Brotia Swinhoei
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''Brotia'' is a genus of Southeast Asian
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s,
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s in the
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea: Pachychilidae)". '' Malacologia'' 48
159
251.
The generic name ''Brotia'' is apparently in honor of a Swiss malacologist
Auguste Louis Brot Auguste Louis Brot (September 18, 1821 ''Sevue Suisse de zoologies'', vol. 4, 1896-189"Auguste Louss Brot"/ref>- August 30, 1896) was a Swiss malacologist ( conchologist). ''The American Naturalist'', March 1897p.270/ref> After several years of st ...
(1821-1896).


Distribution

Species of ''Brotia'' occur in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
habitats of Southeast Asia, ranging from Northern India in the west through to
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
in the east. This range includes India,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...
, Bangladesh, Thailand,
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
, Vietnam,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
(1 species), Cambodia, Malaysia and Indonesia (
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
and Borneo only).


Biology

Species are found predominantly in fast flowing, well oxygenated rivers, occasionally also in lakes. They are
gonochoristic In biology, gonochorism is a sexual system where there are only two sexes and each individual organism is either male or female. The term gonochorism is usually applied in animal species, the vast majority of which are gonochoric. Gonochorism c ...
, and viviparous, retaining developing eggs and youngs in special brood pouch.


Species

Species within the genus ''Brotia'' include: * † '' Brotia alpina'' (F. Sandberger, 1871) * '' Brotia annamita'' Köhler, Holford M., Do & Ho, 2009Köhler F., Holford, Do V. T. & Ho T. H. (2009). "Exploring a largely unknown fauna: On the diversity of pachychilid freshwater gastropods in Vietnam (Caenogastropoda: Cerithioidea)". ''
Molluscan Research Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is est ...
'' 29(3): 121-146
abstract
* '' Brotia angulifera'' Brot, 1872 - incertae sedis * '' Brotia armata'' (Brandt, 1874) * '' Brotia assamensis'' (Nevill, 1885) - incertae sedis * '' Brotia beaumetzi'' (Brot, 1887) - incertae sedis * '' Brotia binodosa'' (Blanford, 1903) * † '' Brotia bittneri'' (Oppenheim, 1895) * '' Brotia boenana'' (Brot, 1881) * '' Brotia borneensis'' (Schepman, 1896) - incertae sedis * † '' Brotia castellaunensis'' (Boussac, 1911) * '' Brotia citrina'' (Brot, 1868) * '' Brotia clavaeformis'' (Brot, 1874) * ''
Brotia costula ''Brotia costula'' is a species of gastropod belonging to the family Pachychilidae Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilida ...
'' (Rafinesque, 1833) * '' Brotia cylindrus'' (Brot, 1886) - incertae sedis * ''
Brotia dautzenbergiana ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Morlet, 1884) * '' Brotia episcopalis'' (H. Lea & I. Lea, 1851) * '' Brotia godwini'' (Brot, 1875) * '' Brotia henriettae'' (Griffith & Pidgeon, 1834) * '' Brotia herculea'' (Gould, 1846) * ''
Brotia hoabinhensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler, Holford, Do & Ho, 2009 * '' Brotia huberi'' Thach, 2021 * '' Brotia indragirica'' (von Martens, 1900) * '' Brotia insolita'' (Brot, 1868) * '' Brotia iravadica'' (Blanford, 1869) * '' Brotia jullieni'' (Deshayes, 1874) * '' Brotia kelantanesis'' (Preston, 1907) * '' Brotia laodelectata'' Köhler, 2008 * † '' Brotia lhazeensis'' Yü, 1982 * † '' Brotia lombersensis'' (Noulet, 1867) * † '' Brotia majevitzae'' (Oppenheim, 1901) * '' Brotia manningi'' Brandt, 1968 * '' Brotia mariae'' Köhler, 2008 * '' Brotia microsculpta'' Brandt, 1968 * ''
Brotia pageli ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea: ...
'' (Thiele, 1908) * ''
