Alex Rasnitsyn
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Alexandr Pavlovich Rasnitsyn (Russian: Александр Павлович Расницын) is a Russian entomologist, expert in palaeoentomology, and Honored Scientist of the Russian Federation (2001).RASNITSYN Александр Павлович (www.paleo.ru)
/ref> His scientific interests are centered on the palaeontology, phylogeny, and taxonomy of
hymenopteran Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
insects and insects in general. He has also studied broader biological problems such as evolutionary theory, the principles of phylogenetics, taxonomy, nomenclature, and palaeoecology. He has published over 300 articles and books in several languages. In August 2008 he was awarded the Distinguished Research Medal of the
International Society of Hymenopterists The International Society of Hymenopterists focuses on the study of the insect order of ''Hymenoptera''. It was founded in 1982. Journal Since 1992, the society publishes the ''Journal of Hymenoptera Research''. The editor-in-chief An editor-in- ...
.


Biography

Alexandr Rasnitsyn was born on 24 September 1936 in Moscow. As a schoolboy Alex was active in the Society of Young Biologists at the
Moscow Zoo The Moscow Zoo or Moskovsky Zoopark (russian: Московский зоопарк) is a zoo founded in 1864 by professor-biologists, K.F. Rulje, S.A. Usov and A.P. Bogdanov, from the Moscow State University. In 1919, the zoo was nationalized. In ...
. In 1955 he became a student at the Biological Faculty of the
Moscow State University M. V. Lomonosov Moscow State University (MSU; russian: Московский государственный университет имени М. В. Ломоносова) is a public research university in Moscow, Russia and the most prestigious ...
and in 1960 he graduated with honors from the Department of Entomology. His Master thesis was "Hibernation in the ichneumon-fly subfamily Ichneumoninae". The same year Rasnitsyn joined the Laboratory of Arthropods at the Paleontological institute, Academy of Sciences of USSR. In 1967 he received his Ph.D. in biology from the Paleontological Institute with the thesis "The Mesozoic Hymenoptera Symphyta and the early evolution of Xyelidae". After defending in 1978 his Dr. hab. (doktor nauk) thesis "The origin and evolution of Hymenoptera" Rasnitsyn became Head of the Laboratory of Arthropods. In 1991 he received the title of a Biology Professor. In 1996 he resigned from heading the Laboratory and continued there as a principal research worker, but after the new leader, Vladimir Zherikhin, died in 2001 Rasnitsyn again became the acting Head of the Laboratory (2002—present). Between 2001 and 2005 Rasnitsyn served as President of the International Palaeoentomological Society. Since 2007 he is serving on the Council of the
Russian Entomological Society The Russian Entomological Society is a Russian scientific society devoted to entomology. The Society was founded in 1859 in St. Petersburg by Karl Ernst von Baer, Johann Friedrich von Brandt who was then the director of the Zoological Museum of th ...
. During more than 20 field seasons between 1956 and 2009 Rasnitsyn conducted field work in various regions of Russia and the former
USSR The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, it was nominally a federal union of fifteen nationa ...
, including
Fergana Valley The Fergana Valley (; ; ) in Central Asia lies mainly in eastern Uzbekistan, but also extends into southern Kyrgyzstan and northern Tajikistan. Divided into three republics of the former Soviet Union, the valley is ethnically diverse and in the ...
,
Issyk Kul Issyk-Kul (also Ysyk-Köl, ky, Ысык-Көл, lit=warm lake, translit=Ysyk-Köl, , zh, 伊塞克湖) is an endorheic lake (i.e., without outflow) in the Northern Tian Shan mountains in Eastern Kyrgyzstan. It is the seventh-deepest lake in th ...
,
Central Asia Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes t ...
,
Transbaikalia Transbaikal, Trans-Baikal, Transbaikalia ( rus, Забайка́лье, r=Zabaykalye, p=zəbɐjˈkalʲjɪ), or Dauria (, ''Dauriya'') is a mountainous region to the east of or "beyond" (trans-) Lake Baikal in Far Eastern Russia. The steppe and ...
, Taimyr,
Okhotsk Okhotsk ( rus, Охотск, p=ɐˈxotsk) is an types of inhabited localities in Russia, urban locality (a urban-type settlement, work settlement) and the administrative center of Okhotsky District of Khabarovsk Krai, Russia, located at the mou ...
,
Sikhote-Alin The Sikhote-Alin (russian: Сихотэ́-Али́нь, , , ) is a mountain range in Primorsky and Khabarovsk Krais, Russia, extending about to the northeast of the Russian Pacific seaport of Vladivostok. The highest summits are Tordoki Yani at ...
, and other parts of
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
,
Far East The ''Far East'' was a European term to refer to the geographical regions that includes East and Southeast Asia as well as the Russian Far East to a lesser extent. South Asia is sometimes also included for economic and cultural reasons. The ter ...
, and
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
.


