Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s commonly known as the mustards, the crucifers, or the cabbage family. Most are

herbaceous plant Herbaceous plants are vascular plants that have no persistent wood, woody stems above ground. This broad category of plants includes many perennial plant, perennials, and nearly all Annual plant, annuals and Biennial plant, biennials. Definition ...

s, while some are shrubs. The leaves are simple (although are sometimes deeply incised), lack

stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...

s, and appear alternately on stems or in rosettes. The

inflorescence An inflorescence is a group or cluster of flowers arranged on a Plant stem, stem that is composed of a main branch or a complicated arrangement of branches. Morphology (biology), Morphologically, it is the modified part of the shoot of sperma ...

s are terminal and lack bracts. The flowers have four free

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coine ...

s, four free alternating petals, two shorter free stamens and four longer free stamens. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

has seeds in rows, divided by a thin wall (or septum). The family contains 372 genera and 4,060 accepted

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

. The largest genera are ''

Draba ''Draba'' is a large genus of flowering plants in the family Brassicaceae, commonly known as whitlow-grasses (though they are not related to the true grasses). Species There are over 400 species: *'' Draba abajoensis'' Windham & Al-Shehbaz *'' ...

'' (440 species), ''

Erysimum ''Erysimum'', or wallflower, is a genus of flowering plants in the cabbage family, Brassicaceae. It includes more than 150 species, both popular garden plants and many wild forms. The genus ''Cheiranthus'' is sometimes included here in whole o ...

'' (261 species), ''

Lepidium ''Lepidium'' is a genus of plants in the mustard/cabbage family, Brassicaceae. The genus is widely distributed in the Americas, Africa, Asia, Europe, and Australia.Cardamine ''Cardamine'' is a large genus of flowering plants in the mustard family, Brassicaceae, known as bittercresses and toothworts. It contains more than 200 species of annuals and perennials. Species in this genus can be found worldwide, except the ...

'' (233 species), and ''

Alyssum ''Alyssum'' is a genus of over a hundred species of flowering plants in the family Brassicaceae, native to Europe, Asia, and northern Africa, with the highest species diversity in the Mediterranean region. The genus comprises annual and perenni ...

'' (207 species). The family contains the

cruciferous vegetables Cruciferous vegetables are vegetables of the family Brassicaceae (also called Cruciferae) with many genera, species, and cultivars being raised for food production such as cauliflower, cabbage, kale, garden cress, bok choy, broccoli, Brusse ...

, including species such as ''

Brassica oleracea ''Brassica oleracea'' is a plant species from family Brassicaceae that includes many common cultivars used as vegetables, such as cabbage, broccoli, cauliflower, kale, Brussels sprouts, collard greens, Savoy cabbage, kohlrabi, and gai lan. ...

'' (cultivated as cabbage, kale, cauliflower,

broccoli Broccoli (''Brassica oleracea'' var. ''italica'') is an edible green plant in the cabbage family (family Brassicaceae, genus ''Brassica'') whose large flowering head, stalk and small associated leaves are eaten as a vegetable. Broccoli is cla ...

and

collards Collard is a group of certain loose-leafed cultivars of '' Brassica oleracea'', the same species as many common vegetables including cabbage ( Capitata group) and broccoli ( Italica group). Collard is a member of the Viridis group of ''Brassic ...

), ''

Brassica rapa ''Brassica rapa'' is a plant species growing in various widely cultivated forms including the turnip (a root vegetable); napa cabbage, bomdong, bok choy, and rapini. ''Brassica rapa'' subsp. ''oleifera'' is an oilseed which has many common n ...

'' (

turnip The turnip or white turnip ('' Brassica rapa'' subsp. ''rapa'') is a root vegetable commonly grown in temperate climates worldwide for its white, fleshy taproot. The word ''turnip'' is a compound of ''turn'' as in turned/rounded on a lathe and ...

Chinese cabbage Chinese cabbage (''Brassica rapa'', subspecies ''pekinensis'' and ''chinensis'') can refer to two cultivar groups of leaf vegetables often used in Chinese cuisine: the Pekinensis Group (napa cabbage) and the Chinensis Group (bok choy). These ...

, etc.), '' Brassica napus'' (

rapeseed Rapeseed (''Brassica napus ''subsp.'' napus''), also known as rape, or oilseed rape, is a bright-yellow flowering member of the family Brassicaceae (mustard or cabbage family), cultivated mainly for its oil-rich seed, which naturally contains a ...

, etc.), ''

Raphanus sativus The radish (''Raphanus raphanistrum'' subsp. ''sativus'') is an edible root vegetable of the family Brassicaceae that was domesticated in Asia prior to Roman times. Radishes are grown and consumed throughout the world, being mostly eaten raw ...

'' (common

radish The radish (''Raphanus raphanistrum'' subsp. ''sativus'') is an edible root vegetable of the family Brassicaceae that was domesticated in Asia prior to Roman times. Radishes are grown and consumed throughout the world, being mostly eaten raw ...

), ''

Armoracia rusticana Horseradish (''Armoracia rusticana'', syn. ''Cochlearia armoracia'') is a perennial plant of the family Brassicaceae (which also includes mustard, wasabi, broccoli, cabbage, and radish). It is a root vegetable, cultivated and used worldwide a ...

'' (

horseradish Horseradish (''Armoracia rusticana'', syn. ''Cochlearia armoracia'') is a perennial plant of the family Brassicaceae (which also includes mustard, wasabi, broccoli, cabbage, and radish). It is a root vegetable, cultivated and used worldwi ...

