family Soricidae) are small mole-like mammals classified in the order Eulipotyphla. True shrews are not to be confused with treeshrews, otter shrews, elephant shrews, West Indies shrews, or marsupial shrews, which belong to dif ...
brown rat
The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat and Norwegian rat, is a widespread species of common rat. One of the largest Muroidea, muroids, it is a brown or grey ...
s do not occur in their diet in proportion to their numbers.Terrestrial animals or burrowing small mammals such as
ground squirrel
Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s and associated species (
marmot
Marmots are large ground squirrels in the genus ''Marmota'', with 15 species living in Asia, Europe, and North America. These herbivores are active during the summer, when they can often be found in groups, but are not seen during the winter, w ...
s,
prairie dog
Prairie dogs (genus ''Cynomys'') are herbivorous burrowing Marmotini, ground squirrels native to the grasslands of North America. There are five recognized species of prairie dog: black-tailed prairie dog, black-tailed, white-tailed prairie dog ...
s,
chipmunk
Chipmunks are small, striped rodents of subtribe Tamiina. Chipmunks are found in North America, with the exception of the Siberian chipmunk which is found primarily in Asia.
Taxonomy and systematics
Chipmunks are classified as four genera: '' ...
s) as well as
vole
Voles are small rodents that are relatives of lemmings and hamsters, but with a stouter body; a longer, hairy tail; a slightly rounder head; smaller eyes and ears; and differently formed molars (high-crowned with angular cusps instead of lo ...
s,
pocket gopher
Pocket gophers, commonly referred to simply as gophers, are burrowing rodents of the family Geomyidae. The roughly 41 speciesSearch results for "Geomyidae" on thASM Mammal Diversity Database are all endemic to North and Central America. They ar ...
s,
kangaroo rat
Kangaroo rats, small mostly nocturnal rodents of genus ''Dipodomys'', are native to arid areas of western North America. The common name derives from their bipedal form. They hop in a manner similar to the much larger kangaroo, but developed thi ...
rattlesnake
Rattlesnakes are venomous snakes that form the genus, genera ''Crotalus'' and ''Sistrurus'' of the subfamily Crotalinae (the pit vipers). All rattlesnakes are vipers. Rattlesnakes are predators that live in a wide array of habitats, hunting sm ...
s. Coyotes kill rattlesnakes mostly for food, but also to protect their pups at their dens, by teasing the snakes until they stretch out and then biting their heads and snapping and shaking the snakes. Birds taken by coyotes may range in size from thrashers,
lark
Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
wild turkey
The wild turkey (''Meleagris gallopavo'') is an upland game bird native to North America, one of two extant species of Turkey (bird), turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic turkey (''M. g. dom ...
s and, rarely, brooding adult swans and pelicans.
If working in packs or pairs, coyotes may have access to larger prey than lone individuals normally take, such as various prey weighing more than . In some cases, packs of coyotes have dispatched much larger prey such as adult '' Odocoileus'' deer, cow elk,
pronghorn
The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...
s and wild sheep, although the young fawn, calves and lambs of these animals are considerably more often taken even by packs, as well as
domestic sheep
Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to ...
and domestic cattle. In some cases, coyotes can bring down prey weighing up to or more. When it comes to adult ungulates such as wild deer, they often exploit them when vulnerable such as those that are infirm, stuck in snow or ice, otherwise winter-weakened or heavily pregnant, whereas less wary domestic ungulates may be more easily exploited.
Although coyotes prefer fresh meat, they will
scavenge
Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding be ...
when the opportunity presents itself. Excluding the insects, fruit, and grass eaten, the coyote requires an estimated of food daily, or annually. The coyote readily cannibalizes the carcasses of conspecifics, with coyote fat having been successfully used by coyote hunters as a lure or poisoned bait. The coyote's winter diet consists mainly of large ungulate carcasses, with very little plant matter. Rodent prey increases in importance during the spring, summer, and fall.
The coyote feeds on a variety of different
produce
In American English, produce generally refers to wikt:fresh, fresh List of culinary fruits, fruits and Vegetable, vegetables intended to be Eating, eaten by humans, although other food products such as Dairy product, dairy products or Nut (foo ...
, including
strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
,
blackberries
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
,
blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
peach
The peach (''Prunus persica'') is a deciduous tree first domesticated and Agriculture, cultivated in China. It bears edible juicy fruits with various characteristics, most called peaches and the glossy-skinned, non-fuzzy varieties called necta ...
es,
pear
Pears are fruits produced and consumed around the world, growing on a tree and harvested in late summer into mid-autumn. The pear tree and shrub are a species of genus ''Pyrus'' , in the Family (biology), family Rosaceae, bearing the Pome, po ...
s,
apple
An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
persimmon
The persimmon () is the edible fruit of a number of species of trees in the genus '' Diospyros''. The most widely cultivated of these is the Chinese and Japanese kaki persimmon, ''Diospyros kaki''. In 2022, China produced 77% of the world's p ...
s,
peanut
The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), goober pea, pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics by small and large ...
s,
watermelon
The watermelon (''Citrullus lanatus'') is a species of flowering plant in the family Cucurbitaceae, that has a large, edible fruit. It is a Glossary of botanical terms#scandent, scrambling and trailing vine-like plant, and is plant breeding ...
s,
cantaloupe
The cantaloupe ( ) is a type of true melon (''Cucumis melo'') with sweet, aromatic, and usually orange flesh. Originally, ''cantaloupe'' refers to the true cantaloupe or European cantaloupe with non- to slightly netted and often ribbed rind. ...
s, and
carrot
The carrot ('' Daucus carota'' subsp. ''sativus'') is a root vegetable, typically orange in colour, though heirloom variants including purple, black, red, white, and yellow cultivars exist, all of which are domesticated forms of the wild ...
s. During the winter and early spring, the coyote eats large quantities of grass, such as green
wheat
Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
blades. It sometimes eats unusual items such as cotton cake,
soybean
The soybean, soy bean, or soya bean (''Glycine max'') is a species of legume native to East Asia, widely grown for its edible bean. Soy is a staple crop, the world's most grown legume, and an important animal feed.
