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Archisargoidea
} Archisargoidea is an extinct superfamily of brachyceran flies known from the late Middle Jurassic (Callovian) to early Late Cretaceous (Turonian). Most flies in the superfamily have large eyes and an elongated abdomen, preserved females have a sharp, piercing oviscapt used for injecting eggs into host matter. Their relationships with other members of Brachycera is controversial, they are usually considered close relatives of either Stratiomyomorpha or Muscomorpha. Internal relationships between the families are uncertain and the topology of the only cladistic analysis of the family was weakly supported, finding that Archisargidae was paraphyletic with respect to Eremochaetidae and miniaturized Tethepomyiidae. The sharp piercing oviscaps possessed by the group had alternatively been suggested to represent evidence of being parasitoids, injecting their eggs into hosts similar to parasitic wasps, or to inject eggs into plant material like rotting wood or fruit, similar to member ...
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Archisargidae
Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis. Taxonomy *†subfamily Archisarginae Rohdendorf 1962 ** †'' Archirhagio'' Rohdendorf 1938 *** †''Archirhagio gracilentus'' Wang et al. 2017 Daohugou, China, Callovian *** †''Archirhagio obscurus'' Rohdendorf 1938 Karabastau Formation, Kazakhstan, Oxfordian *** †''Archirhagio striatus'' Zhang and Zhang 2003J. Zhang and H. Zhang. 2003. Two new species of archisargids (Insecta: Diptera: Archisargidae) from the Upper Jurassic Daohugou Formation (Inner Mongolia, northeastern China). ''Paleontological Journal'' 37:409-412 Daohugou, China, Callovian *** ...
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Eremochaetidae
Eremochaetidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods of Asia. It is part of the extinct superfamily Archisargoidea. The morphology of the ovipositor of the only 3 dimensionally preserved genus ''Zhenia'' was initially interpreted as evidence of being an endoparasitoid of arthropods, however a subsequent study suggested that the ovipositor was used to deposit its eggs in plant material, similar to members of Tephritoidea. In a phylogenetic analysis, Ermochaetidae was found to be monophyletic, surrounded by a paraphyletic Archisargidae. Taxonomy * †''Alleremonomus'' Ren and Guo 1995 ** †''Alleremonomus xingi'' Ren and Guo 1995 Laiyang Formation, Yixian Formation, China, Aptian * †''Dissup'' Evenhuis 1994 ** †''Dissup clausus'' Zhang et al. 2014 Yixian Formation, China, Aptian ** †''Dissup irae'' Kovalev 1989 Turga Formation, Russia, Aptian * †''Eremochaetomima'' Mostovski 1996M. B. Mostovski. 1996. To the knowledge of ...
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Brachycera
The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristics is: * Antenna size (with eight or fewer flagellomeres) is reduced. * The maxillary palp (an elongated appendage near the mouth) has two segments or fewer. * The back portions of the larval head capsule extend into the prothorax (the anterior part of the thorax, which bears the first pair of legs). * Two distinct parts make up of the larval mandible (lower jaw). * The epandrium and hypandrium of the genitalia are separated in males. * No premandible is present on the lower surface of the labrum (the roof of the mouth). * The configuration of the CuA2 and A1 wing veins is distinct. Brachyceran flies can also be distinguished through behavior. Many of the species are predators or scavengers. Classification The structure of subgroups wit ...
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Tethepomyiidae
Tethepomyiidae is an extinct family of small brachyceran flies known from the Cretaceous period of Laurasia. It is part of the extinct superfamily Archisargoidea. The family is characterised by "very large eyes, reduced mouthparts, a highly reduced antennal flagellum, and greatly reduced venation."D. A. Grimaldi and A. Arillo. 2008The Tethepomyiidae, a new family of enigmatic Cretaceous Diptera ''Alavesia'' 2:259-265 The ovipositor of ''Tethepomyia zigrasi'' has a hypodermic morphology likely used for injecting eggs into hosts. Taxonomy * †'' Tethepomima'' Grimaldi and Arillo 2008 ** †''Tethepomima holomma'' Grimaldi and Arillo 2008 Álava amber (Spanish amber), Escucha Formation, Spain, Albian * †'' Tethepomyia'' Grimaldi and Cumming 1999 ** †''Tethepomyia buruhandi'' Grimaldi and Arillo 2008 Álava amber, Escucha Formation, Spain, Albian ** †''Tethepomyia coxa'' Grimaldi 2016 Burmese amber, Myanmar, Cenomanian ** †''Tethepomyia thauma'' Grimaldi and Cumming 1999 ...
