Catfish (or catfishes;
order Siluriformes or Nematognathi) are a diverse group of
ray-finned
Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species.
The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
. Named for their prominent
barbel Barbel may refer to:
*Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles
*Barbel (fish), a common name for certain species of fish
**''Barbus barbus'', a species of cyprinid ...
s, which resemble a
cat
The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
's
whisker
Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser ...
s, catfish range in size and behavior from the three largest species alive, the
Mekong giant catfish
The Mekong giant catfish (''Pangasianodon gigas''; th, ปลาบึก, , ; km, ត្រីរាជ /''trəy riec''/; vi, cá tra dầu), is a large, threatened species of catfish (order Siluriformes) in the shark catfish family (Pang ...
from
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, the
wels catfish
The wels catfish ( or ; ''Silurus glanis''), also called sheatfish or just wels, is a large species of catfish native to wide areas of central, southern, and eastern Europe, in the basins of the Baltic, Black and Caspian Seas. It has been introd ...
of
Eurasia
Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
, and the
piraíba
''Brachyplatystoma filamentosum'', the piraíba or kumakuma, is a species of large catfish of the family Pimelodidae and genus Brachyplatystoma that is native to Amazon and Orinoco River basins and fluvials in Guianas and northeastern Brazil.
D ...
of
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, to
detritivore
Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...
s (species that eat dead material on the bottom), and even to a tiny
parasitic
Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has c ...
species commonly called the
candiru, ''Vandellia cirrhosa''. Neither the armour-plated types nor the naked types have scales. Despite their name, not all catfish have prominent barbels or "whiskers". Members of the Siluriformes order are defined by features of the skull and
swimbladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...
. Catfish are of considerable commercial importance; many of the larger species are
farmed
Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people t ...
or fished for food. Many of the smaller species, particularly the genus ''
Corydoras
''Corydoras'' is a genus of freshwater catfish in the family Callichthyidae and subfamily Corydoradinae. The species usually have more restricted areas of endemism than other callichthyids, but the area of distribution of the entire genus almost ...
'', are important in the
aquarium hobby. Many catfish are
nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.
Nocturnal creatures generally have highly developed sens ...
,
[Catfish Varieties](_blank)
animal-world.com[Wong, Kate (6 June 2001]
"How Nocturnal Catfish Stalk Their Prey"
''Scientific American''. but others (many
Auchenipteridae
The driftwood catfishes are catfishes of the family (biology), family Auchenipteridae. The two genera of the former family Ageneiosidae have been placed here, resulting in a grouping of about 125 species in about 22 genera.
These fish are found ...
) are
crepuscular
In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
or
diurnal (most
Loricariidae
The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are not ...
or
Callichthyidae
Callichthyidae is a family of catfishes ( order Siluriformes), called armored catfishes due to the two rows of bony plates (or scutes) along the lengths of their bodies. It contains some of the most popular freshwater aquarium fish, such as many ...
, for example).
Ecology
Distribution and habitat
Extant catfish species live inland or in coastal waters of every continent except
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
. Catfish have inhabited all continents at one time or another.
They are most diverse in
tropical
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
South America, Asia, and Africa, with one family native to North America and one family in Europe.
[ More than half of all catfish species live in the Americas. They are the only ]ostariophysan
Ostariophysi is the second-largest superorder of fish. Members of this superorder are called ostariophysians. This diverse group contains 10,758 species, about 28% of known fish species in the world and 68% of freshwater species, and are present ...
s that have entered freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
habitats in Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
, Australia, and New Guinea
New Guinea (; Hiri Motu
Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea).
It is a simplified version of ...
.
They are found in fresh water environments, though most inhabit shallow, running water. Representatives of at least eight families are hypogean
Hypogeal, hypogean, hypogeic and hypogeous (; ) are biological terms describing an organism's activity below the soil surface.
In botany, a seed is described as showing hypogeal germination when the cotyledons of the germinating seed remain n ...
(live underground) with three families that are also troglobitic
A troglobite (or, formally, troglobiont) is an animal species, or population of a species, strictly bound to underground habitats, such as caves. These are separate from species that mainly live in above-ground habitats but are also able to live u ...
(inhabiting caves). One such species is ''Phreatobius cisternarum
''Phreatobius cisternarum'' is a species of catfish in the genus ''Phreatobius''. This Brazilian fish is one of the few fish species that Cavefish, live underground in phreatic habitats. It has proved problematic in its classification.Muriel-Cunh ...
'', known to live underground in phreatic
''Phreatic'' is a term used in hydrology to refer to aquifers, in speleology to refer to cave passages, and in volcanology to refer to a type of volcanic eruption.
Hydrology
The term phreatic (the word originates from the Greek , meaning "well" ...
habitats. Numerous species from the families Ariidae
The Ariidae or ariid catfish are a family of catfish that mainly live in marine waters with many freshwater and brackish water species. They are found worldwide in tropical to warm temperate zones. The family includes about 143 species.
Taxonomy ...
and Plotosidae, and a few species from among the Aspredinidae
The Aspredinidae are a small South American family of catfishes (order Siluriformes) also known as the banjo catfishes, with about 43 species.