), but also a cut-flower ''

Matthiola ''Matthiola'' () is a genus of flowering plant in the mustard family Brassicaceae. It is named after Italian naturalist Pietro Andrea Mattioli (1501–1577). The genus contains about 50 species of annual, biennial and perennial herbaceous plants ...

'' (stock) and the model organism '' Arabidopsis thaliana'' (thale cress). ''

Pieris rapae ''Pieris rapae'' is a small- to medium-sized butterfly species of the whites-and-yellows family Pieridae. It is known in Europe as the small white, in North America as the cabbage white or cabbage butterfly, on several continents as the small c ...

'' and other butterflies of the family

Pieridae The Pieridae are a large family of butterflies with about 76 genera containing about 1,100 species, mostly from tropical Africa and tropical Asia with some varieties in the more northern regions of North America and Eurasia.DeVries P. J. in Le ...

are some of the best-known pests of Brassicaceae species planted as commercial crops. ''Trichoplusia ni'' (

cabbage looper The cabbage looper (''Trichoplusia ni'') is a medium-sized moth in the family Noctuidae, a family commonly referred to as owlet moths. Its common name comes from its preferred host plants and distinctive crawling behavior. Cruciferous vegetable ...

) moth is also becoming increasingly problematic for crucifers due to its resistance to commonly used

pest control Pest control is the regulation or management of a species defined as a pest; any animal, plant or fungus that impacts adversely on human activities or environment. The human response depends on the importance of the damage done and will range ...

methods. Some rarer ''Pieris'' butterflies, such as '' P. virginiensis'', depend upon native mustards for their survival in their native habitats. Some non-native mustards such as ''

Alliaria petiolata ''Alliaria petiolata'', or garlic mustard, is a biennial flowering plant in the mustard family (Brassicaceae). It is native to Europe, western and central Asia, north-western Africa, Morocco, Iberia and the British Isles, north to northern S ...

'' (garlic mustard), an extremely invasive species in the United States, can be toxic to their

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. ...

Description

Species belonging to the Brassicaceae are mostly

annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year ** Yearbook ** Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), ...

biennial Biennial means (an event) lasting for two years or occurring every two years. The related term biennium is used in reference to a period of two years. In particular, it can refer to: * Biennial plant, a plant which blooms in its second year and th ...

, or

perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wid ...

s, some are dwarf shrubs or shrubs, and very few

vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themsel ...

s. Although generally terrestrial, a few species such as water awlwort live submerged in fresh water. They may have a

taproot A taproot is a large, central, and dominant root from which other roots sprout laterally. Typically a taproot is somewhat straight and very thick, is tapering in shape, and grows directly downward. In some plants, such as the carrot, the taproo ...

or a sometimes woody caudex that may have few or many branches, some have thin or tuberous rhizomes, or rarely develop runners. Few species have multi-cellular glands.

Hairs Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and f ...

consist of one cell and occur in many forms: from simple to forked, star-, tree- or T-shaped, rarely taking the form of a shield or scale. They are never topped by a gland. The stems may be upright, rise up towards the tip, or lie flat, are mostly herbaceous but sometimes woody. Stems carry leaves or the stems may be leafless (in ''

Caulanthus ''Caulanthus'' is a genus of plants in the family Brassicaceae. Plants of this genus may be known as jewelflowers. They are also often referred to as wild cabbage, although this common name usually refers to wild variants of ''Brassica oleraceae' ...

''), and some species lack stems altogether. The leaves do not have

s, but there may be a pair of glands at base of leafstalks and flowerstalks. The leaf may be seated or have a leafstalk. The

leaf blade A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...

is usually simple, entire or

dissected Dissection (from Latin ' "to cut to pieces"; also called anatomization) is the dismembering of the body of a deceased animal or plant to study its anatomical structure. Autopsy is used in pathology and forensic medicine to determine the cause ...

, rarely

trifoliolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

or pinnately compound. A leaf rosette at the base may be present or absent. The leaves along the stem are almost always alternately arranged, rarely apparently opposite. The stomata are of the anisocytic type. The

genome size Genome size is the total amount of DNA contained within one copy of a single complete genome. It is typically measured in terms of mass in picograms (trillionths (10−12) of a gram, abbreviated pg) or less frequently in daltons, or as the total ...

of Brassicaceae compared to that of other Angiosperm families is very small to small (less than 3.425 million base pairs per cell), varying from 150 Mbp in '' Arabidopsis thaliana'' and '' Sphaerocardamum'' spp., to 2375 Mbp '' Bunias orientalis''. The number of homologous chromosome sets varies from four (n=4) in some ''

Physaria ''Physaria'' is a genus of flowering plants in the family Brassicaceae. Many species are known generally as twinpods, bladderpods, or lesquerella. They are native to the Americas, with many species endemic to western North America. They are dense ...

'' and '' Stenopetalum'' species, five (n=5) in other ''Physaria'' and ''Stenopetalum'' species, ''Arabidopsis thaliana'' and a '' Mathiola'' species, to seventeen (n=17). About 35% of the species in which chromosomes have been counted have eight sets (n=8). Due to

polyploidy Polyploidy is a condition in which the biological cell, cells of an organism have more than one pair of (Homologous chromosome, homologous) chromosomes. Most species whose cells have Cell nucleus, nuclei (eukaryotes) are diploid, meaning they ha ...

, some species may have up to 256 individual chromosomes, with some very high counts in the North American species of ''Cardamine'', such as '' C. diphylla''. Hybridisation is not unusual in Brassicaceae, especially in ''

Arabis ''Arabis'' ,''Sunset Western Garden Book,'' 1995:606–607 or rockcress, is a genus of flowering plants, within the family Brassicaceae. Description The species are herbaceous, annual or perennial plants, growing to 10–80 cm tall, usua ...