Soy is a key source o ...
meal, domestic animal droppings,
bean
A bean is the seed of some plants in the legume family (Fabaceae) used as a vegetable for human consumption or animal feed. The seeds are often preserved through drying (a ''pulse''), but fresh beans are also sold. Dried beans are traditi ...
s, and cultivated
grain
A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
such as
maize
Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
, wheat, and
sorghum
''Sorghum bicolor'', commonly called sorghum () and also known as great millet, broomcorn, guinea corn, durra, imphee, jowar, or milo, is a species in the Poaceae, grass genus ''Sorghum (genus), Sorghum'' cultivated for its grain. The grain i ...
.
In coastal California, coyotes now consume a higher percentage of marine-based food than their ancestors, which is thought to be due to the extirpation of the grizzly bear from this region. In
Death Valley
Death Valley is a desert valley in Eastern California, in the northern Mojave Desert, bordering the Great Basin Desert. It is thought to be the Highest temperature recorded on Earth, hottest place on Earth during summer.
Death Valley's Badwat ...
, coyotes may consume great quantities of
hawkmoth
The Sphingidae are a family of moths commonly called sphinx moths, also colloquially known as hawk moths, with many of their caterpillars known as hornworms. It includes about 1,450 species. It is best represented in the tropics, but species ar ...
caterpillar
Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths).
As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s or
beetle
Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s in the spring flowering months.
Enemies and competitors
In areas where the ranges of coyotes and gray wolves overlap, interference competition and predation by wolves has been hypothesized to limit local coyote densities. Coyote ranges expanded during the 19th and 20th centuries following the extirpation of wolves, while coyotes were driven to extinction on
Isle Royale
Isle Royale (, ) is an Islands of the Great Lakes, island of the Great Lakes located in the northwest of Lake Superior and part of the U.S. state of Michigan. The island and the 450 surrounding smaller islands and waters make up Isle Royale Na ...
after wolves colonized the island in the 1940s. One study conducted in
Yellowstone National Park
Yellowstone National Park is a List of national parks of the United States, national park of the United States located in the northwest corner of Wyoming, with small portions extending into Montana and Idaho. It was established by the 42nd U ...
, where both species coexist, concluded that the coyote population in the Lamar River Valley declined by 39% following the reintroduction of wolves in the 1990s, while coyote populations in wolf inhabited areas of the Grand Teton National Park are 33% lower than in areas where they are absent. Wolves have been observed to not tolerate coyotes in their vicinity, though coyotes have been known to trail wolves to feed on their kills.
Coyotes may compete with
cougar
The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...
s in some areas. In the eastern
Sierra Nevada
The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
, coyotes compete with cougars over
mule deer
The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer.
Unlike the related whit ...
. Cougars normally outcompete and dominate coyotes, and may kill them occasionally, thus reducing coyote predation pressure on smaller carnivores such as foxes and
bobcat
The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...
s. Coyotes that are killed are sometimes not eaten, perhaps indicating that these comprise competitive interspecies interactions, however there are multiple confirmed cases of cougars also eating coyotes. In northeastern
Mexico
Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, cougar predation on coyotes continues apace but coyotes were absent from the prey spectrum of sympatric
jaguar
The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...
s, apparently due to differing habitat usages.
Other than by gray wolves and cougars, predation on adult coyotes is relatively rare but multiple other predators can be occasional threats. In some cases, adult coyotes have been preyed upon by both American black and
grizzly bear
The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America.
In addition to the mainland grizzly (''Ursus arctos horr ...
Canada lynx
The Canada lynx (''Lynx canadensis'') or Canadian lynx is one of the four living species in the genus ''Lynx''. It is a medium-sized wild cat characterized by long, dense fur, triangular ears with black tufts at the tips, and broad, snowshoe- ...
and
golden eagle
The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of pr ...
s. At kill sites and carrion, coyotes, especially if working alone, tend to be dominated by wolves, cougars, bears,
wolverine
The wolverine ( , ; ''Gulo gulo''), also called the carcajou or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species, member of the family Mustelidae. It is a muscular carnivore and a solitary animal. The w ...
s and, usually but not always,
eagle
Eagle is the common name for the golden eagle, bald eagle, and other birds of prey in the family of the Accipitridae. Eagles belong to several groups of Genus, genera, some of which are closely related. True eagles comprise the genus ''Aquila ( ...
s (i.e., bald and golden). When such larger, more powerful or more aggressive predators come to a shared feeding site, a coyote may either try to fight, wait until the other predator is done or occasionally share a kill, but if a major danger such as wolves or an adult cougar is present, the coyote will tend to flee.
