''Brodiechelys'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of terrestrial

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...

belonging to the family

Xinjiangchelyidae Xinjiangchelyidae is an extinct family of turtles known from the Lower Jurassic to the Middle Cretaceous of Asia and western Europe. They have generally been interpreted as either being basal cryptodires or placed outside of crown Testudines. G ...

. Remains of ''Brodiechelys'' dated back to the

Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma. Geology Pro ...

period (

Berriasian In the geological timescale, the Berriasian is an age/stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 145.0 ± 4.0 Ma and 139.8 ± 3.0 Ma (million years ago) ...

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...

stages), and have been found in the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

and

Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

, ''B. brodiei'', was thought to only come from the Wealden of the

, however, specimens of ten shells were found in the locality of

Brighstone Bay Brighstone Bay is a bay on the south west coast of the Isle of Wight, England. It lies to the south and west of the village of Brighstone from which it takes its name. It faces south west towards the English Channel, its shoreline is 7 km ...

in the Wessex Sub-basin of the

Isle of Wight The Isle of Wight ( ) is a county in the English Channel, off the coast of Hampshire, from which it is separated by the Solent. It is the largest and second-most populous island of England. Referred to as 'The Island' by residents, the Isle of ...

, UK. Fragmentary specimens of shell corresponding to an indeterminate species were also found outside Wealden; these specimens were unearthed in the Weald Sub-basin of

Sussex Sussex (), from the Old English (), is a historic county in South East England that was formerly an independent medieval Anglo-Saxon kingdom. It is bounded to the west by Hampshire, north by Surrey, northeast by Kent, south by the English ...

, UK. The discovery of the species ''B. royoi'' in the

Maestrazgo Basin The Maestrazgo () or Maestrat () is a natural and historical mountainous region, located at the eastern end of the Sistema Ibérico mountain range, in Spain. It encompasses the north of the Autonomous Community of Valencia, in Castellón provinc ...

of Morella,

had made

paleontologists Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

realize that some

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

turtle taxa had a wide

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

an distribution during the Early Cretaceous, and at the same time being present in both the UK and the

Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...

. ''B. royoi'' was assigned to ''Brodiechelys'' due to the identified similarities it had with ''B. brodiei'', however, ''B. royoi'' possessed several

autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...

, and therefore its specimens could not have belonged to ''B. brodiei''.

References

Testudinata Prehistoric turtle genera Early Cretaceous turtles Early Cretaceous reptiles of Europe Cretaceous England Fossils of England Cretaceous Spain Fossils of Spain Fossil taxa described in 1928 Taxa named by Franz Nopcsa von Felső-Szilvás {{Paleo-turtle-stub