Archisargidae is an extinct family of

brachycera The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristics i ...

n flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily

Archisargoidea } Archisargoidea is an extinct superfamily of brachyceran flies known from the late Middle Jurassic (Callovian) to early Late Cretaceous (Turonian). Most flies in the superfamily have large eyes and an elongated abdomen, preserved females have a ...

. Most members of the family are known from the Callovian-Oxfordian

Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...

biota of Inner Mongolia, China, and the equivalently aged

Karabastau Formation The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...

of Kazakhstan. The family has been found to be

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

with respect to

Eremochaetidae Eremochaetidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods of Asia. It is part of the extinct superfamily Archisargoidea. The morphology of the ovipositor of the only 3 dimensionally preserved genus '' ...

in a

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...

analysis.

Taxonomy

*†subfamily Archisarginae Rohdendorf 1962 ** †'' Archirhagio'' Rohdendorf 1938 *** †''Archirhagio gracilentus'' Wang et al. 2017

, China,

Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...

*** †''Archirhagio obscurus'' Rohdendorf 1938

, Kazakhstan, Oxfordian *** †''Archirhagio striatus'' Zhang and Zhang 2003J. Zhang and H. Zhang. 2003. Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, northeastern China). ''Paleontological Journal'' 37:409-412 Daohugou, China, Callovian *** †''Archirhagio varius'' Zhang 2014 Daohugou, China, Callovian *** †''Archirhagio zhangi'' Zhang et al. 2009

Tiaojishan Formation The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period (Bathonian- Oxfordian stages). It is known for its exceptionally preserved fossils, including those of ...

, China, Callovian ** †'' Archisargus'' Rohdendorf 1938 *** †''Archisargus maximus'' Mostovski 1997M. B. Mostovski. 1997. On knowledge of fossil flies of the superfamily Archisargoidea (Diptera, Brachycera). ''Paleontological Journal'' 31(1):72-78 Karabastau Formation, Kazakhstan, Oxfordian *** †''Archisargus pulcher'' Rohdendorf 1938 Karabastau Formation, Kazakhstan, Oxfordian *** †''Archisargus spurivenius'' Zhang et al. 2007aK. Y. Zhang, D. Yang, D. Ren and C. K. Shih. 2007. The earliest species of the extinct genus Archisargus from China (Diptera: Brachycera: Archisargidae). ''Annales Zoologici'' 57:827-832 Daohugou, China, Callovian *** †''Archisargus strigatus'' Zhang et al. 2007a Daohugou, China, Callovian ** †'' Calosargus'' Mostovski 1997 *** †''Calosargus (Calosargus)'' Mostovski 1997 **** †''Calosargus (Calosargus) antiquus'' Zhang et al. 2007b Daohugou, China, Callovian **** †''Calosargus (Calosargus) bellus'' Zhang et al. 2007b Daohugou, China, Callovian **** †''Calosargus (Calosargus) daohugouensis'' Zhang et al. 2007b Daohugou, China, Callovian **** †''Calosargus (Calosargus) hani'' Zhang et al. 2007b Daohugou, China, Callovian **** †''Calosargus (Calosargus) niger'' Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian **** †''Calosargus (Calosargus) talbragarensis'' Oberprieler and Yeates 2012 Talbragar Fossil Bed, Australia,

Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...