Distribution
Aspredinids are found throughout the major tropical rivers of South America (e.g., Magda ...
and Bagridae
The Bagridae are a family of catfish that are native to Africa ('' Bagrus'') and Asia (all other genera) from Japan to Borneo. It includes about 245 species. These fish are commonly known as naked catfishes or bagrid catfishes.
Many large bagrid ...
, are found in salt water.
In the Southern United States, catfish species may be known by a variety of slang names, such as "mud cat", "polliwogs", or "chuckleheads". These nicknames are not standardized, so one area may call a bullhead catfish by the nickname "chucklehead", while in another state or region, that nickname refers to the blue catfish.
As invasive species
Representatives of the genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Ictalurus
''Ictalurus'' is a genus of North American freshwater catfishes. It includes the well-known channel catfish (''Ictalurus punctatus'') and blue catfish (''Ictalurus furcatus'').
The catfish genome database (cBARBEL) is a database for the genetic ...
'' have been introduced into European waters in the hope of obtaining a sporting and food resource, but the European stock of American catfishes has not achieved the dimensions of these fish in their native waters, and have only increased the ecological pressure on native European fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is '' funga''. Flora, fauna, funga and other forms of life are collectively referred to as '' biota''. Zoo ...
. Walking catfish
The walking catfish (''Clarias batrachus'') is a species of freshwater airbreathing catfish native to Southeast Asia. It is named for its ability to "walk" and wiggle across dry land, to find food or suitable environments. While it does not trul ...
have also been introduced in the freshwater areas of Florida, with the voracious catfish becoming a major alien pest there. Flathead catfish
The flathead catfish (''Pylodictis olivaris''), also called by several common names including mudcat or shovelhead cat, is a large species of North American freshwater catfish in the family Ictaluridae. It is the only species of the genus ''P ...
, ''Pylodictis olivaris'', is also a North American pest on Atlantic slope drainages.[ '']Pterygoplichthys
''Pterygoplichthys'', sometimes collectively known as janitor fish, is a genus of South American armored catfishes. These fish are commonly known as sailfin armoured catfish or sailfin plecs, especially in the aquarium trade.
Etymology
''Pte ...
'' species, released by aquarium fishkeepers, have also established feral
A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
populations in many warm waters around the world.
Physical characteristics
External anatomy of catfish
Most catfish are bottom feeder
A bottom feeder is an aquatic animal that feeds on or near the bottom of a body of water. Biologists often use the terms ''benthos''—particularly for invertebrates such as shellfish, crabs, crayfish, sea anemones, starfish, snails, bri ...
s. In general, they are negatively buoyant, which means that they usually sink rather than float due to a reduced gas bladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...
and a heavy, bony head. Catfish have a variety of body shapes, though most have a cylindrical body with a flattened ventrum
The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso ...
to allow for benthic feeding.
A flattened head allows for digging through the substrate, as well as perhaps serving as a hydrofoil
A hydrofoil is a lifting surface, or foil, that operates in water. They are similar in appearance and purpose to aerofoils used by aeroplanes. Boats that use hydrofoil technology are also simply termed hydrofoils. As a hydrofoil craft gains sp ...
. Some have a mouth that can expand to a large size and contains no incisiform teeth; catfish generally feed through suction
Suction is the colloquial term to describe the air pressure differential between areas.
Removing air from a space results in a pressure differential. Suction pressure is therefore limited by external air pressure. Even a perfect vacuum cannot ...
or gulping rather than biting and cutting prey. Some families, though, notably the Loricariidae
The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are not ...
and Astroblepidae
''Astroblepus'' is a genus of fish in the family Astroblepidae found in South America and Panama. This genus is the only member of its family. These catfishes are primarily found in torrential streams in the Andean area. ''Astroblepus pholeter'' ...
, have a suckermouth In fish, a suckermouth is a ventrally-oriented (inferior) mouth adapted for grazing on algae and small organisms that grow on submerged objects.
All Loricariidae possess a suckermouth as do the cypriniform algae eaters of the genus ''Gyrinocheilus' ...
that allows them to fasten themselves to objects in fast-moving water. Catfish also have a maxilla
The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
reduced to a support for barbels
In fish anatomy and turtle anatomy, a barbel is a slender, whiskerlike sensory organ near the mouth. Fish that have barbels include the catfish, the carp, the goatfish, the hagfish, the sturgeon, the zebrafish, the black dragonfish and some ...
; this means that they are unable to protrude their mouths as other fish such as carp
Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
.
Catfish may have up to four pairs of barbels - nasal, maxillary (on each side of mouth), and two pairs of chin barbels, though pairs of barbels may be absent depending on the species. Catfish barbels always occur in pairs. Many larger catfish also have chemoreceptor
A chemoreceptor, also known as chemosensor, is a specialized sensory receptor which transduces a chemical substance (endogenous or induced) to generate a biological signal. This signal may be in the form of an action potential, if the chemorecept ...
s across their entire bodies, which means they "taste" anything they touch and "smell" any chemicals in the water. "In catfish, gustation
The gustatory system or sense of taste is the sensory system that is partially responsible for the perception of taste (flavor). Taste is the perception produced or stimulated when a substance in the mouth reacts chemically with taste receptor ...
plays a primary role in the orientation and location of food". Because their barbels and chemoreception are more important in detecting food, the eyes on catfish are generally small. Like other ostariophysan
Ostariophysi is the second-largest superorder of fish. Members of this superorder are called ostariophysians. This diverse group contains 10,758 species, about 28% of known fish species in the world and 68% of freshwater species, and are present ...
s, they are characterized by the presence of a Weberian apparatus
The Weberian apparatus is an anatomical structure that connects the swim bladder to the auditory system in fishes belonging to the superorder Ostariophysi. When it is fully developed in adult fish, the elements of the apparatus are sometimes col ...