Family

Rasnitsyn has one son, Dmitri, who is from a previous marriage. Dmitri married a Masha Kreinin and Rasnitsyn now has three grandchildren; Alexandra, Shelly, and Jonathan Rasnitsyn, who all live in the US.


Research

One of the world's leading paleoentomologists, Rasnitsyn has described ca. 250Nomenclator Zoologicus
/ref> new genera and over 800EDNA Database
/ref> new species of fossil insects from various orders. He is one of the foremost authorities on the paleontology and systematics of
Hymenoptera Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Par ...
whose ideas have formed the foundation of the modern classification of that insect order. Instead of the traditional division into
Symphyta Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay ...
and
Apocrita Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole) formed ...
, he divided the order into “sawflies” (Siricina) and “stinging and parasitic wasps’’ (Vespina), the latter suborder including the parasitic Orussoidea, traditionally placed in the Symphyta. Rasnitsyn suggested his own hypothesis on the origin of insect flight. According to him, the wings first evolved, as a means to control gliding, in relatively large insects that had turned to feeding on generative organs of arboreal plants. Rasnitsyn is one of the most consistent opponents of
cladism Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived chara ...
. He develops an alternative approach to biological systematics, called “phyletics”, which differs from
phenetics In biology, phenetics ( el, phainein – to appear) , also known as taximetrics, is an attempt to classify organisms based on overall similarity, usually in morphology or other observable traits, regardless of their phylogeny or evolutionary rel ...
in taking into account genealogy in addition to similarities and hiatuses. He also contributed significantly to the
epigenetic theory of evolution In biology, epigenetics is the study of stable phenotypic changes (known as ''marks'') that do not involve alterations in the DNA sequence. The Greek prefix '' epi-'' ( "over, outside of, around") in ''epigenetics'' implies features that are "o ...
and, in particular, has put forward the concept of “adaptive compromise” and the notion of macroevolution being irreducible to microevolutionary processes alone. Rasnitsyn has also made a significant contribution to paleoecology and in collaboration with Vladimir Zherikhin developed the theory of ecological crises.


