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Mylodontoidea
Sloths are a group of Neotropical xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America. Sloths are considered to be most closely related to anteaters, together making up the xenarthran order Pilosa. There are six extant sloth species in two genera – '' Bradypus'' (three–toed sloths) and '' Choloepus'' (two–toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb-- although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via parallel evolution from terrestrial ancestors. Besides the extant species, many species of ground sloths ranging up to the size of ele ...
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Ground Sloth
Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbean ground sloths, the most recent survivors, lived in the Antilles, possibly until 1550 BCE. However, radiocarbon dating suggests an age of between 2819 and 2660 BCE for the last occurrence of ''Megalocnus'' in Cuba. Ground sloths had been extinct on the mainland of North and South America for 10,000 years or more. They survived 5,000–6,000 years longer in the Caribbean than on the American mainland, which correlates with the later colonization of this area by humans. Much ground sloth evolution took place during the late Paleogene and Neogene of South America, while the continent was isolated. At their earliest appearance in the fossil record, the ground sloths were already distinct at the family level. The presence ...
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Phyllophaga (mammal)
Sloths are a group of Neotropical xenarthran mammals constituting the suborder Folivora, including the extant arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of movement, tree sloths spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America. Sloths are considered to be most closely related to anteaters, together making up the xenarthran order Pilosa. There are six extant sloth species in two genera – '' Bradypus'' (three–toed sloths) and '' Choloepus'' (two–toed sloths). Despite this traditional naming, all sloths have three toes on each rear limb-- although two-toed sloths have only two digits on each forelimb. The two groups of sloths are from different, distantly related families, and are thought to have evolved their morphology via parallel evolution from terrestrial ancestors. Besides the extant species, many species of ground sloths ranging up to the size of ele ...
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Scelidotheriidae
Scelidotheriidae is a family of extinct ground sloths within the order Pilosa, suborder Folivora and superfamily Mylodontoidea, related to the other extinct mylodontoid family, Mylodontidae, as well as to the living two-toed sloth family Choloepodidae. The only other extant family of the suborder Folivora is the distantly related Bradypodidae. Erected as the family Scelidotheriidae by Ameghino in 1889, the taxon was demoted to a subfamily by Gaudin in 1995. However, recent collagen sequence data indicates the group is less closely related to ''Mylodon'' and ''Lestodon'' than '' Choloepus'' is, and thus it has been elevated back to full family status by Presslee ''et al.'' (2019). Taxonomy Together with Mylodontidae, and the two-toed sloths, the scelidotheriids form the superfamily Mylodontoidea. ''Chubutherium'' is an ancestral and very plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all m ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals . They are the only members of the genus ''Bradypus'' and the family Bradypodidae. The four living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years ago. Relation to the two-toed sloth Both types of sloth t ...
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Pilosa
The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters and sloths (which includes the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The biogeographic origins of the Pilosa are still unclear, but they can be traced back in South America as far as the early Paleogene (about 60 million years ago, only a short time after the end of the Mesozoic Era). The presence of these animals in Central America and their former presence in North America is a result of the Great American Interchange. A number of sloths were also formerly present on the Antilles, which they reached from South America by some combination of rafting or floating with the prevailing currents. Together with the armadillos, which are in the order Cingulata, pilosans are part of the larger superorder Xenarthra, a defining characteristic of which is the presence of xenarthrals (extra formations between lumbar verte ...
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Three-toed Sloth
The three-toed or three-fingered sloths are arboreal neotropical mammals . They are the only members of the genus ''Bradypus'' and the family Bradypodidae. The four living species of three-toed sloths are the brown-throated sloth, the maned sloth, the pale-throated sloth, and the pygmy three-toed sloth. In complete contrast to past morphological studies, which tended to place ''Bradypus'' as the sister group to all other folivorans, molecular studies place them nested within the sloth superfamily Megatherioidea, making them the only surviving members of that radiation. Extant species Evolution A study of mitochondrial cytochrome b and 16S rRNA sequences suggests that '' B. torquatus'' diverged from '' B. variegatus'' and '' B. tridactylus'' about 12 million years ago, while the latter two split 5 to 6 million years ago. The diversification of ''B. variegatus'' lineages was estimated to have started 4 to 5 million years ago. Relation to the two-toed sloth Both types of sloth t ...