Coyotes rarely kill healthy adult
red fox
The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...
es, and have been observed to feed or den alongside them, though they often kill foxes caught in traps. Coyotes may kill fox kits, but this is not a major source of mortality. In southern California, coyotes frequently kill
gray fox
The gray fox (''Urocyon cinereoargenteus''), or grey fox, is an omnivorous mammal of the family Canidae, widespread throughout North America and Central America. This species and its only congener (biology), congener, the diminutive island fox ...
es, and these smaller canids tend to avoid areas with high coyote densities.
In some areas, coyotes share their ranges with bobcats. These two similarly-sized species rarely physically confront one another, though bobcat populations tend to diminish in areas with high coyote densities. However, several studies have demonstrated interference competition between coyotes and bobcats, and in all cases coyotes dominated the interaction. Multiple researchers reported instances of coyotes killing bobcats, whereas bobcats killing coyotes is more rare. Coyotes attack bobcats using a bite-and-shake method similar to what is used on medium-sized prey. Coyotes, both single individuals and groups, have been known to occasionally kill bobcats. In most cases, the bobcats were relatively small specimens, such as adult females and juveniles.
Coyote attacks, by an unknown number of coyotes, on adult male bobcats have occurred. In California, coyote and bobcat populations are not negatively correlated across different habitat types, but predation by coyotes is an important source of mortality in bobcats. Biologist
Stanley Paul Young
Stanley Paul Young (October 31, 1889May 15, 1969) was an American biologist who spent forty-two years working for the U.S. government in several departments which dealt with wildlife management and research. Also known as an author and ecologist, h ...
noted that in his entire trapping career, he had never successfully saved a captured bobcat from being killed by coyotes, and wrote of two incidents wherein coyotes chased bobcats up trees. Coyotes have been documented to directly kill
Canada lynx
The Canada lynx (''Lynx canadensis'') or Canadian lynx is one of the four living species in the genus ''Lynx''. It is a medium-sized wild cat characterized by long, dense fur, triangular ears with black tufts at the tips, and broad, snowshoe- ...
on occasion, and compete with them for prey, especially
snowshoe hare
The snowshoe hare (''Lepus americanus''), also called the varying hare or snowshoe rabbit, is a species of hare found in North America. It has the name "snowshoe" because of the large size of its hind feet. The animal's feet prevent it from sink ...
s. In some areas, including central Alberta, lynx are more abundant where coyotes are few, thus interactions with coyotes appears to influence lynx populations more than the availability of snowshoe hares.
Range
Due to the coyote's wide range and abundance throughout North America, it is listed as
Least Concern
A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
by the
International Union for Conservation of Nature
The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
(IUCN). The coyote's pre-Columbian range was limited to the Southwest and Plains regions of North America, and northern and central Mexico. By the 19th century, the species expanded north and east, expanding further after 1900, coinciding with land conversion and the extirpation of wolves. By this time, its range encompassed the entire North American continent, including all of the contiguous United States and Mexico, southward into Central America, and northward into most of Canada and Alaska. This expansion is ongoing, and the species now occupies the majority of areas between 8°N (Panama) and 70°N (northern Alaska).
Although it was once widely believed that coyotes are recent immigrants to southern Mexico and Central America, aided in their expansion by deforestation, Pleistocene and Early Holocene records, as well as records from the pre-Columbian period and early European colonization show that the animal was present in the area long before modern times. Range expansion occurred south of
Costa Rica
Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
during the late 1970s and northern Panama in the early 1980s, following the expansion of cattle-grazing lands into tropical rain forests.
The coyote is predicted to appear in northern
Belize
Belize is a country on the north-eastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a maritime boundary with Honduras to the southeast. P ...
in the near future, as the habitat there is favorable to the species. Concerns have been raised of a possible expansion into South America through the Panamanian Isthmus, should the
Darién Gap
The Darién Gap (, , ) is a geographic region that connects the Americas, American continents, stretching across southern Panama's Darién Province and the northern portion of Colombia's Chocó Department. Consisting of a large drainage basin, ...
ever be closed by the
Pan-American Highway
The Pan-American Highway is a vast network of roads that stretches about 30,000 kilometers (about 19,000 miles) from Prudhoe Bay, Alaska, in the northernmost part of North America to Ushuaia, Argentina, at the southern tip of South America. I ...
. This fear was partially confirmed in January 2013, when the species was recorded in eastern Panama's Chepo District, beyond the
Panama Canal
The Panama Canal () is an artificial waterway in Panama that connects the Caribbean Sea with the Pacific Ocean. It cuts across the narrowest point of the Isthmus of Panama, and is a Channel (geography), conduit for maritime trade between th ...
.
A 2017 genetic study proposes that coyotes were originally not found in the area of the eastern United States. From the 1890s, dense forests were transformed into agricultural land and wolf control implemented on a large scale, leaving a niche for coyotes to disperse into. There were two major dispersals from two populations of genetically distinct coyotes. The first major dispersal to the northeast came in the early 20th century from those coyotes living in the northern Great Plains. These came to New England via the northern Great Lakes region and southern Canada, and to Pennsylvania via the southern Great Lakes region, meeting together in the 1940s in New York and Pennsylvania.