. Their well-developed Weberian apparatus and reduced gas bladder allow for improved hearing
Hearing, or auditory perception, is the ability to perceive sounds
In physics, sound is a vibration that propagates as an acoustic wave, through a transmission medium such as a gas, liquid or solid.
In human physiology and psycholog ...
and sound production.
Catfish do not have scales; their bodies are often naked. In some species, their mucus
Mucus ( ) is a slippery aqueous secretion produced by, and covering, mucous membranes. It is typically produced from cells found in mucous glands, although it may also originate from mixed glands, which contain both serous and mucous cells. It is ...
-covered skin
Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation.
Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...
is used in cutaneous respiration Cutaneous respiration, or cutaneous gas exchange (sometimes called, skin breathing), is a form of respiration in which gas exchange occurs across the skin or outer integument of an organism rather than gills or lungs. Cutaneous respiration may be th ...
, where the fish breathes through its skin. In some catfish, the skin is covered in bony plates called scute
A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...
s; some form of body armor appears in various ways within the order. In loricarioids and in the Asian genus ''Sisor
''Sisor'' is a genus of catfishes native to Asia.
Taxonomy
''Sisor'' was previously monotypic, containing only ''S. rabdophorus'', prior to a review of the genus in 2003 in which the three species ''S. chennuah'', ''S. rheophilus'', and ''S. tor ...
'', the armor is primarily made up of one or more rows of free dermal
The dermis or corium is a layer of skin between the epidermis (with which it makes up the cutis) and subcutaneous tissues, that primarily consists of dense irregular connective tissue and cushions the body from stress and strain. It is divided in ...
plates. Similar plates are found in large specimens of '' Lithodoras''. These plates may be supported by vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
l process
A process is a series or set of activities that interact to produce a result; it may occur once-only or be recurrent or periodic.
Things called a process include:
Business and management
*Business process, activities that produce a specific se ...
es, as in scoloplacids and in ''Sisor'', but the processes never fuse to the plates or form any external armor. By contrast, in the subfamily Doumeinae (family Amphiliidae
The loach catfishes are a family, Amphiliidae, of catfishes ( order Siluriformes). They are widespread in tropical Africa, but are most common in streams at high elevations; most species are able to cling to rocks in fast-flowing streams. The 13 ...
) and in hoplomyzontines (Aspredinidae
The Aspredinidae are a small South American family of catfishes (order Siluriformes) also known as the banjo catfishes, with about 43 species.
Distribution
Aspredinids are found throughout the major tropical rivers of South America (e.g., Magda ...
), the armor is formed solely by expanded vertebral processes that form plates. Finally, the lateral armor of doradids, ''Sisor'', and hoplomyzontines consists of hypertrophied lateral line
The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
ossicle
The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ...
s with dorsal and ventral lamina
Lamina may refer to:
Science and technology
* Planar lamina, a two-dimensional planar closed surface with mass and density, in mathematics
* Laminar flow, (or streamline flow) occurs when a fluid flows in parallel layers, with no disruption betwee ...
.
All catfish other than members of the Malapteruridae
Electric catfish or Malapteruridae is a Family (biology), family of catfishes (order Siluriformes). This family includes two genus, genera, ''Malapterurus'' and ''Paradoxoglanis'', with 21 species. Several species of this family have the ability ...
(electric
Electricity is the set of physical phenomena associated with the presence and motion of matter that has a property of electric charge. Electricity is related to magnetism, both being part of the phenomenon of electromagnetism, as described by ...
catfish), possess a strong, hollow, bony, leading spine-like ray on their dorsal
Dorsal (from Latin ''dorsum'' ‘back’) may refer to:
* Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism
* Dorsal, positioned on top of an aircraft's fuselage
* Dorsal co ...
and pectoral fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s. As a defense, these spines may be locked into place so that they stick outwards, enabling them to inflict severe wounds.[ In numerous catfish species, these fin rays can be used to deliver a stinging ]protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
if the fish is irritated; as many as half of all catfish species may be venomous in this fashion, making the Siluriformes overwhelmingly the vertebrate order with the largest number of venomous species. This venom
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
is produced by gland
In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland).
Structure
De ...
ular cells in the epidermal
The epidermis is the outermost of the three layers that comprise the skin, the inner layers being the dermis and hypodermis. The epidermis layer provides a barrier to infection from environmental pathogens and regulates the amount of water relea ...
tissue covering the spines. In members of the family Plotosidae and of the genus '' Heteropneustes'', this protein is so strong it may hospitalize humans who receive a sting; in ''Plotosus lineatus
''Plotosus lineatus'', common name striped eel catfish, is a species of eeltail catfishes belonging to the family Plotosidae.
Description
''Plotosus lineatus'' can reach a maximum length of 32 cm (13 in). The body is brown with cream- ...