'', ''

Rorippa ''Rorippa'' is a genus of flowering plants in the family Brassicaceae, native to Europe through central Asia, Africa, and North America. ''Rorippa'' species are annual to perennial herbs, usually with yellow flowers and a peppery flavour. They a ...

'', ''

Cardamine ''Cardamine'' is a large genus of flowering plants in the mustard family, Brassicaceae, known as bittercresses and toothworts. It contains more than 200 species of annuals and perennials. Species in this genus can be found worldwide, except the ...

'' and ''

Boechera ''Boechera'' (rockcress) is a genus of the family Brassicaceae. It was named after the Danish botanist Tyge W. Böcher (1909–1983), who was known for his research in alpine plants, including the mustards ''Draba'' and ''Boechera holboellii''. ...

''. Hybridisation between species originating in Africa and California, and subsequent polyploidisation is surmised for ''

Lepidium ''Lepidium'' is a genus of plants in the mustard/cabbage family, Brassicaceae. The genus is widely distributed in the Americas, Africa, Asia, Europe, and Australia.racemes, panicles, or

corymb Corymb is a botanical term for an inflorescence with the flowers growing in such a fashion that the outermost are borne on longer pedicels than the inner, bringing all flowers up to a common level. A corymb has a flattish top with a superficial ...

s, with pedicels sometimes in the axil of a bract, and few species have flowers that sit individually on flower stems that spring from the axils of rosette leaves. The orientation of the pedicels when fruits are ripe varies dependent on the species. The flowers are bisexual, star symmetrical (zygomorphic in ''

Iberis ''Iberis'' , commonly called candytuft, is a genus of flowering plants belonging to the family Brassicaceae. It comprises annuals, evergreen perennials and subshrubs native to the Old World. The name "candytuft" is not related to candy, but der ...

'' and '' Teesdalia'') and the ovary positioned above the other floral parts. Each flower has four free or seldom merged

s, the lateral two sometimes with a shallow spur, which are mostly shed after flowering, rarely persistent, may be reflexed, spreading, ascending, or erect, together forming a tube-, bell- or urn-shaped calyx. Each flower has four petals, set alternating with the sepals, although in some species these are rudimentary or absent. They may be differentiated into a

blade A blade is the portion of a tool, weapon, or machine with an edge that is designed to puncture, chop, slice or scrape surfaces or materials. Blades are typically made from materials that are harder than those they are to be used on. Histor ...

and a

claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...

or not, and consistently lack basal appendages. The blade is entire or has an indent at the tip, and may sometimes be much smaller than the claws. The mostly six stamens are set in two whorls: usually the two lateral, outer ones are shorter than the four inner stamens, but very rarely the stamens can all have the same length, and very rarely species have different numbers of stamens such as sixteen to twenty four in '' Megacarpaea'', four in '' Cardamine hirsuta'', and two in '' Coronopus''. The filaments are slender and not fused, while the anthers consist of two pollen producing cavities, and open with longitudinal slits. The pollen grains are

tricolpate The eudicots, Eudicotidae, or eudicotyledons are a clade of flowering plants mainly characterized by having two seed leaves upon germination. The term derives from Dicotyledons. Traditionally they were called tricolpates or non-magnoliid dic ...

. The receptacle carries a variable number of nectaries, but these are always present opposite the base of the lateral stamens.

Ovary, fruit and seed

There is one superior pistil that consists of two

carpel Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils' ...

s that may either sit directly above the base of the stamens or on a stalk. It initially consists of only one cavity but during its further development a thin wall grows that divides the cavity, both placentas and separates the two valves (a so-called false septum). Rarely, there is only one cavity without a septum. The 2–600

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the ...

s are usually along the side margin of the carpels, or rarely at the top. Fruits are capsules that open with two valves, usually towards the top. These are called

silique A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit i ...

if at least three times longer than wide, or

silicle A silique or siliqua (plural ''siliques'' or ''siliquae'') is a type of fruit (seed capsule) having two fused carpels with the length being more than three times the width. When the length is less than three times the width of the dried fruit i ...

if the length is less than three times the width. The fruit is very variable in its other traits. There may be one persistent

style Style is a manner of doing or presenting things and may refer to: * Architectural style, the features that make a building or structure historically identifiable * Design, the process of creating something * Fashion, a prevailing mode of clothing ...

that connects the ovary to the globular or conical stigma, which is undivided or has two spreading or connivent lobes. The variously shaped seeds are usually yellow or brown in color, and arranged in one or two rows in each cavity. The

seed leaves A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The num ...

are entire or have a notch at the tip. The seed does not contain endosperm.

Differences with similar families

Brassicaceae have a bisymmetical corolla (left is mirrored by right, stem-side by out-side, but each quarter is not symmetrical), a septum dividing the fruit, lack

s and have simple (although sometimes deeply incised) leaves. The sister family

Cleomaceae The Cleomaceae are a small family of flowering plants in the order Brassicales, comprising about 300 species in 10 genera, or about 150 species in 17 genera.Cleomaceae, Zhang Mingli (张明理)1; Gordon C. Tucker2, Harvard.edu/ref> These genera ...

has bilateral symmetrical corollas (left is mirrored by right, but stem-side is different from out-side), stipules and mostly palmately divided leaves, and mostly no septum. Capparaceae generally have a

gynophore A gynophore is the stalk of certain flowers which supports the gynoecium (the ovule-producing part of a flower), elevating it above the branching points of other floral parts. Plant genera that have flowers with gynophores include '' Telopea'', '' ...

, sometimes an androgynophore, and a variable number of stamens.

