''Canaanimico'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of medium-sized New World monkeys from the Late Oligocene (approximately 26.5 Ma,

Deseadan The Deseadan ( es, Deseadense) age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It fo ...

in the SALMA classification) fossiliferous fluvio- lacustrine

Chambira Formation The Chambira River is a major tributary of the Marañón River, and has been the traditional territory of the Urarina peoples for at least the past 350 years, if not longer. Located in the Amazon jungle of Peru, otherwise known as the Selva, the Ch ...

of the

Ucayali Basin The Ucayali River ( es, Río Ucayali, ) is the main headstream of the Amazon River. It rises about north of Lake Titicaca, in the Arequipa region of Peru and becomes the Amazon at the confluence of the Marañón close to Nauta city. The city ...

in Amazonian

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

. The genus was described by Marivaux et al. in 2016 and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''C. amazonensis''.Marivaux et al., 2016''Canaanimico''
at

Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...

.org

Classification

Marivaux et al. placed the genus in the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Soriacebinae, of the family Homunculidae,Homunculidae
at

.org while Silvestro et al. (2017) attribute an '' incertae sedis'' status for the family and subfamily to the genus.Silvestro et al., 2017, p.13 Marivaux et al. proposed a close relation with the

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

South American genera '' Soriacebus'' (

Santacrucian The Santacrucian age is a period of geologic time (17.5 – 16.3 Ma) within the Early Miocene epoch of the Neogene, used more specifically with SALMA classification in South America. It follows the Colhuehuapian and precedes the Friasian age. ...

) and '' Mazzonicebus'' (

Colhuehuapian The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian ag ...

).Urbani et al., 2018, p.33

Description

''Canaanimico'' was described by Marivaux et al. on the basis of two isolated upper molars collected at the Contamana fossil locality. The authors inferred that based on

dental microwear Dental microwear analysis is a method to infer diet and behavior in extinct animals, especially in fossil specimens. Typically, the patterns of pits and scratches on the occlusal or buccal surface of the enamel are compared with patterns observed ...

patterns recorded on one upper molar ''Canaanimico'' possibly was a fruit and hard-object eater. Silvestro et al. estimated a body mass of for ''Canaanimico'', making it a medium-sized New World monkey.Silvestro et al., 2017, p.14

Paleoecology

Fossils of ''Canaanimico'' were found in the Late Oligocene (

)

of the

in Amazonian Peru. The formation, a sequence of red

claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone, slate, and shale. Most of the particles of which the stone is composed are less than and are too sm ...

s and paleosols with intercalated conglomerates and

sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...

s,Antoine et al., 2016, p.40 and

gypsum Gypsum is a soft sulfate mineral composed of calcium sulfate dihydrate, with the chemical formula . It is widely mined and is used as a fertilizer and as the main constituent in many forms of plaster, blackboard or sidewalk chalk, and drywall. ...

layers,Castro Medina, 2010, p.55 has been dated on the basis of

zircon Zircon () is a mineral belonging to the group of nesosilicates and is a source of the metal zirconium. Its chemical name is zirconium(IV) silicate, and its corresponding chemical formula is Zr SiO4. An empirical formula showing some of the r ...

s in a

tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock cont ...

bed in the formation, providing an age of 26.56 ± 0.07 Ma.Antoine et al., 2016, p.35 The formation was deposited in a

tectonically Tectonics (; ) are the processes that control the structure and properties of the Earth's crust and its evolution through time. These include the processes of mountain building, the growth and behavior of the strong, old cores of continents k ...

relatively calmCastro Medina, 2010, p.71 tropical fluvio- lacustrine

environment Environment most often refers to: __NOTOC__ * Natural environment, all living and non-living things occurring naturally * Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...

with

oxbow lake An oxbow lake is a U-shaped lake or pool that forms when a wide meander of a river is cut off, creating a free-standing body of water. In South Texas, oxbows left by the Rio Grande are called '' resacas''. In Australia, oxbow lakes are call ...

s.Antoine et al., 2016, p.41Boivin et al., 2016, p.103 The Chambira Formation has provided a rich mammal assemblage of '' Chambiramys shipiborum'', '' C. sylvaticus'', '' Deseadomys cf. arambourgi'', '' Loretomys minutus'', '' Palaeosteiromys amazonensis'', '' Plesiosteiromys newelli'', '' Scleromys praecursor'', '' Ucayalimys crassidens'', '' Abderites sp.'', '' aff. Eosallamys sp.'', '' cf. Neoglyptatelus sp.'', and indeterminate Adelphomyinae,

Anthropoidea The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders New World monkeys (Platyrrhini) and Catarrhi ...