New taxa described by Alexandr Rasnitsyn

* Suborder †'' Eolepidopterigina'' Rasn., 1983 – a suborder of
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
with the family ''
Eolepidopterigidae Eolepidopterigoidea is an extinct superfamily of moths, containing the single family Eolepidopterigidae, although the genus ''Undopterix'' is sometimes placed in a separate family Undopterigidae. The type-genus of the family is ''Eolepidopterix'' ...
'' Rasn., 1983Rasnitsyn AP (1983) First record of a moth in the Jurassic. Doklady Akademii Nauk SSSR 269(2): 467–471 n Russian, with English translation in Doklady Biological Sciences [Transactions of the Academy of Sciences of the USSR, Biological Sciences Section 1983]. * Suborder †''Aneuretopsychina'' Rasnitsyn et Kozlov, 1990 – a suborder of Mecoptera with the family ''Aneuretopsychidae'' Rasnitsyn et Kozlov, 1990 * Superfamily †''Karatavitoidea'' Rasn., 1963 – a Mesozoic superfamily of the infraorder '' Orussomorpha'' Newman, 1834 with the family ''
Karatavitidae Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now consid ...
'' Rasn., 1963 * Superfamily †'' Bethylonymoidea'' Rasn., 1975 – a Mesozoic superfamily of Hymenoptera Apocrita with the family '' Bethylonymidae'' Rasn., 1975. The ancestor group of Aculeata.Rasnitsyn AP (1975) Hymenoptera Apocrita of the Mesozoic. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 147: 1–134 n Russian * Family †'' Parapamphiliidae'' Rasn., 1968 – later was included into ''
Sepulcidae Sepulcidae is an extinct family of stem sawflies in the order Hymenoptera. The family is known primarily from late Mesozoic fossils found in 1968 in Transbaikalia. The insects were distant relatives of modern sawflies and are part of the living ...
'' Rasn., 1968 as the subfamily '' Parapamphiliinae'' Rasn., 1968Rasnitsyn AP (1968) New Mesozoic sawflies (Hymenoptera, Symphyta). In: Rohdendorf BB (Ed) Jurassic Insects of Karatau. Nauka Press, Moscow, 190–236 otal pages 252 pp., +25 pls. n Russian * Family †'' Xyelydidae'' Rasn., 1968 – a Mesozoic family of Hymenoptera Symphyta from the superfamily ''
Pamphilioidea The Pamphilioidea are a small superfamily within the Symphyta (the sawflies), containing some 250 living species restricted to the temperate regions of Eurasia and North America. These hymenopterans share the distinctive feature of a very large ...
'' * Family †'' Gigasiricidae'' Rasn., 1968 – a Jurassic family of Hymenoptera Symphyta from the superfamily ''
Siricoidea The superfamily Siricoidea is an archaic group of the order Hymenoptera, consisting of six families (four extinct) of xylophagous sawflies. The group is well represented in early Tertiary and Mesozoic times, but a number of living taxa remain, inc ...
'' * Family †'' Xyelotomidae'' Rasn., 1968 – a Mesozoic family of Hymenoptera Symphyta from the superfamily ''
Tenthredinoidea The Tenthredinoidea are the dominant superfamily of sawflies within the Symphyta, containing some 8,400 species worldwide, primarily in the family Tenthredinidae. All known larvae are phytophagous, and a number are considered pests. The included ...
'' * Family †'' Pararchexyelydae'' Rasn., 1968 – a Mesozoic family of Hymenoptera Symphyta * Family †'' Praeaulacidae'' Rasn., 1972 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' * Family †''
Maimetshidae Maimetshidae is an extinct family of wasps, known from the Cretaceous period.M. S. Engel. 2016A new genus and species of maimetshid wasps in Lebanese Early Cretaceous amber (Hymenoptera: Maimetshidae).''Novitates Paleoentomologicae'' 18:1-14 While ...
'' Rasn., 1975 – a Cretaceous family of Hymenoptera Apocrita from the superfamily ''
Ceraphronoidea The Ceraphronoidea are a small hymenopteran superfamily that includes only two families, and a total of some 800 species, though a great many species are still undescribed. It is a poorly known group as a whole, and most are believed to be parasi ...
'' * Family †'' Cretevaniidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' * Family †'' Kotujellidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' (later included into ''
Gasteruptiidae The Gasteruptiidae are one of the more distinctive families among the apocritan wasps, with surprisingly little variation in appearance for a group that contains around 500 species in two subfamilies ( Gasteruptiinae and Hyptiogastrinae) and wi ...