Phytochemistry

Almost all Brassicaceae have

C3 carbon fixation carbon fixation is the most common of three metabolic pathways for carbon fixation in photosynthesis, along with C4 carbon fixation, and Crassulacean acid metabolism, CAM. This process converts carbon dioxide and ribulose bisphosphate (RuBP, a ...

. The only exceptions are a few '' Moricandia'' species, which have a hybrid system between C3 and C4 carbon fixation, C4 fixation being more efficient in drought, high temperature and low nitrate availability. Brassicaceae contain different cocktails of dozens of glucosinolates. They also contain enzymes called

myrosinase Myrosinase (, ''thioglucoside glucohydrolase'', ''sinigrinase'', and ''sinigrase'') is a family of enzymes involved in plant defense against herbivores, specifically the mustard oil bomb. The three-dimensional structure has been elucidated and is ...

s, that convert the glucosinolates into

isothiocyanate In organic chemistry, isothiocyanate is the functional group , formed by substituting the oxygen in the isocyanate group with a sulfur. Many natural isothiocyanates from plants are produced by enzymatic conversion of metabolites called glucosi ...

thiocyanate Thiocyanate (also known as rhodanide) is the anion . It is the conjugate base of thiocyanic acid. Common derivatives include the colourless salts potassium thiocyanate and sodium thiocyanate. Mercury(II) thiocyanate was formerly used in pyr ...

s and nitriles, which are toxic to many organisms, and so help guard against herbivory.

Taxonomy

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...

in 1753 regarded the Brassicaceae as a natural group, naming them "Klass" Tetradynamia. Alfred Barton Rendle placed the family in the order Rhoedales, while

George Bentham George Bentham (22 September 1800 – 10 September 1884) was an English botanist, described by the weed botanist Duane Isely as "the premier systematic botanist of the nineteenth century". Born into a distinguished family, he initially studie ...

and

Joseph Dalton Hooker Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For twenty years he served as director of ...

in their system published from 1862–1883, assigned it to their cohort Parietales (now the class

Violales Violales is a botanical name of an order of flowering plants and takes its name from the included family Violaceae; it was proposed by Lindley (1853). The name has been used in several systems, although some systems used the name Parietales for ...

). Following Bentham and Hooker, John Hutchinson in 1948 and again in 1964 thought the Brassicaceae to stem from near the

Papaveraceae The Papaveraceae are an economically important family of about 42 genera and approximately 775 known species of flowering plants in the order Ranunculales, informally known as the poppy family. The family is cosmopolitan, occurring in temperat ...

. In 1994, a group of scientists including Walter Stephen Judd suggested to include the

Capparaceae The Capparaceae (or Capparidaceae), commonly known as the caper family, are a family of plants in the order Brassicales. As currently circumscribed, the family contains 33 genera and about 700 species. The largest genera are ''Capparis'' (about ...

in the Brassicaceae. Early DNA-analysis showed that the Capparaceae—as defined at that moment—were paraphyletic, and it was suggested to assign the genera closest to the Brassicaceae to the

. The Cleomaceae and Brassicaceae diverged approximately 41 million years ago. All three families have consistently been placed in one order (variably called

Capparales Capparales is a botanical name of an order of flowering plants. It was used in the Cronquist system for an order in subclass Dilleniidae and in the Kubitzki system, nowadays. In the 1981 version of this system it included : * order Capparales *: ...

Brassicales The Brassicales (or Cruciales) are an order of flowering plants, belonging to the eurosids II group of dicotyledons under the APG II system. One character common to many members of the order is the production of glucosinolate (mustard oil) compo ...

). The

APG II The APG II system (Angiosperm Phylogeny Group II system) of plant classification is the second, now obsolete, version of a modern, mostly molecular-based, system of plant taxonomy that was published in April 2003 by the Angiosperm Phylogeny Gr ...

system merged Cleomaceae and Brassicaceae. Other classifications have continued to recognize the Capparaceae, but with a more restricted circumscription, either including ''Cleome'' and its relatives in the Brassicaceae or recognizing them in the segregate family

. The APG III system has recently adopted this last solution, but this may change as a consensus arises on this point. Current insights in the relationships of the Brassicaceae, based on a 2012 DNA-analysis, are summarized in the following tree.

Relationships within the family

Early classifications depended on morphological comparison only, but because of extensive

convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...

, these do not provide a reliable

phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...

. Although a substantial effort was made through molecular phylogenetic studies, the relationships within the Brassicaceae have not always been well resolved yet. It has long been clear that the '' Aethionema'' are

sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a family, familial relationship, it is sometimes used endearingly to r ...

of the remainder of the family. One analysis from 2014 represented the relation between 39 tribes with the following tree.

Genera

Version 1 of The Plant List lists 349 genera.

Etymology

The name ''Brassicaceae'' comes to international scientific vocabulary from

New Latin New Latin (also called Neo-Latin or Modern Latin) is the revival of Literary Latin used in original, scholarly, and scientific works since about 1500. Modern scholarly and technical nomenclature, such as in zoological and botanical taxonomy ...

, from ''

Brassica ''Brassica'' () is a genus of plants in the cabbage and mustard family ( Brassicaceae). The members of the genus are informally known as cruciferous vegetables, cabbages, or mustard plants. Crops from this genus are sometimes called ''cole c ...

'', the

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal ...

, + '' -aceae'', a standardized suffix for plant family names in modern taxonomy. The genus name comes from the

Classical Latin Classical Latin is the form of Literary Latin recognized as a literary standard by writers of the late Roman Republic and early Roman Empire. It was used from 75 BC to the 3rd century AD, when it developed into Late Latin. In some later period ...

word ''

brassica ''Brassica'' () is a genus of plants in the cabbage and mustard family ( Brassicaceae). The members of the genus are informally known as cruciferous vegetables, cabbages, or mustard plants. Crops from this genus are sometimes called ''cole c ...