'', the stings can be lethal. The dorsal- and pectoral-fin spines are two of the most conspicuous features of siluriforms, and differ from those in other fish groups. Despite the widespread use of the spines for taxonomic and phylogenetic studies the fields have struggled to effectively use the information due to a lack of consistency in the nomenclature, with a general standard for the descriptive anatomy of catfish spines proposed in 2022 to try and resolve this problem.
Juvenile catfish, like most fish, have relatively large heads, eyes, and posterior median fins in comparison to larger, more mature individuals. These juveniles can be readily placed in their families, particularly those with highly derived fin or body shapes; in some cases, identification of the genus is possible. As far as known for most catfish, features that are often characteristic of species, such as mouth and fin positions, fin shapes, and barbel lengths, show little difference between juveniles and adults. For many species, pigmentation pattern is also similar in juveniles and adults. Thus, juvenile catfish generally resemble and develop smoothly into their adult form without distinct juvenile specializations. Exceptions to this are the ariid catfish, where the young retain yolk sacs late into juvenile stages, and many pimelodids, which may have elongated barbels and fin filaments or coloration patterns.
Sexual dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is reported in about half of all families of catfish. The modification of the anal fin
Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
into an intromittent organ
An intromittent organ is any external organ of a male organism that is specialized to deliver sperm during copulation. Intromittent organs are found most often in terrestrial species, as most non-mammalian aquatic species fertilize their eggs e ...
(in internal fertilizers) as well as accessory structures of the reproductive apparatus (in both internal and external fertilizers) have been described in species belonging to 11 different families.
Size
Catfish have one of the greatest ranges in size within a single order of bony fish
Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...
. Many catfish have a maximum length of under . Some of the smallest species of the Aspredinidae
The Aspredinidae are a small South American family of catfishes (order Siluriformes) also known as the banjo catfishes, with about 43 species.
Distribution
Aspredinids are found throughout the major tropical rivers of South America (e.g., Magda ...
and Trichomycteridae
Trichomycteridae is a family of catfishes commonly known as pencil catfishes or parasitic catfishes. This family includes the candiru fish (''Vandellia cirrhosa''), feared by some people for its alleged habit of entering into the urethra of hum ...
reach sexual maturity at only .[
The ]wels catfish
The wels catfish ( or ; ''Silurus glanis''), also called sheatfish or just wels, is a large species of catfish native to wide areas of central, southern, and eastern Europe, in the basins of the Baltic, Black and Caspian Seas. It has been introd ...
, ''Silurus glanis'', and the much smaller related Aristotle's catfish
Aristotle's catfish (''Silurus aristotelis'') is a species of fish in the family Siluridae. It is endemic to Greece, where it occurs in the Acheloos River drainage. Its natural habitat is freshwater lakes. It is threatened by habitat loss. This ...
, are the only catfish indigenous to Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
; the former ranges throughout Europe, and the latter is restricted to Greece
Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
. Mythology
Myth is a folklore genre consisting of narratives that play a fundamental role in a society, such as foundational tales or origin myths. Since "myth" is widely used to imply that a story is not objectively true, the identification of a narrat ...
and literature record wels catfish of astounding proportions, yet are to be proven scientifically. The typical size of the species is about , and fish more than are rare. However, they are known to exceed in length and in weight. In July 2009, a catfish weighing was caught in the River Ebro
, name_etymology =
, image = Zaragoza shel.JPG
, image_size =
, image_caption = The Ebro River in Zaragoza
, map = SpainEbroBasin.png
, map_size =
, map_caption = The Ebro ...
, Spain, by an 11-year-old British schoolgirl.
In North America, the largest ''Ictalurus furcatus
The blue catfish (''Ictalurus furcatus'') is the largest species of North American catfish, reaching a length of and a weight of . The typical length is about 25–46 in (64–117 cm). The fish can live to 20 years. The native distribution o ...
'' (blue catfish) caught in the Missouri River on 20 July 2010, weighed . The largest flathead catfish
The flathead catfish (''Pylodictis olivaris''), also called by several common names including mudcat or shovelhead cat, is a large species of North American freshwater catfish in the family Ictaluridae. It is the only species of the genus ''P ...
, ''Pylodictis olivaris'', ever caught was in Independence, Kansas
Independence is a city in and the county seat of Montgomery County, Kansas, United States. As of the 2020 census, the population of the city was 8,548. It was named in commemoration of the Declaration of Independence.
History
Independence w ...
, weighing .
These records pale in comparison to a Mekong giant catfish
The Mekong giant catfish (''Pangasianodon gigas''; th, ปลาบึก, , ; km, ត្រីរាជ /''trəy riec''/; vi, cá tra dầu), is a large, threatened species of catfish (order Siluriformes) in the shark catfish family (Pang ...
caught in northern Thailand
Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
on 1 May 2005, and reported to the press almost 2 months later, that weighed . This is the largest giant Mekong catfish caught since Thai officials started keeping records in 1981. Also in Asia, Jeremy Wade
Jeremy John Wade (born 23 March 1956) is an English television presenter, an author of books on angling, a freshwater detective, and a biologist. He is known for his television series ''River Monsters'', ''Mighty Rivers'', and ''Dark Waters''. He ...
caught a goonch following three fatal attacks on humans in the Kali River on the India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
-Nepal
Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne,
सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
border. Wade was of the opinion that the offending fish must have been significantly larger than this to have taken an 18-year-old boy, as well as a water buffalo.
