Chialingosaurus
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''Chialingosaurus'' (meaning "Chialing Lizard") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of herbivorous
stegosauria Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, predominantly in what is now North America, Europe, ...
n
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
similar to ''
Kentrosaurus ''Kentrosaurus'' ( ; ) is a genus of stegosaurid dinosaur from the Late Jurassic in Lindi Region of Tanzania. The type species is ''K. aethiopicus'', named and described by German palaeontologist Edwin Hennig in 1915. Often thought to be a " pri ...
'' from the Upper
Shaximiao Formation The Shaximiao Formation () is a Middle Jurassic, Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in ...
,
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
beds in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
Province in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. Its age makes it one of the oldest species of stegosaurs, living about 160 million years ago. Since it was an
herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, scientists think that ''Chialingosaurus'' probably ate ferns and
cycads Cycads are seed plants that typically have a stout and woody (ligneous) trunk with a crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants of a species are either male or ...
, which were plentiful during the period when ''Chialingosaurus'' was alive.


Discovery and species

The fossils of ''Chialingosaurus'' were collected by the geologist Kuan Yaowu or Guan Yao-Wu in 1957, at Taipingstai in Quxian County, while surveying the Chialing River in southern China. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Chialingosaurus kuani'' was named and described by paleontologist
Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. H ...
, ("C. C. Young") two years later in 1959. The generic name refers to the Chialing. The specific name honours Kuan. ''Chialingosaurus'' was the first stegosaurian described from China.C.-C. Young, 1959, "On a new Stegosauria from Szechuan, China", ''Vertebrata PalAsiatica'' 3(1): 1-8 The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, IVPP 2300, was found in a layer of the Upper Shaximiao Formation, dating from the Oxfordian-
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
. It consists of a partial skeleton lacking the skull. It contains six vertebrae, the coracoids, the humeri, a right radius and three spines. The original material has been supplemented after November 1978 by
Zhou Shiwu Zhou may refer to: Chinese history * King Zhou of Shang () (1105 BC–1046 BC), the last king of the Shang dynasty * Predynastic Zhou (), 11th-century BC precursor to the Zhou dynasty * Zhou dynasty () (1046 BC–256 BC), a dynasty of China ** West ...
of the Municipal Museum of Chongqing. A second specimen, CV 202, was referred. It represents a skeleton with a partial skull and lower jaws, some vertebrae and limb elements, and four plates. Zhou considered it likely that the holotype and CV 202 represented a single individual, as the material was not overlapping. A third specimen, CV 203, a partial skeleton lacking the skull, was made the
paratype In zoology and botany, a paratype is a specimen of an organism that helps define what the scientific name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Of ...
. All specimens are juvenile or subadult. In 1999, Li Kui,
Zhang Yuguang Zhang may refer to: Chinese culture, etc. * Zhang (surname) (張/张), common Chinese surname ** Zhang (surname 章), a rarer Chinese surname * Zhang County (漳县), of Dingxi, Gansu * Zhang River (漳河), a river flowing mainly in Henan * ''Z ...
and Cai Kaiji mentioned a second species: ''Chialingosaurus guangyuanensis''. A description was not provided, however, and the name is seen as a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
''.


Description

Yang in 1959 provided a diagnosis, emphasising the slender build of the animal. It might have been only four metres (thirteen feet) long. However, the gracile proportions may have been caused by the subadult age and the remainder of the diagnostic traits were in fact shared by other stegosaurs. In 1990,
Peter Malcolm Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosa ...
identified just a single
autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
: the lesser trochanter of the thighbone is triangular with a broad base.
Susannah Maidment Susannah Catherine Rose Maidment is a British palaeontologist at the Natural History Museum, London. She is internationally recognised for her research on ornithischian dinosaur evolution, and was awarded the 2016 Hodson Award of the Palaeontologic ...
in 2008 even listed ''Chialingosaurus kuani'' among the invalid stegosaur taxa in her phylogeny of Stegosauria. This was based upon previous work by Maidment & Wei in 2006 in which the authors were unable to identify any autapomorphies or unique character combinations for ''Chialingosaurus'', therefore making it a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
''. Yang modelled ''Chialingosaurus'' after ''Kentrosaurus''. Some plates and spine bases having been found, he suggested that the plates were placed on the front upper parts of the animal, the spines at the hip and tail.


Phylogeny

Yang in 1959 placed ''Chialingosaurus'' in the
Stegosaurinae Stegosauridae is a family of thyreophoran dinosaurs (armoured dinosaurs) within the suborder Stegosauria. The clade is defined as all species of dinosaurs more closely related to ''Stegosaurus'' than ''Huayangosaurus''.David B. Weishampel, Peter ...
within the
Stegosauridae Stegosauridae is a family of thyreophoran dinosaurs (armoured dinosaurs) within the suborder Stegosauria. The clade is defined as all species of dinosaurs more closely related to ''Stegosaurus'' than ''Huayangosaurus''.David B. Weishampel, Peter ...
. In 1969, Rodney Steel suggested that ''Chialingosaurus'' might have actually been an early ancestor of other stegosaurs. However, Galton in 2004 recovered a rather derived position, albeit indeed close to ''Kentrosaurus''.Galton, P.M., and P. Upchurch, 2004, "Stegosauria", pp. 343-362 in: D.B. Weishampel, P. Dodson, and H. Osmolska (eds.), ''The Dinosauria'', 2nd Edition. University of California Press, Berkeley, CA Dong confirmed the placement in the
Stegosaurinae Stegosauridae is a family of thyreophoran dinosaurs (armoured dinosaurs) within the suborder Stegosauria. The clade is defined as all species of dinosaurs more closely related to ''Stegosaurus'' than ''Huayangosaurus''.David B. Weishampel, Peter ...
.


See also

*
Timeline of stegosaur research This timeline of stegosaur research is a chronological listing of events in the history of paleontology focused on the stegosaurs, the iconic plate-backed, spike-tailed herbivorous eurypod dinosaurs that predominated during the Jurassic period ...


Sources

*


Notes

{{Taxonbar, from=Q132494 Late Jurassic dinosaurs of Asia Stegosaurs Fossil taxa described in 1959 Taxa named by Yang Zhongjian Paleontology in Sichuan Nomina dubia Ornithischian genera