'', referring to cabbage and other

. The alternative older name,

Cruciferae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous plants, while some are shrubs. The leav ...

, meaning "cross-bearing", describes the four petals of

mustard Mustard may refer to: Food and plants * Mustard (condiment), a paste or sauce made from mustard seeds used as a condiment * Mustard plant, one of several plants, having seeds that are used for the condiment ** Mustard seed, seeds of the mustard p ...

flowers, which resemble a

cross A cross is a geometrical figure consisting of two intersecting lines or bars, usually perpendicular to each other. The lines usually run vertically and horizontally. A cross of oblique lines, in the shape of the Latin letter X, is termed a sa ...

. Cruciferae is one of eight plant family names, not derived from a genus name and without the suffix ''-aceae'' that are authorized alternative names.

Distribution

Brassicaceae can be found almost on the entire land surface of the planet, but the family is absent from Antarctica, and also absent from some areas in the tropics i.e. northeastern Brazil, the Congo basin,

Maritime Southeast Asia Maritime Southeast Asia comprises the countries of Brunei, Indonesia, Malaysia, the Philippines, Singapore, and East Timor. Maritime Southeast Asia is sometimes also referred to as Island Southeast Asia, Insular Southeast Asia or Oceanic Sout ...

and tropical

Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologi ...

. The area of origin of the family is possibly the

Irano-Turanian Region The Irano-Turanian Region is a floristic region located within the Tethyan Subkingdom of the Holarctic Kingdom. It is divided into 12 floristic provinces according to the work of Armen Takhtajan, a Soviet-Armenian botanist who created a classifi ...

, where approximately 900 species occur in 150 different genera. About 530 of those 900 species are

endemics Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...

. Next in abundance comes the

Mediterranean Region In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and wa ...

, with around 630 species (290 of which are endemic) in 113 genera. The family is less prominent in the Saharo-Arabian Region—65 genera, 180 species of which 62 are endemic—and North America (comprising the

North American Atlantic Region North American Atlantic Region is a floristic region within the Holarctic Kingdom identified by Armen Takhtajan and Robert F. Thorne, spanning from the Atlantic and Gulf coasts to the Great Plains and comprising a major part of the United States ...

and the Rocky Mountain Floristic Region)—99 genera, 780 species of which 600 are endemic. South-America has 40 genera containing 340 native species, Southern Africa 15 genera with over 100 species, and Australia and New-Zealand have 19 genera with 114 species between them.

Ecology

Brassicaceae are almost exclusively

pollinated by insects Entomophily or insect pollination is a form of pollination whereby pollen of plants, especially but not only of flowering plants, is distributed by insects. Flowers pollinated by insects typically advertise themselves with bright colours, some ...

. A chemical mechanism in the pollen is active in many species to avoid

selfing Autogamy, or self-fertilization, refers to the fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a reproductive mechanism employed by many flowering plants. However, species ...

. Two notable exceptions are exclusive self-pollination in closed flowers in '' Cardamine chenopodifolia'', and wind pollination in ''

Pringlea antiscorbutica ''Pringlea antiscorbutica'', commonly known as Kerguelen cabbage, is a flowering plant and the sole member of the monotypic genus ''Pringlea'' in the family Brassicaceae. Its common name comes from the archipelago of its discovery, the Kerguelen ...

''. Although it can be cross-pollinated, ''

'' (garlic mustard) is

self-fertile Self-incompatibility (SI) is a general name for several genetic mechanisms that prevent self-fertilization in sexually reproducing organisms, and thus encourage outcrossing and allogamy. It is contrasted with separation of sexes among individuals ...

. Most species reproduce sexually through seed, but '' Cardamine bulbifera'' produces gemmae and in others, such as '' Cardamine pentaphyllos'', the coral-like roots easily break into segments, that will grow into separate plants. In some species, such as in the genus ''

'', seed pods open with force and so catapult the seeds quite far. Many of these have sticky seed coats, assisting long distance dispersal by animals, and this may also explain several intercontinental dispersal events in the genus, and its near global distribution. Brassicaceae are common on serpentine and

dolomite Dolomite may refer to: *Dolomite (mineral), a carbonate mineral *Dolomite (rock), also known as dolostone, a sedimentary carbonate rock *Dolomite, Alabama, United States, an unincorporated community *Dolomite, California, United States, an unincor ...

rich in

magnesium Magnesium is a chemical element with the symbol Mg and atomic number 12. It is a shiny gray metal having a low density, low melting point and high chemical reactivity. Like the other alkaline earth metals (group 2 of the periodic ta ...

. Over a hundred species in the family accumulate heavy metals, particularly

zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...

and

nickel Nickel is a chemical element with symbol Ni and atomic number 28. It is a silvery-white lustrous metal with a slight golden tinge. Nickel is a hard and ductile transition metal. Pure nickel is chemically reactive but large pieces are slow ...