, Astrapotheriidae, Caenolestidae, Caviomorpha,

Chinchilloidea Caviomorpha is the rodent infraorder or parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now ...

, Emballonuridae,

Erethizontoidea The New World porcupines, family Erethizontidae, are large arboreal rodents, distinguished by their spiny coverings from which they take their name. They inhabit forests and wooded regions across North America, and into northern South America. A ...

Herpetotheriidae Herpetotheriidae is an extinct family of metatherians, closely related to marsupials. Species of this family are generally reconstructed as terrestrial, and are considered morphologically similar to modern opossums. Fossils of herpetotheriids com ...

Interatheriinae Interatheriinae is an extinct subfamily of interatheriids that consisted of notoungulates dating from the Early Eocene (Casamayoran SALMA) to the Early Pliocene (Montehermosan SALMA). The subfamily includes the genera '' Archaeophylus'', '' Argy ...

Litopterna Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until the G ...

, Marsupialia, Microbiotheria,

Mylodontidae Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters a ...

, Mylodontoidea, Notoungulata,

Octodontoidea Caviomorpha is the rodent infraorder or parvorder that unites all New World hystricognaths. It is supported by both fossil and molecular evidence. The Caviomorpha was for a time considered to be a separate order outside the Rodentia, but is now ...

, Palaeothentidae, Pampatheriidae,

Rhinolophoidea Rhinolophoidea is a superfamily of bats. It contains the following families: Craseonycteridae, Hipposideridae, Megadermatidae, Rhinolophidae, Rhinonycteridae, and Rhinopomatidae. It is one of two superfamilies that comprise the suborder Yinpte ...

Tolypeutinae Tolypeutinae is a subfamily of armadillos in the family Chlamyphoridae, consisting of the giant, three-banded and naked-tailed armadillos. __TOC__ Taxonomy It contains the following genera: *''Cabassous'' *''Kuntinaru'' *''Priodontes'' *''To ...

, Toxodontidae,

Typotheria Typotheria is a suborder of the extinct mammalian order Notoungulata and includes five families: Archaeopithecidae, Campanorcidae, Interatheriidae, Mesotheriidae, and Oldfieldthomasiidae. CifelliCifelli, Richard L. 1993. The phylogeny of the nat ...

and

Vespertilionoidea Vespertilionoidea is a superfamily of bats containing five families: Cistugidae, Miniopteridae, Molossidae, Natalidae, and Vespertilionidae. It is one of three superfamilies in the suborder Yangochiroptera, the others being Noctilionoidea Noc ...

.Boivin et al., 2016CTA-61
at

.orgCTA-32
at

.org Additionally, crocodylians and

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...

s of '' ?Balanerodus sp.'', '' Podocnemis sp.'', '' cf. Purussaurus sp.'', '' cf. Sebecus sp.'', and indeterminate frogs,

Booidea The Booidea, also known as booid snakes, are a superfamily of snakes that contains boas (family Boidae) and other closely related boa-like snakes (but not pythons, which are in a separate superfamily called Pythonoidea). As of 2017, Booidea conta ...

, Caimaninae,

Colubroidea Colubroidea is a superfamily of snakes in the clade Colubroides that includes Colubridae, with some studies splitting Colubridae into multiple families that make up Colubroidea. Historically, Colubroidea also included other caenophidian snakes su ...

Gavialoidea Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are ...

and

snakes Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joi ...

were found in the formation, as well as fossil fishes of '' cf. Hydrolycus sp.'', '' Leporinus sp.'', '' cf. Phractocephalus sp.'', '' Potamotrygon sp.'', and indeterminate

Actinopterygii Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...

Cichlidae Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted thi ...

Erythrinidae The Erythrinidae are a family of fishes found in rivers and other freshwater habitats from Costa Rica south as far as Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the s ...

, Loricariidae and

Pimelodidae The Pimelodidae, commonly known as the long-whiskered catfishes, are a family of catfishes ( order Siluriformes). Taxonomy The family Pimelodidae has undergone much revision. Currently, it contains about 30 genera and about 90 recognized and kno ...

References

Bibliography

* * * * * *

Classification

Description

Paleoecology

See also

References

Bibliography

Further reading