Megatheriidae is a family of

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

ground sloth Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribb ...

s that lived from approximately 23 mya—11,000 years ago. Megatheriids appeared during the Late Oligocene (

Deseadan The Deseadan ( es, Deseadense) age is a period of geologic time (29.0–21.0 Ma) within the Oligocene epoch of the Paleogene to the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification of South America. It fol ...

in the SALMA classification), some 29 million years ago, in

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...

. The group includes the heavily built '' Megatherium'' (given its name 'great beast' by

Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French naturalist and zoologist, sometimes referred to as the "founding father of paleontology". Cuvier was a major figure in na ...

) and ''

Eremotherium ''Eremotherium'' (from Greek for "steppe" or "desert beast": ἔρημος "steppe or desert" and θηρίον "beast") is an extinct genus of giant ground sloth, in the family Megatheriidae, the largest and most heavily built family of sloths. ' ...

''. An early genus that was originally considered a megatheriid, the more slightly built ''

Hapalops ''Hapalops'' is an extinct genus of ground sloth from the Early to Late Miocene of Brazil (Solimões Formation), Bolivia ( Honda Group), Colombia ( Honda Group),Croft, 2007, p.300 and Argentina (Santa Cruz Formation) in South America.

'', reached a length of about . The nothrotheres have recently been placed in their own family,

Nothrotheriidae Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately . Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, th ...

. The skeletal structure of these ground sloths indicates that the animals were massive. Their thick

bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...

s and even thicker

joint A joint or articulation (or articular surface) is the connection made between bones, ossicles, or other hard structures in the body which link an animal's skeletal system into a functional whole.Saladin, Ken. Anatomy & Physiology. 7th ed. McGraw- ...

s (especially those on the hind legs) gave their appendages tremendous power that, combined with their size and fearsome

claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...

s, provided a formidable defense against predators. The earliest megatheriid in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...

was '' Sibotherium'' which arrived 5.3 million years ago, after crossing the recently formed

Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Co ...

nian land bridge. At more than five tons in weight, 6 meters in length, and able to reach as high as , ''

'' was taller than an

African bush elephant The African bush elephant (''Loxodonta africana'') is one of two extant African elephant species and one of three extant elephant species. It is the largest living terrestrial animal, with bulls reaching a shoulder height of up to and a body ...

bull. Unlike relatives, this species retained a

plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, ...

ic extra claw. While other species of ''Eremotherium'' had four fingers with only two or three claws, ''E. eomigrans'' had five fingers, four of them with claws up to nearly a foot long.De Iuliis and Cartelle (1999)

Phylogeny

The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data (see Fig. 4 of Presslee ''et al''., 2019). The following phylogeny is based on Varela ''et al''. 2019.

References

External links

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CTD: Megatheriidae

Memidex: Megatheriidae
{{Taxonbar, from=Q784732 Prehistoric mammal families Burdigalian first appearances Holocene extinctions Neogene mammals of South America Neogene mammals of North America Pleistocene mammals of South America Pleistocene mammals of North America Taxa named by John Edward Gray