, which is a record percentage. Several ''Alyssum'' species can accumulate nickel up to 0.3% of their dry weight, and may be useful in

soil remediation Environmental remediation deals with the removal of pollution or contaminants from environmental media such as soil, groundwater, sediment, or surface water. Remedial action is generally subject to an array of regulatory requirements, and may also ...

or even bio-mining. Brassicaceae contain glucosinolates as well as

s inside their cells. When the cell is damaged, the myrosinases hydrolise the glucosinolates, leading to the synthesis of

s, which are compounds toxic to most animals, fungi and bacteria. Some insect herbivores have developed counter adaptations such as rapid absorption of the glucosinates, quick alternative breakdown into non-toxic compounds and avoiding cell damage. In the whites family (Pieridae), one counter mechanism involves glucosinolate sulphatase, which changes the glucosinolate, so that it cannot be converted to isothiocyanate. A second is that the glucosinates are quickly broken down, forming nitriles. Differences between the mixtures of glucosinolates between species and even within species is large, and individual plants may produce in excess of fifty individual substances. The energy penalty for synthesising all these glucosinolates may be as high as 15% of the total needed to produce a leaf. ''

Barbarea vulgaris ''Barbarea vulgaris'', also called wintercress (usual common name), or alternatively herb barbara, rocketcress, yellow rocketcress, winter rocket, yellow rocket, and wound rocket, is a biennial herb of the genus '' Barbarea'', belonging to the fa ...

'' (bittercress) also produces

triterpenoid saponin Saponins (Latin "sapon", soap + "-in", one of), also selectively referred to as triterpene glycosides, are bitter-tasting usually toxic plant-derived organic chemicals that have a foamy quality when agitated in water. They are widely distributed ...

s. These adaptations and counter adaptations probably have led to extensive diversification in both the Brassicaceae and one of its major pests, the butterfly family

. A particular cocktail of volatile glucosinates triggers egg-laying in many species. Thus a particular crop can sometimes be protected by planting bittercress as a deadly bait, for the saponins kill the caterpillars, but the butterfly is still lured by the bittercress to lay its egg on the leaves. A moth that feeds on a range of Brassicaceae is the

diamondback moth The diamondback moth (''Plutella xylostella''), sometimes called the cabbage moth, is a moth species of the family Plutellidae and genus '' Plutella''. The small, grayish-brown moth sometimes has a cream-colored band that forms a diamond along ...

(''Plutella xylostella''). Like the Pieridae, it is capable of converting isothiocyanates into less problematic nitriles. Managing this pest in crops became more complicated after resistance developed against a toxin produced by '' Bacillus thuringiensis'', which is used as a wide spectrum biological

plant protection Crop protection is the science and practice of managing plant diseases, weeds, and other pests (both vertebrate and invertebrate) that damage crops and forestry. Crops include field crops (maize, wheat, rice, etc.), vegetable crops ( potatoes, ca ...

against caterpillars. Parasitoid wasps that feed on such insect herbivores are attracted to the chemical compounds released by the plants, and thus are able to locate their prey. The

cabbage aphid ''Brevicoryne brassicae'', commonly known as the cabbage aphid or cabbage aphis, is a destructive aphid (plant louse) native to Europe that is now found in many other areas of the world. The aphids feed on many varieties of produce, including cab ...

(''Brevicoryne brassicae'') stores glucosinolates and synthesises its own myrosinases, which may deter its potential predators. Since its introduction in the 19th century, ''Alliaria petiolata'' has been shown to be extremely successful as an invasive species in temperate North America due, in part, to its secretion of allelopathic chemicals. These inhibit the germination of most competing plants and kill beneficial soil

fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...

needed by many plants, such as many tree species, to successfully see their seedlings grow to maturity. The

monoculture In agriculture, monoculture is the practice of growing one crop species in a field at a time. Monoculture is widely used in intensive farming and in organic farming: both a 1,000-hectare/acre cornfield and a 10-ha/acre field of organic kale are ...

formation of an herb layer carpet by this plant has been shown to dramatically alter forests, making them wetter, having fewer and fewer trees, and having more vines such as poison ivy (''

Toxicodendron radicans ''Toxicodendron radicans'', commonly known as eastern poison ivy or poison ivy, is an allergenic Asian and Eastern North American flowering plant in the genus ''Toxicodendron''. The species is well known for causing urushiol-induced contact derm ...

''). The overall herb layer

biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...

is also drastically reduced, particularly in terms of

sedge The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus '' Carex'' ...

s and

forb A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...

s. Research has found that removing 80% of the garlic mustard

infestation Infestation is the state of being invaded or overrun by pests or parasites. It can also refer to the actual organisms living on or within a host. Terminology In general, the term "infestation" refers to parasitic diseases caused by animals s ...

plants did not lead to a particularly significant recovery of that

diversity Diversity, diversify, or diverse may refer to: Business *Diversity (business), the inclusion of people of different identities (ethnicity, gender, age) in the workforce *Diversity marketing, marketing communication targeting diverse customers * ...

. Instead, it required around 100% removal. Given that not one of an estimated 76

that

prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...

on the plant has been approved for

biological control Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also i ...

in North America and the variety of mechanisms the plant has to ensure its dominance without them (e.g. high seed production, self-fertility, allelopathy, spring growth that occurs before nearly all native plants, roots that break easily when pulling attempts are made, a complete lack of palatability for herbivores at all life stages, etc.) it is unlikely that such a high level of control can be established and maintained on the whole.Becker, R., Gerber E., Hinz H., Katovich E., Panke B., Reardon R., Renz R., Van Riper L., 2013. Biology and Biological Control of Garlic Mustard. The Forest Technology Enterprise Team. https://www.fs.fed.us/foresthealth/technology/pdfs/FS_garlicmustard.pdfDriesche, F.V.; Blossey, B.; Hoodle, M.; Lyon, S.; Reardon, R., 2010. Biological Control of Invasive Plants in the Eastern United States. USDA Forest Service. Forest Health Technology Enterprise Team. http://wiki.bugwood.org/Archive:BCIPEUS It is estimated that adequate control can be achieved with the introduction of two European

weevil Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...

s, including one that is

monophagous A generalist species is able to thrive in a wide variety of environmental conditions and can make use of a variety of different resources (for example, a heterotroph with a varied diet). A specialist species can thrive only in a narrow range of e ...