Piraíba ''(Brachyplatystoma, Brachyplatystoma filamentosum)'' can grow exceptionally large and are native to the Amazon Basin. They can occasionally grow to 200 kg (440 lb), as evidenced by numerous catches. Deaths from being swallowed by these fish have been reported in the region.
Internal anatomy
In many catfish, the "humeral process" is a bony process extending backward from the pectoral girdle immediately above the base of the pectoral fin. It lies beneath the skin, where its outline may be determined by dissecting the skin or probing with a needle.
The retinae of catfish are composed of single cone cell, cones and large rod cell, rods. Many catfish have a tapetum lucidum, which may help enhance photon capture and increase low-light sensitivity. Double cone (biology), Double cones, though present in most teleosts, are absent from catfish.
The anatomical organization of the testicle, testis in catfish is variable among the families of catfish, but the majority of them present fringed testis: Ictaluridae, Claridae, Auchenipteridae, Doradidae, Pimelodidae, and Pseudopimelodidae. In the testes of some species of Siluriformes, organs and structures such as a spermatogenic cranial region and a secretory caudal region are observed, in addition to the presence of seminal vesicles in the caudal region. The total number of fringes and their length are different in the Caudal (anatomical term), caudal and Skull, cranial portions between species. Fringes of the caudal region may present tubules, in which the lumen is filled by secretion and Spermatozoon, spermatozoa. Spermatocysts are formed from cytoplasmic extensions of Sertoli cells; the release of spermatozoa is allowed by breaking of the cyst walls.
The occurrence of seminal vesicles, in spite of their interspecific variability in size, gross morphology, and function, has not been related to the mode of fertilization. They are typically paired, multichambered, and connected with the sperm duct, and have been reported to play glandular and storage functions. Seminal vesicle secretion may include steroids and steroid glucuronides, with hormonal and pheromonal functions, but it appears to be primarily constituted of mucoproteins, acid mucopolysaccharides, and phospholipids.
Fish ovaries may be of two types - gymnovarian or cystovarian. In the first type, the oocytes are released directly into the coelomic cavity and then eliminated. In the second type, the oocytes are conveyed to the exterior through the oviduct. Many catfish are cystovarian in type, including ''Pseudoplatystoma corruscans'', ''Pseudoplatystoma fasciatum, P. fasciatum'', ''Lophiosilurus alexandri'', and ''Loricaria lentiginosa''.
Communication
Catfish can produce different types of sounds and also have well-developed auditory reception used to discriminate between sounds with different pitches and velocities. They are also able to determine the distance of the sound's origin and from what direction it originated. This is a very important fish communication mechanism, especially during agonistic behavior, agonistic and distress behaviors. Catfish are able to produce a variety of sounds for communication that can be classified into two groups: drumming sounds and stridulation sounds. The variability in catfish sound signals differs due to a few factors: the mechanism by which the sound is produced, the function of the resulting sound, and physiological differences such as size, sex, and age. To create a drumming sound, catfish use an indirect vibration mechanism using a swimbladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth w ...
. In these fishes, sonic muscles insert on the ramus Mulleri, also known as the elastic spring. The sonic muscles pull the elastic spring forward and extend the swimbladder. When the muscles relax, the tension in the spring quickly returns the swimbladder to its original position, which produces the sound.
Catfish also have a sound-generating mechanism in their pectoral fins. Many species in the catfish family possess an enhanced first pectoral fin ray, called the spine, which can be moved by large Abduction (kinesiology), abductor and Adduction, adductor muscles. The base of the catfishes' spines has a sequence of ridges, and the spine normally slides within a groove on the fish's pelvic girdle during routine movement; but, pressing the ridges on the spine against the pelvic girdle groove creates a series of short pulses. The movement is analogous to a finger moving down the teeth of a comb, and consequently a series of sharp taps is produced.
Sound-generating mechanisms are often different between the sexes. In some catfish, pectoral fins are longer in males than in females of similar length, and differences in the characteristic of the sounds produced were also observed. Comparison between families of the same order of catfish demonstrated family and species-specific patterns of vocalization, according to a study by Maria Clara Amorim. During courtship behavior in three species of ''Corydoras'' catfish, all males actively produced stridulation sounds before egg fertilization, and the species' songs were different in pulse number and sound duration.
Sound production in catfish may also be correlated with fighting and alarm calls. According to a study by Kaatz, sounds for disturbance (e.g. alarm) and agonistic behavior were not significantly different, which suggests distress sounds can be used to sample variation in agonistic sound production. However, in a comparison of a few different species of tropical catfish, some fish put under distress conditions produced a higher intensity of stridulatory sounds than drumming sounds. Differences in the proportion of drumming versus stridulation sounds depend on morphology (biology), morphological constraints, such as different sizes of drumming muscles and pectoral spines. Due to these constraints, some fish may not even be able to produce a specific sound. In several different species of catfish, aggressive sound production occurs during cover site defense or during threats from other fish. More specifically, in long-whiskered catfish, drumming sounds are used as a threatening signal and stridulations are used as a defense signal. Kaatz investigated 83 species from 14 families of catfish, and determined that catfish produce more stridulatory sounds in disturbance situations and more swimbladder sounds in intraspecific conflicts.