'' Ashmead, 1900) * Family †'' Anomopterellidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' * Family †'' Baissidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' (later was included into ''
Aulacidae The Aulacidae are a small, cosmopolitan family of wasps, with two extant genera containing some 200 known species. They are primarily endoparasitoids of wood wasps (Xiphydriidae) and xylophagous beetles (Cerambycidae and Buprestidae). They are c ...
'' Schuckard, 1841) * Family †'' Ichneumonomimidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita. Systematic position uncertain, probably a relative of ''
Ichneumonoidea The superfamily Ichneumonoidea contains one extinct and three extant families, including the two largest families within Hymenoptera: Ichneumonidae and Braconidae. The group is thought to contain as many as 100,000 species, many of which have not ...
'' * Family †''
Angarosphecidae Angarosphecidae is an extinct family of Mesozoic and early Cenozoic wasps in the superfamily Apoidea. Genera * †'' Angarosphex'' Rasnitsyn 1975 Weald Clay, United Kingdom, Hauterivian/Barremian La Pedrera de Rúbies Formation, Spain, Barremi ...
'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita from the superfamily ''
Scolioidea The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
'' * Family †'' Falsiformicidae'' Rasn., 1975 – a Cretaceous family of Hymenoptera Apocrita from the superfamily ''
Scolioidea The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
'' * Family †'' Baissodidae'' Rasn., 1975 – a Mesozoic family of Hymenoptera Apocrita * Family †'' Evenkiidae'' Rasn., 1977 – a Carboniferous family of the order '' Protortoptera''Rasnitsyn AP (1977) New Paleozoic and Mesozoic insects. Paleontologicheskiy Zhurnal 1977(1): 64–77 n Russian, with English translation in Paleontological Journal, 1978, 11: 60–72 * Family †'' Permonkidae'' Rasn., 1977 – a family of ''
Miomoptera Miomoptera is an extinct order of insects. Although it is thought to be a common ancestor of all holometabolous insects, because no smooth transition between Miomoptera and other holometabolous insect orders is known, it is considered to be in a ...
'' * Family †'' Palaeomantiscidae'' Rasn., 1977 – a family of ''
Miomoptera Miomoptera is an extinct order of insects. Although it is thought to be a common ancestor of all holometabolous insects, because no smooth transition between Miomoptera and other holometabolous insect orders is known, it is considered to be in a ...
'' * Family †'' Karataidae'' Rasn., 1977 – a Mesozoic family of Hymenoptera Apocrita * Family †'' Electrotomidae'' Rasn., 1977 – a family of ''
Tenthredinoidea The Tenthredinoidea are the dominant superfamily of sawflies within the Symphyta, containing some 8,400 species worldwide, primarily in the family Tenthredinidae. All known larvae are phytophagous, and a number are considered pests. The included ...
'' from Baltic amberRasnitsyn AP (1977) A new family of sawflies (Hymenoptera, Tenthredinoidea, Electrotomidae) from the Baltic amber. Zoologicheskiy Zhurnal 56(9): 1304–1308 n Russian, with English summary * Family †''
Praeichneumonidae ''Praeichneumon'' is an extinct genus of ichneumon wasps from the Lower Cretaceous of Mongolia and the Russian region of Transbaikalia. It was originally described by Alexandr Pavlovich Rasnitsyn Alexandr Pavlovich Rasnitsyn (Russian: Але ...
'' Rasn., 1983 – a Lower Cretaceous family of the superfamily ''
Ichneumonoidea The superfamily Ichneumonoidea contains one extinct and three extant families, including the two largest families within Hymenoptera: Ichneumonidae and Braconidae. The group is thought to contain as many as 100,000 species, many of which have not ...
'' * Family †''
Strashilidae Strashilidae is an extinct family of Jurassic flies from Siberia and China. They were originally believed to represent a distinct order called Nakridletia, but subsequent research determined that they were nematoceran flies related to the extant ...
'' Rasnitsyn, 1992 – a family of Mecopteroidea. Hypothetical ancestors of
Anoplura Sucking lice (Anoplura, formerly known as Siphunculata) have around 500 species and represent the smaller of the two traditional superfamily (taxonomy), superfamilies of lice. As opposed to the paraphyletic chewing lice, which are now divided amo ...
* Family †'' Saurodectidae'' Rasnitsyn et Zherikhin, 2000 – a Mesozoic family of