These coyotes have hybridized with the remnant gray wolf and eastern wolf populations, which has added to coyote genetic diversity and may have assisted adaptation to the new niche. The second major dispersal to the southeast came in the mid-20th century from Texas and reached the Carolinas in the 1980s. These coyotes have hybridized with the remnant red wolf populations before the 1970s when the red wolf was extirpated in the wild, which has also added to coyote genetic diversity and may have assisted adaptation to this new niche as well. Both of these two major coyote dispersals have experienced rapid population growth and are forecast to meet along the mid-Atlantic coast. The study concludes that for coyotes the long range dispersal, gene flow from local populations, and rapid population growth may be inter-related.
Diseases and parasites
Among large North American carnivores, the coyote probably carries the largest number of diseases and parasites, likely due to its wide range and varied diet.
Viral diseases
A viral disease (or viral infection) occurs when an organism's body is invaded by pathogenic viruses, and infectious virus particles (virions) attach to and enter susceptible cells.
Examples include the common cold, gastroenteritis, COVID-19, ...
known to infect coyotes include
rabies
Rabies is a viral disease that causes encephalitis in humans and other mammals. It was historically referred to as hydrophobia ("fear of water") because its victims panic when offered liquids to drink. Early symptoms can include fever and abn ...
, canine distemper, infectious canine hepatitis, four strains of equine encephalitis, and oral papillomatosis. By the late 1970s, serious rabies outbreaks in coyotes had ceased to be a problem for over 60 years, though sporadic cases every 1–5 years did occur. Distemper causes the deaths of many pups in the wild, though some specimens can survive infection. ''
Tularemia
Tularemia, also known as rabbit fever, is an infectious disease caused by the bacterium '' Francisella tularensis''. Symptoms may include fever, skin ulcers, and enlarged lymph nodes. Occasionally, a form that results in pneumonia or a throat ...
'', a bacterial disease, infects coyotes from tick bites and through their rodent and lagomorph prey, and can be deadly for pups.
Coyotes can be infected by both demodectic and sarcoptic mange, the latter being the most common.
Mite
Mites are small arachnids (eight-legged arthropods) of two large orders, the Acariformes and the Parasitiformes, which were historically grouped together in the subclass Acari. However, most recent genetic analyses do not recover the two as eac ...
infestations are rare and incidental in coyotes, while
tick
Ticks are parasitic arachnids of the order Ixodida. They are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, and species, but can become larger when engorged. Ticks a ...
infestations are more common, with seasonal peaks depending on locality (May–August in the Northwest, March–November in Arkansas). Coyotes are only rarely infested with
lice
Louse (: lice) is the common name for any member of the infraorder Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera was previously recognized as an order, until a 2021 genetic study determined th ...
, while
flea
Flea, the common name for the order (biology), order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by hematophagy, ingesting the blood of their hosts. Adult f ...
s infest coyotes from puphood, though they may be more a source of irritation than serious illness. '' Pulex simulans'' is the most common species to infest coyotes, while '' Ctenocephalides canis'' tends to occur only in places where coyotes and dogs (its primary host) inhabit the same area. Although coyotes are rarely host to flukes, they can nevertheless have serious effects on coyotes, particularly '' Nanophyetus salmincola'', which can infect them with salmon poisoning disease, a disease with a 90% mortality rate. Trematode ''
Metorchis conjunctus
''Metorchis conjunctus'', common name Canadian liver fluke, is a species of trematode parasite in the family Opisthorchiidae. It can infect mammals that eat raw fish in North America. The first intermediate host is a freshwater snail and the seco ...
'' can also infect coyotes.
Tapeworms have been recorded to infest 60–95% of all coyotes examined. The most common species to infest coyotes are '' Taenia pisiformis'' and '' Taenia crassiceps'', which uses cottontail rabbits and rodents as intermediate hosts. The largest species known in coyotes is '' T. hydatigena'', which enters coyotes through infected
ungulates
Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to b ...
, and can grow to lengths of . Although once largely limited to wolves, ''
Echinococcus granulosus
''Echinococcus granulosus'', also called the hydatid worm or dog tapeworm, is a cyclophyllid cestode that dwells in the small intestine of canids as an adult, but which has important intermediate hosts such as livestock and humans, where it caus ...
'' has expanded to coyotes since the latter began colonizing former wolf ranges.
The most frequent ascaroid roundworm in coyotes is '' Toxascaris leonina'', which dwells in the coyote's small intestine and has no ill effects, except for causing the host to eat more frequently.
Hookworm
Hookworms are Gastrointestinal tract, intestinal, Hematophagy, blood-feeding, parasitic Nematode, roundworms that cause types of infection known as helminthiases. Hookworm infection is found in many parts of the world, and is common in areas with ...
s of the genus '' Ancylostoma'' infest coyotes throughout their range, being particularly prevalent in humid areas. In areas of high moisture, such as coastal Texas, coyotes can carry up to 250 hookworms each. The blood-drinking '' A. caninum'' is particularly dangerous, as it damages the coyote through blood loss and lung congestion. A 10-day-old pup can die from being host to as few as 25 ''A. caninum'' worms.
Relationships with humans
In folklore and mythology
Coyote features as a
trickster
In mythology and the study of folklore and religion, a trickster is a character in a story (god, goddess, spirit, human or anthropomorphisation) who exhibits a great degree of intellect or secret knowledge and uses it to play tricks or otherw ...
figure and
skin-walker
In Navajo culture, a skin-walker () is a type of harmful witch who has the ability to turn into, possess, or disguise themselves as an animal. The term is never used for healers.