**** †''Calosargus (Calosargus) tatianae'' Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian **** †''Calosargus (Calosargus) tenuicellulatus'' Zhang et al. 2007b Daohugou, China, Callovian **** †''Calosargus (Calosargus) validus'' Zhang et al. 2007b Daohugou, China, Callovian *** †''Calosargus (Pterosargus)'' Mostovski 1997 **** †''Calosargus (Pterosargus) sinicus'' Zhang 2010 Daohugou, China, Callovian **** †''Calosargus (Pterosargus) thanasymus'' Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian *** †''Calosargus tatianae'' Mostovski 1997 Karabastau Formation, Kazakhstan, Oxfordian ** †'' Flagellisargus'' Zhang 2012 *** †''Flagellisargus robustus'' Zhang 2012 Daohugou, China, Callovian *** †''Flagellisargus sinicus'' Zhang 2012 Daohugou, China, Callovian *** †''Flagellisargus venustus'' Zhang 2012 Daohugou, China, Callovian ** †'' Mesosolva'' Hong 1983 *** †''Mesosolva angustocellulata'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva balyshevae'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva daohugouensis'' Zhang and Zhang 2003 Daohugou, China, Callovian *** †''Mesosolva dolosa'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva hennigi'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva huabeiensis'' Hong 1983

Haifanggou Formation The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ...

, China, Callovian *** †''Mesosolva imperfecta'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva karataviensis'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva longivena'' Mostovski 1996 Shar Teeg, Mongolia, Tithonian *** †''Mesosolva parva'' Hong 1983 Haifanggou Formation, China, Callovian *** †''Mesosolva rohdendorfi'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Mesosolva sinensis'' Zhang et al. 2010 Daohugou, China, Callovian *** †''Mesosolva zhangae'' Zhang 2012 Daohugou, China, Callovian ** †'' Novisargus'' Zhang 2014 *** †''Novisargus rarus'' Zhang 2014 Daohugou, China, Callovian ** †'' Origoasilus'' Zhang et al. 2011K. Y. Zhang, D. Yang, and D. Ren. 2011. A new brachyceran family Origoasilidae Fam. Nov. from the late Mesozoic of China (Insecta: Diptera). ''Acta Geologica Sinica'' 85(5):994-997 *** †''Origoasilus pingquanensis'' Zhang et al. 2011

Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...

, China,

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

** †'' Parvisargus'' Mostovski 1996M. B. Mostovski. 1996. To the knowledge of Archisargoidea (Diptera, Brachycera). Families Eremochaetidae and Archisargidae. ''Russian Entomological Journal'' 5:117-124 *** †''Parvisargus malus'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian *** †''Parvisargus peior'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian ** †'' Sharasargus'' Mostovski 1996 *** †''Sharasargus fortis'' Zhang et al. 2008 Daohugou, China, Callovian *** †''Sharasargus maculus'' Zhang 2014 Daohugou, China, Callovian *** †''Sharasargus oresbius'' Ren 1998D. Ren. 1998. Late Jurassic Brachycera from northeastern China (Insecta: Diptera). ''Acta Zootaxonomica Sinica'' 23:65-82 Yixian Formation, China, Aptian *** †''Sharasargus ruptus'' Mostovski 1996 Shar Teeg, Mongolia, Tithonian *** †''Sharasargus spiniger'' Mostovski 1996 Karabastau Formation, Kazakhstan, Oxfordian ** †'' Sinallomyia'' Zhang 2012 *** †''Sinallomyia ruderalis'' Ren 1998 Yixian Formation, China, Aptian ** †'' Tabanisargus'' Zhang 2014 *** †''Tabanisargus daohugouensis'' Zhang 2014 Daohugou, China, Callovian * †subfamily Uranorhagioninae Zhang et al. 2010 Grimaldi 2016 suggests that this grouping does not belong to the superfamily. ** †'' Daohugosargus'' Zhang 2012 *** †''Daohugosargus eximius'' (Zhang et al. 2008) Daohugou, China, Callovian ** †'' Uranorhagio'' Zhang et al. 2010 *** †''Uranorhagio asymmetricus'' Zhang et al. 2010 Daohugou, China, Callovian *** †''Uranorhagio daohugouensis'' Zhang et al. 2010 Daohugou, China, Callovian *** †''Uranorhagio deviatus'' Zhang et al. 2010 Daohugou, China, Callovian

References

{{Taxonbar, from=Q4120835 † Prehistoric Diptera Prehistoric insect families