Mallophaga The Mallophaga are a possibly paraphyletic section of lice Louse ( : lice) is the common name for any member of the clade Phthiraptera, which contains nearly 5,000 species of wingless parasitic insects. Phthiraptera has variously been reco ...
* Family †'' Andreneliidae'' Rasnitsyn et Martinez-Delclos, 2000 – a family of Hymenoptera Apocrita of the superfamily ''
Evanioidea The Evanioidea are a small hymenopteran superfamily that includes three families, two of which (Aulacidae and Gasteruptiidae) are much more closely related to one another than they are to the remaining family, Evaniidae. The rich fossil record, h ...
'' * Family †'' Tshekarcephalidae'' Novokshonov et Rasnitsyn, 2000 – a Paleozoic family of uncertain systematic position * Family †'' Daohugoidae'' Rasn. et Zhang Haichun, 2004 – a family of Hymenoptera Symphyta of the superfamily ''
Siricoidea The superfamily Siricoidea is an archaic group of the order Hymenoptera, consisting of six families (four extinct) of xylophagous sawflies. The group is well represented in early Tertiary and Mesozoic times, but a number of living taxa remain, inc ...
'' * Family †''
Khutelchalcididae ''Khutelchalcis'' is an extinct genus of chalcid wasp, it is the only member of the family Khutelchalcididae. It is only known from a single species, (''Khutelchalcis gobiensis'') described from the Early Cretaceous Tsagaantsav Formation The T ...
'' Rasnitsyn, Basibuyuk et Quicke, 2004 – a family of
Chalcidoidea Chalcid wasps (, , for their metallic colour) are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, me ...
* Family †''
Radiophronidae Radiophronidae is an extinct family of wasps known from two genera found in Cretaceous (Albian) aged amber from Spain. While originally classified in Ceraphronoidea, they were later considered to probably be members of Chrysidoidea The super ...
'' Ortega-Blanco, Rasnitsyn et Delclos, 2010 – a Mesozoic family of ''
Ceraphronoidea The Ceraphronoidea are a small hymenopteran superfamily that includes only two families, and a total of some 800 species, though a great many species are still undescribed. It is a poorly known group as a whole, and most are believed to be parasi ...
'' * Subfamily †'' Archexyelinae'' Rsan., 1964 – a subfamily of ''
Xyelidae The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 Extant taxon, extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprisi ...
'' Newman, 1834 * Subfamily †'' Dolichostigmatinae'' Rasn., 1968 – a subfamily of ''
Anaxyelidae Anaxyelidae is a family of incense cedar wood wasps in the order Hymenoptera. It contains only one living genus, '' Syntexis'', which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jura ...
'' Martynov., 1925 * Subfamily †'' Karatavitinae'' Rasn., 1968 –a subfamily of ''
Karatavitidae Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now consid ...
'' Rasn., 1963 * Subfamily †'' Sepulcinae'' Rasn., 1968 – was described as a subfamily of ''
Karatavitidae Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now consid ...
'' Rasn., 1963, later was raised to the family rank, ''
Sepulcidae Sepulcidae is an extinct family of stem sawflies in the order Hymenoptera. The family is known primarily from late Mesozoic fossils found in 1968 in Transbaikalia. The insects were distant relatives of modern sawflies and are part of the living ...
'' Rasn., 1968 * Subfamily †'' Auliscinae'' Rasn., 1968 – a subfamily of ''
Karatavitidae Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now consid ...
'' Rasn., 1963 * Subfamily †'' Praesiricinae'' Rasn., 1968 – was described as a subfamily of ''
Karatavitidae Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now consid ...
'' Rasn., 1963, later was raised to the family rank, '' Praesiricidae'' Rasn. 1968 * Subfamily †'' Madygellinae'' Rasn., 1969 – a subfamily of ''
Xyelidae The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 Extant taxon, extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprisi ...
'' Newman, 1834Rasnitsyn AP (1969) The origin and evolution of lower Hymenoptera. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 123: 1–196 n Russian, with English translation by Amerind Co., New Delhi, 1979 * Subfamily †'' Cleistogastrinae'' Rasn., 1975 – a subfamily of ''
Megalyridae Megalyroidea is a small hymenopteran superfamily of wasps that includes a single family, Megalyridae, with eight extant genera (plus around a dozen extinct ones) and 49 described species. Modern megalyrids are found primarily in the southern hem ...