. The

USDA The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of com ...

's TAG group has blocked these introductions since 2004. In addition to being invasive, garlic mustard also is a threat to native North American '' Pieris'' butterflies such as '' P. oleracea'', as they preferentially

oviposit The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typical ...

on it, although it is toxic to their

e. Invasive aggressive mustard species are known for being self-fertile, seeding very heavily with small seeds that have a lengthy lifespan coupled with a very high rate of viability and germination, and for being completely unpalatable to both herbivores and insects in areas to which they are not native. Garlic mustard is toxic to several rarer North American ''Pieris'' species.

Uses

This family includes important agricultural crops, among which many vegetables such as cabbage,

, cauliflower, kale,

Brussels sprouts The Brussels sprout is a member of the Gemmifera cultivar group of cabbages (''Brassica oleracea''), grown for its edible buds. The leaf vegetables are typically 1.5–4.0 cm (0.6–1.6 in) in diameter and resemble miniature cabbag ...

collard greens Collard is a group of certain loose-leafed cultivars of ''Brassica oleracea'', the same species as many common vegetables including cabbage ( Capitata group) and broccoli ( Italica group). Collard is a member of the Viridis group of ''Brassica ...

, Savoy,

kohlrabi Kohlrabi (pronounced ; scientific name ''Brassica oleracea'' Gongylodes Group (horticulture), Group), also called German turnip or turnip cabbage, is a Biennial plant, biennial vegetable, a low, stout cultivar of wild cabbage. It is a cultivar o ...

, and

gai lan Gai lan, kai-lan, Chinese broccoli, Chinese kale, or jie lan (''Brassica oleracea'' var. ''alboglabra'') is a leaf vegetable with thick, flat, glossy blue-green leaves with thick stems, and florets similar to (but much smaller than) broccoli. ...

(''

''),

napa cabbage Napa cabbage (''Brassica rapa'' subsp. ''pekinensis'' or ''Brassica rapa'' Pekinensis Group) is a type of Chinese cabbage originating near the Beijing region of China that is widely used in East Asian cuisine. Since the 20th century, it has als ...

bomdong Bomdong () also known as spring cabbage is a hardy cabbage with tough, sweet leaves. The leaves of bomdong, unlike those of regular napa cabbages, fall to the sides, giving the plant a flat shape. This cabbage is primarily used in the making of ...

bok choy Bok choy (American English, Canadian English, and Australian English), pak choi (British English) or pok choi (''Brassica rapa'' subsp. ''chinensis'') is a type of Chinese cabbage, used as food. ''Chinensis'' varieties do not form heads and have ...

and

rapini Rapini or broccoli rabe () is a green cruciferous vegetable, with the leaves, buds, and stems all being edible; the buds somewhat resemble broccoli, but do not form a large head. Rapini is known for its bitter taste, and is particularly associ ...

(''

''), rocket salad/arugula (''

Eruca sativa Arugula (American English) or rocket (Commonwealth English) (''Eruca vesicaria''; syns. ''Eruca sativa'' Mill., ''E. vesicaria'' subsp. ''sativa'' (Miller) Thell., ''Brassica eruca'' L.) is an edible annual plant in the family Brassicaceae used ...

''),

garden cress Cress (''Lepidium sativum''), sometimes referred to as garden cress (or curly cress) to distinguish it from similar plants also referred to as cress (from old Germanic ''cresso'' which means sharp, spicy), is a rather fast-growing, edible herb. ...

(''

Lepidium sativum Cress (''Lepidium sativum''), sometimes referred to as garden cress (or curly cress) to distinguish it from similar plants also referred to as cress (from old Germanic ''cresso'' which means sharp, spicy), is a rather fast-growing, edible herb ...

''),

watercress Watercress or yellowcress (''Nasturtium officinale'') is a species of aquatic flowering plant in the cabbage family Brassicaceae. Watercress is a rapidly growing perennial plant native to Europe and Asia. It is one of the oldest known leaf v ...

(''

Nasturtium officinale Watercress or yellowcress (''Nasturtium officinale'') is a species of aquatic flowering plant in the cabbage family Brassicaceae. Watercress is a rapidly growing perennial plant native to Europe and Asia. It is one of the oldest known leaf ve ...

'') and

(''

Raphanus ''Raphanus'' (Latin for "radish") is a genus within the flowering plant family Brassicaceae. Carl Linnaeus described three species within the genus: the cultivated radish (''Raphanus sativus''), the wild radish or jointed charlock (''Raphanus ra ...

'') and a few spices like

(''

''), ''Brassica'',

wasabi Wasabi ( Japanese: , , or , ; ''Eutrema japonicum'' or ''Wasabia japonica'') or Japanese horseradish is a plant of the family Brassicaceae, which also includes horseradish and mustard in other genera. The plant is native to Japan and the Russi ...

('' Eutrema japonicum''), white, Indian and black mustard (''

Sinapis alba White mustard (''Sinapis alba'') is an annual plant of the family Brassicaceae. It is sometimes also referred to as ''Brassica alba'' or ''B. hirta''. Grown for its seeds, used to make the condiment mustard, as fodder crop, or as a green manu ...

'', ''

Brassica juncea ''Brassica juncea'', commonly brown mustard, Chinese mustard, Indian mustard, leaf mustard, Oriental mustard and vegetable mustard, is a species of mustard plant. Cultivar ''Brassica juncea'' cultivars can be divided into four major subgroup ...