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Muscomorpha
The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important. The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha. Classification The name Cyclorrhapha is used, in various modern classifications, to represent either a subgroup within the infraorder Muscomorpha, or simply a rankless group within the Brachycera. In either ca ...
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Prehistoric Diptera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Tephritoidea
The Tephritoidea are a superfamily of flies. It has over 7,800 species, the majority of them in family Tephritidae. The following families are included: * Ctenostylidae * Eurygnathomyiidae * Lonchaeidae - lance flies * Pallopteridae — flutter flies * Piophilidae — skippers * Platystomatidae — signal flies * Pyrgotidae * Richardiidae * Tephritidae — fruit flies * Ulidiidae (Otitidae) — picture-winged flies The Tachiniscinae, formerly ranked as the family Tachiniscidae, are now included in the Tephritidae. Description Tephritoidea are generally rather hairy flies with setae weakly differentiated. They have the following synapomorphies: male tergum 6 strongly reduced or absent; surstylus or medial surstylus with toothlike prensisetae (in Piophilidae only in one genus); female sterna 4-6 with anterior rodlike apodemes; female tergosternum 7 consisting of two portions, the anterior forming a tubular oviscape and the posterior consisting of two pairs of longitudinal taenia ...
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Parasitoid Wasp
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders. Parasitoid wasp species differ in which host life-stage they attack: eggs, larvae, pupae, or adults. They mainly follow one of two major strategies within parasitism: either they are endoparasitic, developing inside the host, and koinobiont, allowing the host to continue to feed, develop, and moult; or they are ectoparasitic, developing outside the host, and idiobiont, paralysing the host immediately. Some endoparasitic wasps of the superfamily Ichneumonoidea have a mutualistic relationship with polydnaviruses, the vir ...
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Parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable strategy, evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation. Among parasitoids, strategies range from living inside the host (''endoparasitism''), allowing it to continue growing before emerging as an adult, to Paralysis, paralysing the host and living outside it (''ectoparasitism''). Hosts can include other parasitoids, resulting in hyperparasitism; in the case of oak galls, up to five levels of parasitism are possible. Some parasitoids Behavior-altering parasite, influence their host's behaviour in ways that favour the propagation of the parasitoid. Parasitoids are found in a variety of Taxon, taxa across the insect superorder Endopterygota, whose compl ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Turonian
The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by the Cenomanian Stage and underlies the Coniacian Stage. At the beginning of the Turonian an oceanic anoxic event (OAE 2) took place, also referred to as the Cenomanian-Turonian boundary event or the "Bonarelli Event". Stratigraphic definition The Turonian (French: ''Turonien'') was defined by the French paleontologist Alcide d'Orbigny (1802–1857) in 1842. Orbigny named it after the French city of Tours in the region of Touraine (department Indre-et-Loire), which is the original type locality. The base of the Turonian Stage is defined as the place where the ammonite species '' Watinoceras devonense'' first appears in the stratigraphic column. The official reference profile (the GSSP) for the base of the Turonian is located in the Roc ...
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Stratiomyoidea
Stratiomyoidea is a superfamily of flies (order Diptera). The antennae have a primitive structure. A characteristic morphological characteristic of one family, Pantophthalmidae, is the size of the body: this family includes some species that are among the largest Diptera, reaching wingspans of up to 10 cm. Stratiomyoidea larvae live in aquatic or terrestrial habitats and are mostly scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...s that feed on organic material. References Diptera superfamilies Taxa named by Friedrich Georg Hendel {{Stratiomyomorpha-stub ...
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