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Mylodontidae
Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters and sloths (which includes the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The b ..., living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another family of extinct ground sloths, Scelidotheriidae, as well as to the extant arboreal two-toed sloths, family Choloepus, Choloepodidae; together these make up the superfamily Mylodontoidea. Phylogenetic analyses based on morphology uncovered the relationship between Mylodontidae and Scelidotheriidae; in fact, the latter was for a time considered a subfamily of mylodontids. However, molecular sequence comparisons were needed for the correct placement of Choloepodidae. These studies have been car ...
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Nothrotheriidae
Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately . Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built ''Nothrotheriops'', which reached a length of about . While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today. Evolution During the late Miocene and Pliocene, the sloth genus ''Thalassocnus'' of the west coast of South America became adapted to a shallow-water marine lifestyle. However, the family placement of ''Thalassocnus'' has been disputed; while long considered a nothrotheriid, one recent analysis moves it to ...
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Megatheriidae
Megatheriidae is a family of extinct ground sloths that lived from approximately 23 mya—11,000 years ago. Megatheriids appeared during the Late Oligocene (Deseadan in the SALMA classification), some 29 million years ago, in South America. The group includes the heavily built '' Megatherium'' (given its name 'great beast' by Georges Cuvier) and ''Eremotherium''. An early genus that was originally considered a megatheriid, the more slightly built ''Hapalops'', reached a length of about . The nothrotheres have recently been placed in their own family, Nothrotheriidae. The skeletal structure of these ground sloths indicates that the animals were massive. Their thick bones and even thicker joints (especially those on the hind legs) gave their appendages tremendous power that, combined with their size and fearsome claws, provided a formidable defense against predators. The earliest megatheriid in North America was ''Sibotherium'' which arrived 5.3 million years ago, after cros ...
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Megalonychidae
Megalonychidae is an extinct family of sloths including the extinct ''Megalonyx''. Megalonychids first appeared in the early Oligocene, about 35 million years (Ma) ago, in southern Argentina (Patagonia). There is actually one possible find dating to the Eocene, about 40 Ma ago, on Seymour Island in Antarctica (which was then still connected to South America). They first reached North America by island-hopping across the Central American Seaway, about 9 million years ago, prior to formation of the Isthmus of Panama about 2.7 million years ago (which led to the main pulse of the Great American Interchange). Some megalonychid lineages increased in size as time passed. The first species of these were small and may have been partly tree-dwelling, whereas the Pliocene (about 5 to 2 million years ago) species were already approximately half the size of the huge Late Pleistocene ''Megalonyx jeffersonii'' from the last ice age.J.L. White (1993) It was formerly believed, based on morpholo ...
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Megalocnidae
Megalocnidae is an extinct family of sloths, native to the islands of the Greater Antilles from the Early Oligocene to the Mid-Holocene. They are known from Cuba, Hispaniola and Puerto Rico, but are absent from Jamaica. While they were formerly placed in the Megalonychidae alongside two-toed sloths and ground sloths like ''Megalonyx'', recent mitochondrial DNA and collagen sequencing studies place them as the earliest diverging group basal to all other sloths. They displayed significant diversity in body size and lifestyle, with ''Megalocnus'' being terrestrial and probably weighing several hundred kilograms, while ''Neocnus'' was likely arboreal and similar in weight to extant tree sloths, at less than 10 kilograms. Origin It is thought that sloths arrived in the Caribbean from South America (where they arose) around the Eocene-Oligocene boundary about 33 million years ago, when there was a significant sea level drop caused by a glaciation episode. This has been associate ...
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Two-toed Sloth
''Choloepus'' is a genus of xenarthran mammals of Central and South America within the monotypic family Choloepodidae, consisting of two-toed sloths, sometimes also called two-fingered sloths. The two species of ''Choloepus'' (which means "lame foot"), Linnaeus's two-toed sloth (''Choloepus didactylus'') and Hoffmann's two-toed sloth (''Choloepus hoffmanni''), were formerly believed on the basis of morphological studies to be the only surviving members of the sloth family Megalonychidae, but have now been shown by molecular results to be closest to extinct ground sloths of the family Mylodontidae. Extant species Evolution A study of retrovirus and mitochondrial DNA suggests that ''C. didactylus'' and ''C. hoffmani'' diverged 6 to 7 million years ago. Furthermore, based on cytochrome c oxidase subunit I sequences, a similar divergence date ( years ago) between the two populations of ''C. hofmanni'' separated by the Andes has been reported. Their ancestors evolved with mari ...
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