Brotia pagodula ''Brotia pagodula'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Taxonomy ''Brotia pagodula'' is the type species of the genus ''Brotia''.Köhler F., Holford, Do V. T. & Ho T. ...
'' (Gould, 1847) - type species * † '' Brotia palaeocostula'' Gurung, Takayasu & Matsuoka, 1997 * '' Brotia paludiformis'' (Solem, 1966) * '' Brotia peninsularis'' (Brandt, 1974) * '' Brotia persculpta'' (Ehrmann, 1922) * '' Brotia praetermissa'' Köhler & Glaubrecht, 2002 * '' Brotia pseudoasperata'' Brandt, 1968 * '' Brotia pseudosulcospira'' (Brandt, 1968) * '' Brotia siamensis'' (Brot, 1886) * '' Brotia solemiana'' (Brandt, 1968) * '' Brotia sooloensis'' (Reeve, 1859) - incertae sedis * '' Brotia spinata'' (Godwin-Austen, 1872) - incertae sedis * '' Brotia subgloriosa'' (Brandt, 1968) * '' Brotia sumatrensis'' (Brot, 1875) * '' Brotia torquata'' (von dem Busch, 1842) * '' Brotia verbecki'' (Brot, 1886) * '' Brotia wykoffi'' (Brandt, 1974) * '' Brotia yunnanensis'' Köhler, Du & Yang, 2010Köhler F., Du L.-N. & Yang J.-X. (2010). "A new species of ''Brotia'' from Yunnan, China (Caenogastropoda, Pachychilidae)". '' Zoosystematics and Evolution'' 86(2): 295-300. . * '' Brotia zonata'' (Benson, 1836) - incertae sedis Synonyms: * ''Brotia asperata'' (Lamarck, 1822): synonym of '' Jagora asperata'' (Lamarck, 1822) * ''Brotia baccata'' (Gould, 1847): synonym of '' Brotia henriettae'' (Gray, 1834) * ''Brotia delavayana'' Heude, 1888 : synonym of '' Sulcospira delavayana'' (Heude, 1888) * † ''Brotia escheri aquitanica'' (Noulet, 1846): synonym of † '' Tinnyea aquitanica'' (Noulet, 1846) * † ''Brotia escheri auingeri'' (Handmann, 1882): synonym of † '' Tinnyea lauraea'' (Matheron, 1842) * ''Brotia hainanensis'' (Brot, 1872): synonym of ''
Sulcospira hainanensis ''Sulcospira'' is a genus of freshwater snails which have an operculum, aquatic gastropod mollusks in the family Pachychilidae. Distribution The distribution of this genus includes: Malaysia, China, Thailand, Indonesia, and Vietnam. Speci ...
'' (Brot, 1874) * ''Brotia hamonvillei'' : synonym of '' Sulcospira tonkiniana'' (Morelet, 1886) * ''Brotia infracostata'' (Mousson, 1848): synonym of '' Sulcospira infracostata'' (Mousson, 1848) * ''Brotia proteus'' (Bavay & Dautzenberg, 1908): synonym of ''
Sulcospira dakrongensis ''Sulcospira'' is a genus of freshwater snails which have an operculum, aquatic gastropod mollusks in the family Pachychilidae. Distribution The distribution of this genus includes: Malaysia, China, Thailand, Indonesia, and Vietnam. Speci ...
'' Köhler, Holford, Do & Ho, 2009 * ''Brotia swinhoei'' (H. Adams, 1870): synonym of '' Sulcospira swinhoei'' (H. Adams, 1870) * ''Brotia testudinaria'' (von dem Busch, 1842):Köhler F. & Glaubrecht M. (2001). "Toward a systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' H. – Adams, 1866 (Cerithioidea: Pachychilidae): an account of species from around the South China Sea". '' Journal of Molluscan Studies'' 67(3): 281-318. synonym of '' Sulcospira testudinaria'' (von dem Busch, 1842) * ''Brotia tourannensis'': synonym of '' Sulcospira tourannensis'' (Souleyet, 1852) * ''Brotia variabilis'' (Benson, 1836): synonym of ''
Brotia costula ''Brotia costula'' is a species of gastropod belonging to the family Pachychilidae Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilida ...
'' (Rafinesque, 1833) Variability of shells of ''Brotia'' (images in the first row are in the same ratio): File:Brotia armata shell.png, '' Brotia armata'' File:Brotia binodosa shell.png, '' Brotia binodosa'' File:Brotia herculea shell.png, '' Brotia herculea''


References


External links

* Köhler F. & Deein G. (2010). "Hybridisation as potential source of incongruence in the morphological and mitochondrial diversity of a Thai freshwater gastropod (Pachychilidae, ''Brotia'' H. Adarns, 1866)". '' Zoosystematics and Evolution'' 86(2): 301-314. . {{Taxonbar, from=Q990852 Gastropod genera
Brotia ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
Freshwater snails