The yee naaldlooshii, translating to "by means of it, it goes ...
in the folktales of some Native Americans, notably several nations in the Southwestern and Plains regions, where he alternately assumes the form of an actual coyote or that of a man. As with other trickster figures, Coyote acts as a picaresque hero who rebels against social convention through deception and humor. Folklorists such as Harris believe coyotes came to be seen as tricksters due to the animal's intelligence and adaptability. After the European colonization of the Americas,
Anglo-America
Anglo-America most often refers to a region in the Americas in which English is the main language and British culture and the British Empire have had significant historical, ethnic, linguistic, and cultural impact."Anglo-America", vol. 1, Mic ...
n depictions of Coyote are of a cowardly and untrustworthy animal. Unlike the gray wolf, which has undergone a radical improvement of its public image, Anglo-American cultural attitudes towards the coyote remain largely negative.
In the
Maidu
The Maidu are a Native Americans in the United States, Native American people of northern California. They reside in the central Sierra Nevada (U.S.), Sierra Nevada, in the watershed area of the Feather River, Feather and American River, American ...
creation story, Coyote introduces work, suffering, and death to the world. Zuni lore has Coyote bringing winter into the world by stealing light from the
kachina
A kachina (; Hopi language, Hopi: ''katsina'' , plural ''katsinim'' ) is a spirit being in the religious beliefs of the Pueblo people, Native Americans in the United States, Native American cultures located in the south-western part of the Unite ...
Pawnee Pawnee initially refers to a Native American people and its language:
* Pawnee people
* Pawnee language
Pawnee is also the name of several places in the United States:
* Pawnee, Illinois
* Pawnee, Kansas
* Pawnee, Missouri
* Pawnee City, Nebraska
* ...
Ute
Ute or UTE may refer to:
* Ute people, a Native American people of the Great Basin
* Ute Indian Tribe of the Uintah and Ouray Reservation, Utah
* Ute Mountain Ute Tribe, Colorado, New Mexico, and Utah
* Southern Ute Indian Tribe of the Southern ...
portray the coyote as the companion of The Creator. A Tohono O'odham
flood story
A flood myth or a deluge myth is a myth in which a great flood, usually sent by a deity or deities, destroys civilization, often in an act of divine retribution. Parallels are often drawn between the flood waters of these myths and the primeval ...
has Coyote helping Montezuma survive a global deluge that destroys humanity. After The Creator creates humanity, Coyote and Montezuma teach people how to live. The
Crow
A crow is a bird of the genus ''Corvus'', or more broadly, a synonym for all of ''Corvus''. The word "crow" is used as part of the common name of many species. The related term "raven" is not linked scientifically to any certain trait but is rathe ...
creation story portrays Old Man Coyote as The Creator. In The Dineh creation story, Coyote was present in the First World with First Man and First Woman, though a different version has it being created in the Fourth World. The Navajo Coyote brings death into the world, explaining that without death, too many people would exist, thus no room to plant corn.
Prior to the
Spanish conquest of the Aztec Empire
The Spanish conquest of the Aztec Empire was a pivotal event in the history of the Americas, marked by the collision of the Aztec Triple Alliance and the Spanish Empire. Taking place between 1519 and 1521, this event saw the Spanish conquistad ...
, Coyote played a significant role in Mesoamerican cosmology. The coyote symbolized military might in Classic era
Teotihuacan
Teotihuacan (; Spanish language, Spanish: ''Teotihuacán'', ; ) is an ancient Mesoamerican city located in a sub-valley of the Valley of Mexico, which is located in the State of Mexico, northeast of modern-day Mexico City.
Teotihuacan is ...
, with warriors dressing up in coyote costumes to call upon its predatory power. The species continued to be linked to Central Mexican warrior cults in the centuries leading up to the post-Classic Aztec rule.Schwartz, M. (1998). ''A History of Dogs in the Early Americas''. Yale University Press. pp. 146–149. .
In
Aztec mythology
Aztec mythology is the body or collection of myths of the Aztec civilization of Central Mexico. The Aztecs were a culture living in central Mexico and much of their mythology is similar to that of other Mesoamerican cultures. According to legend ...
, Huehuecóyotl (meaning "old coyote"), the god of dance, music and carnality, is depicted in several codices as a man with a coyote's head. He is sometimes depicted as a womanizer, responsible for bringing war into the world by seducing Xochiquetzal, the goddess of love.
Epigrapher
Epigraphy () is the study of inscriptions, or epigraphs, as writing; it is the science of identifying graphemes, clarifying their meanings, classifying their uses according to dates and cultural contexts, and drawing conclusions about the wr ...
David H. Kelley argued that the god Quetzalcoatl owed its origins to pre-Aztec
Uto-Aztecan
The Uto-Aztecan languages are a family of native American languages, consisting of over thirty languages. Uto-Aztecan languages are found almost entirely in the Western United States and Mexico. The name of the language family reflects the common ...
mythological depictions of the coyote, which is portrayed as mankind's "Elder Brother", a creator, seducer, trickster, and culture hero linked to the morning star.
Attacks on humans
Coyote attacks on humans are uncommon and rarely cause serious injuries, due to the relatively small size of the coyote, but have been increasingly frequent, especially in
California
California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. By the middle of the 19th century, the coyote was already marked as an enemy by humans. (Sharp & Hall, 1978 Pg. 41-54) There have been only two confirmed fatal attacks: one on three-year-old Kelly Keen in
Glendale, California
Glendale is a city located primarily in the Verdugo Mountains region, with a small portion in the San Fernando Valley, of Los Angeles County, California, United States. It is located about north of downtown Los Angeles.