Economic importance
Aquaculture
Catfish are easy to farm in warm climates, leading to inexpensive and safe food at local grocers. About 60% of U.S. farm-raised catfish are grown within a 65-mile (100-km) radius of Belzoni, Mississippi. Channel catfish (''Ictalurus punctatus'') supports a $450 million/yr aquaculture industry.[ The largest producers are located in the Southern United States, including Mississippi, Alabama, and Arkansas.
Catfish raised in inland tanks or channels are usually considered safe for the environment, since their waste and disease should be contained and not spread to the wild.
In Asia, many catfish species are important as food. Several airbreathing catfish (Clariidae) and shark catfish (Pangasiidae) species are heavily cultured in Africa and Asia. Exports of one particular shark catfish species from Vietnam, ''Pangasius bocourti'', have met with pressures from the U.S. catfish industry. In 2003, The United States Congress passed a law preventing the imported fish from being labeled as catfish. As a result, the Vietnamese exporters of this fish now label their products sold in the U.S. as "basa fish." Trader Joe's has labeled frozen fillets of Vietnamese ''Pangasius hypophthalmus'' as "striper."
There is a large and growing ornamental fish trade, with hundreds of species of catfish, such as '']Corydoras
''Corydoras'' is a genus of freshwater catfish in the family Callichthyidae and subfamily Corydoradinae. The species usually have more restricted areas of endemism than other callichthyids, but the area of distribution of the entire genus almost ...
'' and Loricariidae, armored suckermouth catfish (often called plecos), being a popular component of many aquarium, aquaria. Other catfish commonly found in the aquarium trade are Aspredinidae, banjo catfish, Doradidae, talking catfish, and Pimelodidae, long-whiskered catfish.
Catfish as food
Catfish have widely been caught and farmed for food for hundreds of years in Africa, Asia, Europe, and North America. Judgments as to the quality and flavor vary, with some food critics considering catfish excellent to eat, while others dismiss them as watery and lacking in flavor.[Jenny Baker (1988), ''Simply Fish'' p 36-37. Faver & Faber, London.] Catfish is high in vitamin D. Farm-raised catfish contains low levels of omega-3 fatty acids and a much higher proportion of omega-6 fatty acids.
In Central Europe, catfish were often viewed as a delicacy to be enjoyed on Feast Day, feast days and holidays. Migrants from Europe and Africa to the United States brought along this tradition, and in the Southern United States, catfish is an extremely popular food.
The most commonly eaten species in the United States are the channel catfish and the blue catfish, both of which are common in the wild and increasingly widely farmed. Farm-raised catfish became such a staple of the U.S. diet that President Ronald Reagan established National Catfish Day on June 25, 1987 to recognize "the value of farm-raised catfish."
Catfish is eaten in a variety of ways. In Europe, it is often cooked in similar ways to carp
Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
, but in the United States it is popularly crumbed with cornmeal and fried.[
In Indonesia, catfish is usually served fried or grilled in street stalls called ''warung'' and eaten with vegetables, sambal (a spicy relish or sauce), and usually ''nasi uduk'' (traditional coconut rice). The dish is called ''pecel lele'' or ''pecak lele''. ''Lele'' is the Indonesian language, Indonesian word for catfish. The same dish can also be called as ''lele penyet'' (squashed catfish) if the fish is lightly squashed along with sambal with a stone Mortar and pestle, mortar-and-pestle. The ''pecel''/''pecak'' version presents the fish in a separate plate while the mortar is solely for sambal.
In Malaysia, catfish is called ''ikan keli'' and is fried with spices or grilled and eaten with tamarind and Bird's eye chili, Thai chili gravy and is also often eaten with steamed rice.
In Bangladesh and the ]India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
n states of Odisha, West Bengal and Assam, catfish (locally known as ''magur'') is eaten as a favored delicacy during the Monsoon, monsoons. In the Indian state of Kerala, the local catfish, known as ''thedu or ''etta'' in Malayalam, is also popular.
In Hungary, catfish is often cooked in paprika sauce (Harcsapaprikás) typical of Hungarian cuisine. It is traditionally served with pasta smothered with curd cheese (túrós csusza).
In Myanmar (formally Burma), catfish is usually used in ''mohinga'', a traditional noodle fish soup cooked with lemon grass, ginger, garlic, pepper, banana stem, onions, and other local ingredients.
Vietnamese catfish, of the genus ''Pangasius'', cannot be legally marketed as catfish in the United States, and so is referred to as Iridescent shark, ''swai'' or ''Basa (fish), basa''. Only fish of the family ''Ictaluridae'' may be marketed as catfish in the United States.[Public Law 107-171, § 10806, 116 United States Statutes at Large, Stat. 526-527, codified in and ] In the UK, Vietnamese catfish is sometimes sold as "Vietnamese river cobbler", although more commonly as Basa.
In Nigeria, catfish is often cooked in a variety of Stew, stews. It is particularly cooked in a delicacy popularly known as "catfish pepper soup" which is enjoyed throughout the nation.