'' Schletterer, 1889 * Subfamily '' Proscoliinae'' Rasn., 1977 – a recent subfamily of
Scoliidae The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
* Subfamily †'' Juralydinae'' Rasn., 1977 – a subfamily of''
Pamphiliidae Pamphiliidae (sometimes incorrectly spelled Pamphilidae) is a small wasp family within Symphyta, containing some 200 species from the temperate regions of North America and Eurasia. The larvae feed on plants (often conifers), using silk to bu ...
'' Cameron, 1890 * Subfamily †'' Mesorussinae'' Rasn., 1977 – a subfamily of ''
Orussidae The Orussidae or the parasitic wood wasps represent a small family of sawflies (" Symphyta"). Currently, about 93 extant and four fossil species are known. They take a key position in phylogenetic analyses of Hymenoptera, because they form the s ...
'' Mewman, 1834 * Subfamily †'' Cretogonalinae'' Rasn., 1977 – a subfamily of ''
Trigonalidae Trigonalidae (the spelling Trigonalyidae is incorrect under Article 29.5 of the ICZN,ICZN Code
29.5. Maintena ...
'' Cresson, 1867 * Subfamily †'' Manlayinae'' Rasn., 1986 – a subfamily of ''
Aulacidae The Aulacidae are a small, cosmopolitan family of wasps, with two extant genera containing some 200 known species. They are primarily endoparasitoids of wood wasps (Xiphydriidae) and xylophagous beetles (Cerambycidae and Buprestidae). They are c ...
'' Schuckard, 1841 * Subfamily †'' Ghilarellinae'' Rasn., 1986 – a subfamily of ''
Sepulcidae Sepulcidae is an extinct family of stem sawflies in the order Hymenoptera. The family is known primarily from late Mesozoic fossils found in 1968 in Transbaikalia. The insects were distant relatives of modern sawflies and are part of the living ...
'' Rasn., 1968Rasnitsyn AP (1988) Sepulcidae and origin of Cephidae (Hymenoptera: Cephoidea). Transactions of the All-Union Entomological Society 70: 480–497 n Russian obias VI, editor (1988) Taxonomy of Insects and Mites. Nauka Press, Moscow * Subfamily †'' Trematothoracinae'' Rasn., 1986 – a subfamily of ''
Sepulcidae Sepulcidae is an extinct family of stem sawflies in the order Hymenoptera. The family is known primarily from late Mesozoic fossils found in 1968 in Transbaikalia. The insects were distant relatives of modern sawflies and are part of the living ...
'' Rasn., 1968 * Subfamily †'' Priorvespinae'' Carpenter et Rasnitsyn, 1990 – a subfamily of ''
Vespidae The Vespidae are a large (nearly 5000 species), diverse, cosmopolitan family of wasps, including nearly all the known eusocial wasps (such as ''Polistes fuscatus'', ''Vespa orientalis'', and ''Vespula germanica'') and many solitary wasps. Each ...
'' Latrielle, 1802 * Subfamily †'' Archaeoscoliinae'' Rasnitsyn, 1993 – a subfamily of
Scoliidae The Scoliidae, the scoliid wasps, are a family of about 560 species found worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than females, wi ...
* Subfamily †'' Karataoserphinae'' Rasnitsyn, 1994 – a subfamily of '' Mesoserphidae'' Kozlov, 1970 * Subfamily †'' Iscopininae'' Rasnitsyn, 1980 – a subfamily of ''
Pelecinidae Pelecinidae is a family of parasitic wasps in the superfamily Proctotrupoidea. It contains only one living genus, '' Pelecinus'', with three species known from the Americas. The earliest fossil species are known from the Jurassic, and the group ...
'' Haliday, 1840. * Tribe †'' Angaridyelini'' Rasn., 1966 – a tribe of the subfamily '' Macroxyelinae'' Ashmead, 1898 of the family ''
Xyelidae The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 Extant taxon, extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprisi ...
'' * Tribe †'' Cretodinapsini'' Rasn., 1977 – a tribe of the subfamily '' Megalyrinae'' of the family ''
Megalyridae Megalyroidea is a small hymenopteran superfamily of wasps that includes a single family, Megalyridae, with eight extant genera (plus around a dozen extinct ones) and 49 described species. Modern megalyrids are found primarily in the southern hem ...
'' * Tribe †'' Gigantoxyelini'' Rasn., 1969 – a tribe of the subfamily '' Macroxyelinae'' Ashmead, 1898 of the family ''
Xyelidae The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 Extant taxon, extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprisi ...
'' Newmann, 1834 A.P. Rasnitsyn has also described ca. 250 new genera and over 800 new species of arthropods, mainly fossil.