As of 2024, Glendale ha ...
and another on nineteen-year-old singer-songwriter Taylor Mitchell in
Nova Scotia
Nova Scotia is a Provinces and territories of Canada, province of Canada, located on its east coast. It is one of the three Maritime Canada, Maritime provinces and Population of Canada by province and territory, most populous province in Atlan ...
, Canada. In the 30 years leading up to March 2006, at least 160 attacks occurred in the United States, mostly in the
Los Angeles County
Los Angeles County, officially the County of Los Angeles and sometimes abbreviated as LA County, is the most populous county in the United States, with 9,663,345 residents estimated in 2023. Its population is greater than that of 40 individua ...
area. Data from
United States Department of Agriculture
The United States Department of Agriculture (USDA) is an executive department of the United States federal government that aims to meet the needs of commercial farming and livestock food production, promotes agricultural trade and producti ...
(USDA) Wildlife Services, the California Department of Fish and Game, and other sources show that while 41 attacks occurred during the period of 1988–1997, 48 attacks were verified from 1998 through 2003. The majority of these incidents occurred in Southern California near the suburban-wildland interface.
In the absence of the harassment of coyotes practiced by rural people, urban coyotes are losing their fear of humans, which is further worsened by people intentionally or unintentionally feeding coyotes. In such situations, some coyotes have begun to act aggressively toward humans, chasing joggers and bicyclists, confronting people walking their dogs, and stalking small children. Albeit rarely, coyotes in these areas have targeted small children, mostly under the age of 10, though some adults have been bitten.
Although media reports of such attacks generally identify the animals in question as simply "coyotes", research into the genetics of the eastern coyote indicates those involved in attacks in northeast North America, including Pennsylvania, New York, New England, and eastern Canada, may have actually been coywolves, hybrids of ''Canis latrans'' and ''C. lupus,'' not fully coyotes.
Livestock and pet predation
, coyotes were the most abundant livestock predators in western North America, causing the majority of sheep, goat, and cattle losses. For example, according to the
National Agricultural Statistics Service
The National Agricultural Statistics Service (NASS) is the statistical branch of the U.S. Department of Agriculture and a principal agency of the U.S. Federal Statistical System. NASS has 12 regional offices throughout the United States and P ...
, coyotes were responsible for 60.5% of the 224,000 sheep deaths attributed to predation in 2004. The total number of sheep deaths in 2004 comprised 2.22% of the total sheep and lamb population in the United States, which, according to the National Agricultural Statistics Service USDA report, totaled 4.66 million and 7.80 million heads respectively as of July 1, 2005.
Because coyote populations are typically many times greater and more widely distributed than those of wolves, coyotes cause more overall predation losses. United States government agents routinely shoot, poison, trap, and kill about 90,000 coyotes each year to protect livestock. An Idaho census taken in 2005 showed that individual coyotes were 5% as likely to attack livestock as individual wolves. In Utah, more than 11,000 coyotes were killed for bounties totaling over $500,000 in the fiscal year ending June 30, 2017.Livestock guardian dogs are commonly used to aggressively repel predators and have worked well in both fenced pasture and range operations. A 1986 survey of sheep producers in the USA found that 82% reported the use of dogs represented an economic asset.
Re-wilding cattle, which involves increasing the natural protective tendencies of cattle, is a method for controlling coyotes discussed by Temple Grandin of Colorado State University. This method is gaining popularity among producers who allow their herds to calve on the range and whose cattle graze open pastures throughout the year.
Coyotes typically bite the throat just behind the jaw and below the ear when attacking adult sheep or goats, with death commonly resulting from suffocation. Blood loss is usually a secondary cause of death. Calves and heavily fleeced sheep are killed by attacking the flanks or hindquarters, causing shock and blood loss. When attacking smaller prey, such as young lambs, the kill is made by biting the skull and spinal regions, causing massive tissue and bone damage. Small or young prey may be completely carried off, leaving only blood as evidence of a kill. Coyotes usually leave the hide and most of the skeleton of larger animals relatively intact, unless food is scarce, in which case they may leave only the largest bones. Scattered bits of wool, skin, and other parts are characteristic where coyotes feed extensively on larger carcasses.
Tracks are an important factor in distinguishing coyote from dog predation. Coyote tracks tend to be more oval-shaped and compact than those of domestic dogs, and their claw marks are less prominent and the tracks tend to follow a straight line more closely than those of dogs. With the exception of
sighthound
Sighthounds (also called gazehounds) are a Dog type, type of hound dog that hunts primarily by sight and speed, unlike scent hound, scent hounds, which rely on scent and endurance.
Appearance
These dogs specialize in pursuing prey, keeping it ...
s, most dogs of similar weight to coyotes have a slightly shorter stride. Coyote kills can be distinguished from wolf kills by less damage to the underlying tissues in the former. Also, coyote scat tends to be smaller than wolf scat.
Coyotes are often attracted to
dog food
Dog food is specifically formulated food intended for consumption by dogs and other related canines. Dogs are considered to be omnivores with a carnivorous bias. They have the sharp, pointed teeth and shorter gastrointestinal tracts of carn ...
and animals that are small enough to appear as prey. Items such as garbage, pet food, and sometimes feeding stations for birds and squirrels attract coyotes into backyards. About three to five pets attacked by coyotes are brought into the Animal Urgent Care hospital of South Orange County (California) each week, the majority of which are dogs, since cats typically do not survive the attacks. Scat analysis collected near
Claremont, California
Claremont () is a suburban city in eastern Los Angeles County, California, United States, east of Los Angeles. It lies in the Pomona Valley at the foothills of the San Gabriel Mountains. As of the 2010 United States Census, 2010 census it had ...