'' and '' B. nigra'' respectively). Vegetable oil is produced from the seeds of several species such as '' Brassica napus'' (rapeseed oil), perhaps providing the largest volume of vegetable oils of any species.

Woad ''Isatis tinctoria'', also called woad (), dyer's woad, or glastum, is a flowering plant in the family Brassicaceae (the mustard family) with a documented history of use as a blue dye and medicinal plant. Its genus name, Isatis, derives from ...

(''

Isatis tinctoria ''Isatis tinctoria'', also called woad (), dyer's woad, or glastum, is a flowering plant in the family Brassicaceae (the mustard family) with a documented history of use as a blue dye and medicinal plant. Its genus name, Isatis, derives from ...

'') was used in the past to produce a blue textile dye (

indigo Indigo is a deep color close to the color wheel blue (a primary color in the RGB color space), as well as to some variants of ultramarine, based on the ancient dye of the same name. The word "indigo" comes from the Latin word ''indicum'', m ...

), but has largely been replaced by the same substance from unrelated tropical species like ''

Indigofera tinctoria ''Indigofera tinctoria'', also called true indigo, is a species of plant from the bean family that was one of the original sources of indigo dye. Description True indigo is a shrub one to two meters high. It may be an annual plant, annual, bi ...

''.

Brassinosteroids Brassinosteroids (BRs or less commonly BS) are a class of polyhydroxysteroids that have been recognized as a sixth class of plant hormones and may have utility as an anticancer drug for endocrine-responsive cancers to induce apoptosis and inhibi ...

are growing in agricultural and gardening importance. ''Pringlea antiscorbutica'', commonly known as Kerguelen cabbage, is edible, containing high levels of

potassium Potassium is the chemical element with the symbol K (from Neo-Latin ''kalium'') and atomic number19. Potassium is a silvery-white metal that is soft enough to be cut with a knife with little force. Potassium metal reacts rapidly with atmosph ...

. Its leaves contain a vitamin C-rich oil, a fact which, in the days of sailing ships, made it very attractive to sailors suffering from

scurvy Scurvy is a disease resulting from a lack of vitamin C (ascorbic acid). Early symptoms of deficiency include weakness, feeling tired and sore arms and legs. Without treatment, decreased red blood cells, gum disease, changes to hair, and bleeding ...

, hence the species name's epithet ''antiscorbutica'', which means "against scurvy" in

Low Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in ...

. It was essential to the diets of the whalers on

Kerguelen The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the sub-Antarctic constituting one of the two exposed parts of the Kerguelen Plateau, a large ...

when pork, beef, or

seal meat Seal meat is the flesh, including the blubber and organs, of seals used as food for humans or other animals. It is prepared in numerous ways, often being hung and dried before consumption. Historically, it has been eaten in many parts of the world, ...

was used up. The Brassicaceae also includes ornamentals, such as species of '' Aethionema'', ''

'', ''

'', '' Aubrieta'', '' Aurinia'', '' Cheiranthus'', ''

'', ''

Hesperis ''Hesperis'' is a genus of flowering plants in the family Brassicaceae. Most are native to Eurasia, with several endemic to Greece & Turkey. Many plants of this genus bear showy, fragrant flowers in shades of purple and white. One of the more wi ...

'', ''

'', '' Lobularia'', ''

Lunaria ''Lunaria'', common names honesty, dollar plant, money-in-both-pockets, money plant, moneywort, moonwort, and silver dollar; is a genus of flowering plants in the family Brassicaceae. It is native to central and southern Europe and North Americ ...

'', '' Malcolmia'', and ''

''.

Honesty Honesty or truthfulness is a facet of moral character that connotes positive and virtuous attributes such as integrity, truthfulness, straightforwardness, including straightforwardness of conduct, along with the absence of lying, cheating, theft ...

(''

Lunaria annua ''Lunaria annua'', called honesty or annual honesty in English, is a species of flowering plant in the cabbage and mustard family Brassicaceae. It is native to the Balkans and south west Asia, and cultivated throughout the temperate world. Desc ...

'') is cultivated for the decorative value of the translucent remains of the fruits after drying. It can be a pest species in areas where it is not native. The small Eurasian weed '' Arabidopsis thaliana'' is widely used as model organism in the study of the

molecular biology Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including biomolecular synthesis, modification, mechanisms, and interactions. The study of chemical and physi ...

of flowering plants ( Angiospermae). Some species are useful as food plants for Lepidoptera, such as certain wild mustard and cress species, such as '' Turritis glabra'' and ''

Boechera laevigata ''Boechera laevigata'' is a species of flowering plant in the mustard family known by the common name smooth rockcress. It is native to many areas of the eastern United States and Canada, where it grows in calcareous rocky woods and bluffs. It is ...

'' that are utilized by several North American butterflies.

Gallery

File:Brassicaceae_Coast_Sand_Loving_wallflower_erysimum_ammophilum.jpg, Coast/sand-loving wallflower '' Erysimum ammophilum'' File:Brassicaceae_Money_Plant_Honesty_Lunaria_annua.jpg, Honesty ''

'' File:Brassicaceae_Western_wallflower_erysimum_capitatum_var_capitatum.jpg, Western wallflower ''

Erysimum capitatum ''Erysimum capitatum'' is a species of wallflower known commonly as the sanddune wallflower, western wallflower, or prairie rocket. This species can be found in regions across North America, from the Great Lakes to the West Coast of the United S ...

var. capitatum''

References

External links

BrassiBase
a collection of resources on Brassicaceae biology

Description

Ovary, fruit and seed

Differences with similar families

Phytochemistry

Taxonomy

Relationships within the family

Genera

Etymology

Distribution

Ecology

Uses

Gallery

References

External links

Further reading