''Caypullisaurus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

platypterygiine Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currentl ...

ophthalmosaurid Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the ex ...

ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, altho ...

from the

Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...

to the

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

(

Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...

and

Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...

stages) of

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

. Its

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

was collected from the

Vaca Muerta Formation The Vaca Muerta Formation, commonly known as Vaca Muerta (Spanish for ''Dead Cow''), is a geologic formation of Late Jurassic to Early Cretaceous age, located in the Neuquén Basin in northern Patagonia, Argentina. It is well known as the host roc ...

of Cerro Lotena, Neuquen, dating to the early

stage of the

, about 150

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. ''Caypullisaurus'' was first named by Marta Fernández in

1997 File:1997 Events Collage.png, From left, clockwise: The movie set of ''Titanic'', the highest-grossing movie in history at the time; ''Harry Potter and the Philosopher's Stone'', is published; Comet Hale-Bopp passes by Earth and becomes one of t ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Caypullisaurus bonapartei''. It was a large ichthyosaur, measuring about long.

Classification

It is a member of the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Ophthalmosauridae Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) worldwide. Almost all ichthyosaurs from the Middle Jurassic onwards belong to the family, until the ex ...

, and closely related to ''

Platypterygius ''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus or ...

'' and ''

Brachypterygius ''Brachypterygius'' (meaning ″short wing/paddle″ in Greek) is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England.McGowan, C. & Motani, R. ''Ichthyopterygia''. In Sues, H.-D. (ed.) Handb ...

''.Fernández M. 2007. Redescription and phylogenetic position of ''Caypullisaurus'' (Ichthyosauria: Ophthalmosauridae). ''Journal of Paleontology'' 81 (2): 368-375. In 2012, ''Caypullisaurus'' was found to be most closely related to ''

Athabascasaurus ''Athabascasaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from Alberta, Canada. Discovery and etymology ''Athabascasaurus'' is known from the holotype TMP 2000.29.01, articulated nearly complete postcr ...

'' and "''Platypterygius''" ''australis'', and to nest within the subfamily

Platypterygiinae Platypterygiinae is an extinct subfamily of ophthalmosaurid thunnosaur ichthyosaurs from the early Late Jurassic to the early Late Cretaceous (Kimmeridgian - Cenomanian) of Asia, Australia, Europe, North America and South America. Currently, the ...

Phylogeny

The following cladogram shows a possible phylogenetic position of ''Caypullisaurus'' in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).

References

Early Cretaceous ichthyosaurs Late Jurassic ichthyosaurs Berriasian life Tithonian life Late Jurassic reptiles of South America Early Cretaceous reptiles of South America Cretaceous Argentina Jurassic Argentina Fossils of Argentina Neuquén Basin Fossil taxa described in 1997 Ophthalmosauridae Ichthyosauromorph genera {{cretaceous-reptile-stub

Classification

Phylogeny

See also

References