, revealed that coyotes relied heavily on pets as a food source in winter and spring.
At one location in Southern California, coyotes began relying on a colony of
feral cat
A feral cat or a stray cat is an unowned domestic cat (''Felis catus'') that lives outdoors and avoids human contact; it does not allow itself to be handled or touched, and usually remains hidden from humans. Feral cats may breed over dozens ...
s as a food source. Over time, the coyotes killed most of the cats and then continued to eat the cat food placed daily at the colony site by people who were maintaining the cat colony.
Coyotes usually attack smaller-sized dogs, but they have been known to attack even large, powerful breeds such as the Rottweiler in exceptional cases. Dogs larger than coyotes, such as
greyhound
The English Greyhound, or simply the Greyhound, is a dog breed, breed of dog, a sighthound which has been bred for coursing, greyhound racing and hunting. Some are kept as show dogs or pets.
Greyhounds are defined as a tall, muscular, smooth-c ...
s, are generally able to drive them off and have been known to kill coyotes. Smaller breeds are more likely to suffer injury or death.
Hunting
Coyote hunting is one of the most common forms of predator hunting that humans partake in. There are not many regulations with regard to the taking of the coyote which means there are many different methods that can be used to hunt the animal. The most common forms are
trapping
Animal trapping, or simply trapping or ginning, is the use of a device to remotely catch and often kill an animal. Animals may be trapped for a variety of purposes, including for meat, fur trade, fur/feathers, sport hunting, pest control, and w ...
hound
A hound is a type of hunting dog used by hunters to track or chase prey.
Description
Hounds can be contrasted with gun dogs that assist hunters by identifying prey and/or recovering shot quarry. The hound breeds were the first hunting dogs. ...
hunting. Since coyotes are colorblind, seeing only in shades of gray and subtle blues, open camouflages and plain patterns can be used. As the average male coyote weighs 8 to 20 kg (18 to 44 lbs) and the average female coyote 7 to 18 kg (15 to 40 lbs), a universal projectile that can perform between those weights is the .223 Remington, so that the projectile expands in the target after entry, but before the exit, thus delivering the most energy.
Coyotes being the light and agile animals they are, they often leave a very light impression on terrain. The coyote's footprint is oblong, approximately 6.35 cm (2.5-inches) long and 5.08 cm (2-inches) wide. There are four claws in both their front and hind paws. The coyote's center pad is relatively shaped like that of a rounded triangle. Like the domestic dog the coyote's front paw is slightly larger than the hind paw. The coyote's paw is most similar to that of the domestic dog.
The hunting of coyotes often results in grey wolves being shot in places where the two species still coexist, as a result of mistaken identity.
Fur uses
Prior to the mid-19th century, coyote fur was considered worthless. This changed with the diminution of
beaver
Beavers (genus ''Castor'') are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver (''Castor canadensis'') and the Eurasian beaver (''C. fiber''). Beavers are the second-large ...
s, and by 1860, the hunting of coyotes for their fur became a great source of income (75 cents to $1.50 per skin) for wolfers in the
Great Plains
The Great Plains is a broad expanse of plain, flatland in North America. The region stretches east of the Rocky Mountains, much of it covered in prairie, steppe, and grassland. They are the western part of the Interior Plains, which include th ...
. Coyote pelts were of significant economic importance during the early 1950s, ranging in price from $5 to $25 per pelt, depending on locality. The coyote's fur is not durable enough to make rugs, but can be used for coats and jackets, scarves, or muffs. The majority of pelts are used for making trimmings, such as coat collars and sleeves for women's clothing. Coyote fur is sometimes dyed black as imitation silver fox.
Coyotes were occasionally eaten by trappers and mountain men during the western expansion. Coyotes sometimes featured in the feasts of the
Plains Indians
Plains Indians or Indigenous peoples of the Great Plains and Canadian Prairies are the Native American tribes and First Nations peoples who have historically lived on the Interior Plains (the Great Plains and Canadian Prairies) of North ...
, and coyote pups were eaten by the indigenous people of
San Gabriel, California
San Gabriel (Spanish language, Spanish for "Gabriel, St. Gabriel") is a city located in the San Gabriel Valley of Los Angeles County, California. At the 2020 United States census, 2020 census, the population was 39,568.
San Gabriel was founded ...
. The taste of coyote meat has been likened to that of the wolf and is more tender than
pork
Pork is the culinary name for the meat of the pig (''Sus domesticus''). It is the most commonly consumed meat worldwide, with evidence of pig animal husbandry, husbandry dating back to 8000–9000 BCE.
Pork is eaten both freshly cooke ...
when boiled. Coyote fat, when taken in the fall, has been used on occasion to grease leather or eaten as a spread.