In Jewish dietary law, known as kashrut, fish must have fins and scales to be kashrut, kosher. Since catfish lacks scales, they are not kosher.
Dangers to humans
While the vast majority of catfish are harmless to humans, a few species are known to present some risk. Many catfish species have "stings" (actually non-venomous in most cases) embedded behind their fins; thus precautions must be taken when handling them. Stings by the venomous striped eel catfish have killed people in rare cases.
Taxonomy
The catfish are a monophyly, monophyletic group. This is supported by molecular evidence.
Catfish belong to a superorder called the Ostariophysi, which also includes the Cypriniformes, Characiformes, Gonorynchiformes and Gymnotiformes, a superorder characterized by the Weberian apparatus
The Weberian apparatus is an anatomical structure that connects the swim bladder to the auditory system in fishes belonging to the superorder Ostariophysi. When it is fully developed in adult fish, the elements of the apparatus are sometimes col ...
. Some place Gymnotiformes as a sub-order of Siluriformes, however this is not as widely accepted. Currently, the Siluriformes are said to be the sister group to the Gymnotiformes, though this has been debated due to more recent molecular evidence. there are about 36 extant taxon, extant catfish families, and about 3,093 extant species have been described.[ This makes the catfish order the second or third most diverse vertebrate order; in fact, 1 out of every 20 vertebrate species is a catfish.]
The taxonomy of catfish is quickly changing. In a 2007 and 2008 paper, ''Horabagrus'', ''Phreatobius'', and ''Conorhynchos'' were not classified under any current catfish families. There is disagreement on the family status of certain groups; for example, Nelson (2006) lists Auchenoglanididae and Heteropneustidae as separate families, while the All Catfish Species Inventory (ACSI) includes them under other families. Also, FishBase and the Integrated Taxonomic Information System lists Parakysidae as a separate family, while this group is included under Akysidae by both Nelson (2006) and ACSI. Many sources do not list the recently revised family Anchariidae. The family Horabagridae, including ''Horabagrus'', ''Pseudeutropius'', and ''Platytropius'', is also not shown by some authors but presented by others as a true group. Thus, the actual number of families differs between authors. The species count is in constant flux due to Taxonomy (biology), taxonomic work as well as description of new species. On the other hand, our understanding of catfish should increase in the next few years due to work by the ACSI.
Between 2003 and 2005, over 100 species have been named, a rate three times faster than that of the past century. In June 2005, researchers named the newest family of catfish, Lacantuniidae, only the third new family of fish distinguished in the last 70 years (others being the coelacanth in 1938 and the megamouth shark in 1983). The new species in Lacantuniidae, ''Lacantunia enigmatica'', was found in the Lacantun river in the Mexican state of Chiapas.
The higher-level phylogeny of Siluriformes has gone through several recent changes, mainly due to Molecular phylogenetics, molecular phylogenetic studies. While most studies, both morphological and molecular, agree that catfishes are arranged into three main Lineage (evolution), lineages, the relationship among these lineages has been a contentious point in which morphological and molecular phylogenetic studies, performed for example by Rui Diogo, differ. The three main lineages in Siluriformes are the family Diplomystidae, the denticulate catfish suborder Loricarioidea, Loricarioidei (which includes the families Nematogenyidae, Trichomycteridae
Trichomycteridae is a family of catfishes commonly known as pencil catfishes or parasitic catfishes. This family includes the candiru fish (''Vandellia cirrhosa''), feared by some people for its alleged habit of entering into the urethra of hum ...
, Callichthyidae
Callichthyidae is a family of catfishes ( order Siluriformes), called armored catfishes due to the two rows of bony plates (or scutes) along the lengths of their bodies. It contains some of the most popular freshwater aquarium fish, such as many ...
, Scoloplacidae, Astroblepidae
''Astroblepus'' is a genus of fish in the family Astroblepidae found in South America and Panama. This genus is the only member of its family. These catfishes are primarily found in torrential streams in the Andean area. ''Astroblepus pholeter'' ...
, and Loricariidae
The Loricariidae is the largest family of catfish (order Siluriformes), with 92 genera and just over 680 species. Loricariids originate from freshwater habitats of Costa Rica, Panama, and tropical and subtropical South America. These fish are not ...
, which is sometimes referred to as the superfamily Loricarioidea), and the suborder Siluroidei, which contains the remaining families of the order. According to morphology (biology), morphological data, Diplomystidae is usually considered to be the earliest branching catfish lineage and the cladistics, sister group to the other two lineages, Loricarioidei and Siluroidei. Molecular evidence usually contrasts with this hypothesis, and shows the suborder Loricarioidei as the earliest branching catfish lineage, and sister to a clade that includes the Diplomystidae and Siluroidei. While in the first study this relationship was proposed the "morphological" hypothesis could not be rejected, the new, "molecular" phylogenetic hypothesis was later obtained in numerous other phylogenetic studies based on genetic data. However, a recent study based on molecular data argued that the previous "molecular" hypothesis is the result of phylogenetic artifacts due to a strong heterogeneity in Rate of evolution, evolutionary rates among siluriform lineages. In that study it was suggested that the fast evolution of the Loricarioidei suborder was attracting this clade to the outgroups through long branch attraction, incorrectly placing it as the earliest-branching catfish lineage. When a data filtering method was used to reduce lineage rate heterogeneity (the potential source of bias) on their dataset, a final phylogeny was recovered which showed the Diplomystidae are the earliest-branching catfish, followed by Loricarioidea, Loricarioidei and Siluroidei as sister lineages. Thus, there is currently both morphological ''and'' molecular evidence for a higher-level phylogenetic arrangement of Siluriformes in which Diplomystidae is the earliest branching catfish, sister to a clade including the Loricarioidea, Loricarioidei and Siluroidei suborders.