Animal names in honor of Alexandr Rasnitsyn

Over 50 species of animals have been named in honor of Rasnitsyn, as well as some taxa of higher rank: * '' Rasnicynipidae'' Kovalev, 1996 (a replacement name for '' Rasnitsyniidae'' Kovalev, 1994)— a fossil family of
Cynipoidea The Cynipoidea are a moderate-sized hymenopteran superfamily that presently includes five modern families and three extinct families, though others have been recognized in the past. The most familiar members of the group are phytophagous, especia ...
with the genus '' Rasnicynips'' Kovalev, 1996 (a replacement name for '' Rasnitsynia'' Kovalev 1994) * '' Alexarasniidae'' Gorochov, 2011 — a fossil family of ''
Polyneoptera The cohort Polyneoptera is a proposed taxonomic ranking for the Orthoptera (grasshoppers, crickets, etc.) and all other Neopteran insects believed to be more closely related to Orthoptera than to any other insect orders. These winged insects, no ...
'' with the genus '' Alexarasnia'' Gorochov, 2011 * '' Alexrasnitsyniidae'' Prokop & Nel, 2011 — a family of the Paleozoic order ''
Diaphanopterodea The Diaphanopterodea or Paramegasecoptera are an extinct order of moderate to large-sized Palaeozoic insects. They are first known from the Middle Carboniferous (late Serpukhovian or early Bashkirian in age), and include some of the earliest know ...
'' with the genus '' Alexrasnitsynia'' Prokop & Nel, 2011 * ''
Plumalexiidae ''Plumalexius'' is a genus of wasps in the extinct monotypic family Plumalexiidae, containing two species: the type species ''Plumalexius rasnitsyni'', known from the Late Cretaceous White Oaks Pit in Sayreville, New Jersey, and ''Plumalexius ohm ...
'' Brothers, 2011 — a fossil family of Hymenoptera with the species '' Plumalexius rasnitsyni'' Brothers, 2011 * '' Rasnitsynaphididae'' Homan & Wegierek, 2011 — a fossil family of Aphidoidea with the species '' Rasnitsynaphis'' Homan & Wegierek, 2011 * '' Rasnitsynitini'' Kasparyan, 1994 — a tribe of the fossil subfamily '' Townesitinae'' of the family ''
Ichneumonidae The Ichneumonidae, also known as the ichneumon wasps, Darwin wasps, or ichneumonids, are a family (biology), family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 2 ...
'' with the genus '' Rasnitsynites'' Kasparyan, 1994 * '' Rasnitsynella'' Krivolutzkii, 1976 — a fossil genus of Acarina * '' Rasnitsynia'' Pagliano et Scaramozzino 1989 (a replacement name for '' Oligoneuroides'' Zhang 1985) — a fossil genus of ''
Braconidae The Braconidae are a family of parasitoid wasps. After the closely related Ichneumonidae, braconids make up the second-largest family in the order Hymenoptera, with about 17,000 recognized species and many thousands more undescribed. One analysis ...
'' * '' Rasnitsynitilla'' Lelej, 2006 — a genus of ''
Mutillidae The Mutillidae are a family of more than 7,000 species of wasps whose wingless females resemble large, hairy ants. Their common name velvet ant refers to their dense pile of hair, which most often is bright scarlet or orange, but may also be bla ...
'' * '' Alicodoxa rasnitsyni'' Emeljanov et Shcherbakov, 2011 — fossil genus and species of Fulgoroidea of the family ''
Dictyopharidae Dictyopharidae is a family of planthoppers, related to the Fulgoridae. The family comprises nearly 760 species in more than 150 genera which are grouped into two subfamilies, Dictyopharinae and Orgeriinae. Description Like all other fulgoroi ...
'' * '' Palerasnitsynus'' Wichard, Ross et Ross, 2011 — a fossil genus of
Trichoptera The caddisflies, or order Trichoptera, are a group of insects with aquatic larvae and terrestrial adults. There are approximately 14,500 described species, most of which can be divided into the suborders Integripalpia and Annulipalpia on the b ...
of the family ''
Psychomyiidae The Psychomyiidae are a family of tube-making caddisflies. Members of this family are typically very similar to polycentropodids, most of them can be differentiated by the spur formula is 2-4-4, thyridial cell short, absence of the forewing for ...
'' * '' Rasnitsynala'' Zessin, Brauckmann et Groening, 2011 — a genus of Odonata from the family '' Erasipteridae''


Publications

A.P. Rasnitsyn is an author of more than 300 books and articles, including 17 monographs.


Major works

* Rasnitsyn AP (1969) Origin and evolution of lower Hymenoptera. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 123: 1–196 n Russian, with English translation by Amerind Co., New Delhi, 1979 * Rasnitsyn AP (1975) Hymenoptera Apocrita of the Mesozoic. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 147: 1–134 n Russian * Rasnitsyn AP (1980) Origin and evolution of Hymenoptera. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 174: 1–192 n Russian * Rohdendorf BB, Rasnitsyn AP, editors (1980) Historical development of the class Insecta. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 175: 1–269, +8 pls. n Russian * Rasnitsyn AP, Quicke DLJ, editors (2002) History of Insects. Kluwer Academic Publishers, Dordrecht, xii+517 pp. . * Rasnitsyn AP (2005) Selected Works on Evolutionary Biology. KMK Scientific Press, Moscow, Russia, iv+347 pp n Russian ollection of earlier papers, except for: “Dynamics of taxonomic diversity: An afterword of 2004”, pp. 247–248 * Zherikhin VV, Ponomarenko AG, Rasnitsyn AP (2008) Introduction to Palaeoentomology. KMK Scientific Press, Moscow, 371 pp. n Russian


References


External links


Personal web pagePalaeoentomology in RussiaInternational Palaeoentomological Society
{{DEFAULTSORT:Rasnitsyn, Alex Living people Russian paleontologists Russian entomologists Hymenopterists 1936 births Soviet paleontologists Soviet entomologists