Tameability
Coyotes were likely semidomesticated by various pre-Columbian cultures. Some 19th-century writers wrote of coyotes being kept in native villages in the Great Plains. The coyote is easily tamed as a pup, but can become destructive as an adult. Both full-blooded and hybrid coyotes can be playful and confiding with their owners, but are suspicious and shy of strangers, though coyotes being tractable enough to be used for practical purposes like retrieving and pointing have been recorded. A tame coyote named "Butch", caught in the summer of 1945, had a short-lived career in cinema, appearing in '' Smoky'' (1946) and ''
Ramrod
A ramrod (or scouring stick) is a metal or wooden device used with muzzleloader, muzzleloading firearms to push the projectile up against the propellant (mainly blackpowder). The ramrod was used with weapons such as muskets and cannons and was u ...
'' (1947) before being shot while raiding a henhouse.
Looney Tunes
''Looney Tunes'' is an American media franchise produced and distributed by Warner Bros. The franchise began as a series of animated short films that originally ran from 1930 to 1969, alongside its spin-off series ''Merrie Melodies'', during t ...
'' and ''
Merrie Melodies
''Merrie Melodies'' is an American animated comedy short film series distributed by Warner Bros. Pictures. It was part of the ''Looney Tunes'' franchise and featured many of the same characters. Originally running from August 2, 1931, to Septem ...
'' series of animated short films, in which he makes numerous ill-fated attempts to capture an elusive roadrunner.
*Dag and his coyote pack are the main antagonists in Nickelodeon's 2006 animated film '' Barnyard''.
*The
NHL
The National Hockey League (NHL; , ''LNH'') is a professional ice hockey league in North America composed of 32 teams25 in the United States and 7 in Canada. The NHL is one of the major professional sports leagues in the United States and Cana ...
team in
Arizona
Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
(1996–2024) was named the
Arizona Coyotes
The Arizona Coyotes are an inactive professional ice hockey team based in the Phoenix metropolitan area. They competed in the National Hockey League (NHL) as a member of the Central Division (NHL), Central Division (1996–1998, 2021–2024) and ...
to pay tribute to the large population of coyotes in the region. The team's official suited mascot was Howler.
*The famous ''oo-wee-oo-wee-oo wah-wah-wah'' scream in ''
The Good, The Bad and The Ugly
''The Good, the Bad and the Ugly'' (, literally "''The good, the ugly, the bad''") is a 1966 Italian epic spaghetti Western film directed by Sergio Leone and starring Clint Eastwood as "the Good", Lee Van Cleef as "the Bad", and Eli Wallach a ...
'' (1966) was inspired by the howl of the coyote.
*Copper, a coyote, was one of three mascots for the
2002 Winter Olympics
The 2002 Winter Olympics, officially the XIX Olympic Winter Games and commonly known as Salt Lake 2002 (; Gosiute dialect, Gosiute Shoshoni: ''Tit'-so-pi 2002''; ; Shoshoni language, Shoshoni: ''Soónkahni 2002''), were an international wi ...
NBA
The National Basketball Association (NBA) is a professional basketball league in North America composed of 30 teams (29 in the United States and 1 in Canada). The NBA is one of the major professional sports leagues in the United States and Ca ...
's
San Antonio Spurs
The San Antonio Spurs are an American professional basketball team based in San Antonio. The Spurs compete in the National Basketball Association (NBA) as a member of the Southwest Division (NBA), Southwest Division of the Western Conference (NBA ...
. The mascot made his debut in 1983 and was inducted in to the
Mascot Hall of Fame
The Mascot Hall of Fame was an interactive children's museum for sports mascots between 2018 and 2024. It was founded by David Raymond, who was the original Phillie Phanatic from 1978 to 1993.
Each year (beginning in 2005), mascots are elected ...
in 2007.
*An animated coyote voiced by
Johnny Cash
John R. Cash (born J. R. Cash; February 26, 1932 – September 12, 2003) was an American singer-songwriter. Most of his music contains themes of sorrow, moral tribulation, and redemption, especially songs from the later stages of his career. ...
plays a pivotal role as a spirit guide to Homer Simpson in the '' Simpsons'' episode " El Viaje Misterioso de Nuestro Jomer".
*The West Canaan Coyotes are the fictional high school Varsity football team in the 1999 sports comedy-drama film '' Varsity Blues''; the film focuses on players of the team under pressure of a ruthless coach.
*The 2013 documentary film '' Bad Coyote'' profiles the expansion of coyotes into
Atlantic Canada
Atlantic Canada, also called the Atlantic provinces (), is the list of regions of Canada, region of Eastern Canada comprising four provinces: New Brunswick, Newfoundland and Labrador, Nova Scotia, and Prince Edward Island. As of 2021, the landma ...
SaltWire Network
SaltWire Network Inc. was a Canadian newspaper publishing company. The company was formed in Halifax, Nova Scotia on April 13, 2017, via its purchase of 27 newspapers from Transcontinental (company), Transcontinental. The company owned 23 daily a ...
Aeon
The word aeon , also spelled eon (in American and Australian English), originally meant "life", "vital force" or "being", "generation" or "a period of time", though it tended to be translated as "age" in the sense of "ages", "forever", "timele ...
''.
Audiobooks
* Olson, Jack (May 2015). ''The Last Coyote'' (8 hours). Narrated by Gary MacFadden. Originally published as ''Slaughter the Animals, Poison the Earth'', Simon & Schuster, Oct. 11, 1971. .
, Wolf and Coyote DNA Bank @ Trent University
* View occurrences of Canis latrans ' in the
Biodiversity Heritage Library
The Biodiversity Heritage Library (BHL) is the world’s largest open-access digital library for biodiversity literature and archives. BHL operates as a worldwide consortium of natural history, botanical, research, and national libraries working ...