Below is a list of family relationships by different authors. Lacantuniidae is included in the Sullivan scheme based on recent evidence that places it sister to Claroteidae.
Phylogeny
Phylogeny of living Siluriformes based on 2017 and extinct families based on Nelson, Grande & Wilson 2016.
Unassigned families:
* Scoloplacidae (Loricarioidei)
* Akysidae (Sisoroidea)
* Amblycipitidae (Sisoroidea)
* Anchariidae (Arioidea)
* Ariidae
The Ariidae or ariid catfish are a family of catfish that mainly live in marine waters with many freshwater and brackish water species. They are found worldwide in tropical to warm temperate zones. The family includes about 143 species.
Taxonomy ...
(Arioidea)
* Amphiliidae
The loach catfishes are a family, Amphiliidae, of catfishes ( order Siluriformes). They are widespread in tropical Africa, but are most common in streams at high elevations; most species are able to cling to rocks in fast-flowing streams. The 13 ...
(Big African catfishes)
* Austroglanididae (Arioidea)
* Chacidae (Siluroidei)
* ''Conorhynchos'' (Pimelodoidea)
* Cranoglanididae (Ictaluroidea)
* Heteropneustidae (Clarioidea)
* Horabagridae (Sisoroidea)
* Kryptoglanidae (Siluroidea)
* Lacantuniidae (Big African catfishes)
* Malapteruridae
Electric catfish or Malapteruridae is a Family (biology), family of catfishes (order Siluriformes). This family includes two genus, genera, ''Malapterurus'' and ''Paradoxoglanis'', with 21 species. Several species of this family have the ability ...
(Big African catfishes)
* Phreatobiidae (Pimelodoidea)
* ''Rita (fish), Rita'' (Sisoroidea)
* Schilbeidae (Big African catfishes)
Timeline
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from: -99.6 till: -65.5 color:latecretaceous text:Late Cretaceous, Late
from: -65.5 till: -55.8 color:paleocene text:Paleocene, Paleo.
from: -55.8 till: -33.9 color:eocene text:Eocene, Eo.
from: -33.9 till: -23.03 color:oligocene text:Oligocene, Oligo.
from: -23.03 till: -5.332 color:miocene text:Miocene, Mio.
from: -5.332 till: -2.588 color:pliocene text:Pliocene, Pl.
from: -2.588 till: -0.0117 color:pleistocene text:Pleistocene, Pl.
from: -0.0117 till: 0 color:holocene text:Holocene, H.
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from: -145.5 till: -65.5 color:cretaceous text:Cretaceous
from: -65.5 till: -23.03 color:paleogene text:Paleogene
from: -23.03 till: -2.588 color:neogene text:Neogene
from: -2.588 till: 0 color:quaternary text:Quaternary, Q.
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from: -99.6 till: -65.5 color:latecretaceous text:Late Cretaceous, Late
from: -65.5 till: -55.8 color:paleocene text:Paleocene, Paleo.
from: -55.8 till: -33.9 color:eocene text:Eocene, Eo.
from: -33.9 till: -23.03 color:oligocene text:Oligocene, Oligo.
from: -23.03 till: -5.332 color:miocene text:Miocene, Mio.
from: -5.332 till: -2.588 color:pliocene text:Pliocene, Pl.
from: -2.588 till: -0.0117 color:pleistocene text:Pleistocene, Pl.
from: -0.0117 till: 0 color:holocene text:Holocene, H.
bar:era
from: -145.5 till: -65.5 color:cretaceous text:Cretaceous
from: -65.5 till: -23.03 color:paleogene text:Paleogene
from: -23.03 till: -2.588 color:neogene text:Neogene
from: -2.588 till: 0 color:quaternary text:Quaternary, Q.
Catfish fishing records
By information from International Game Fish Association IGFA the most outstanding record:
* The biggest flathead catfish caught was by Ken Paulie in the Elk City Reservoir in Kansas, USA on 19 May 1998 that weighed 55.79 kg (123 lbs. 0 oz.)
References
External links
All catfish species inventory
"Giant Baghair caught in Jamuna"
in ''The Daily Star'' (Bangladesh), 12 May 2009
* Skelton, Paul H. and Teugels, Guy G. 1992
Ichthyological Bulletin; No. 56: Neotype description for the African catfish Clarias Gariepinus (Burchell, 1822) (Pisces: Siluroidei: Clariidae)
J.L.B. Smith Institute of Ichthyology, Rhodes University, Grahamstown, South Africa
{{Authority control
Siluriformes, .
Edible fish
Commercial fish
Ostariophysi
Late Cretaceous fish
Extant Late Cretaceous first appearances
Taxa named by Georges Cuvier