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Brachycera
The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristics is: * Antenna size (with eight or fewer flagellomeres) is reduced. * The maxillary palp (an elongated appendage near the mouth) has two segments or fewer. * The back portions of the larval head capsule extend into the prothorax (the anterior part of the thorax, which bears the first pair of legs). * Two distinct parts make up of the larval mandible (lower jaw). * The epandrium and hypandrium of the genitalia are separated in males. * No premandible is present on the lower surface of the labrum (the roof of the mouth). * The configuration of the CuA2 and A1 wing veins is distinct. Brachyceran flies can also be distinguished through behavior. Many of the species are predators or scavengers. Classification The structure of subgroups wit ...
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Diptera
Flies are insects of the order Diptera, the name being derived from the Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwings having evolved into advanced mechanosensory organs known as halteres, which act as high-speed sensors of rotational movement and allow dipterans to perform advanced aerobatics. Diptera is a large order containing an estimated 1,000,000 species including horse-flies, crane flies, hoverflies and others, although only about 125,000 species have been described. Flies have a mobile head, with a pair of large compound eyes, and mouthparts designed for piercing and sucking (mosquitoes, black flies and robber flies), or for lapping and sucking in the other groups. Their wing arrangement gives them great maneuverability in flight, and claws and pads on their feet enable them to cling to smooth surfaces. Flies undergo complete metamorphosis; the eggs are often laid on the l ...
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Brachycera
The Brachycera are a suborder of the order Diptera. It is a major suborder consisting of around 120 families. Their most distinguishing characteristic is reduced antenna segmentation. Description A summary of the main physical characteristics is: * Antenna size (with eight or fewer flagellomeres) is reduced. * The maxillary palp (an elongated appendage near the mouth) has two segments or fewer. * The back portions of the larval head capsule extend into the prothorax (the anterior part of the thorax, which bears the first pair of legs). * Two distinct parts make up of the larval mandible (lower jaw). * The epandrium and hypandrium of the genitalia are separated in males. * No premandible is present on the lower surface of the labrum (the roof of the mouth). * The configuration of the CuA2 and A1 wing veins is distinct. Brachyceran flies can also be distinguished through behavior. Many of the species are predators or scavengers. Classification The structure of subgroups wit ...
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Archisargoidea
} Archisargoidea is an extinct superfamily of brachyceran flies known from the late Middle Jurassic (Callovian) to early Late Cretaceous (Turonian). Most flies in the superfamily have large eyes and an elongated abdomen, preserved females have a sharp, piercing oviscapt used for injecting eggs into host matter. Their relationships with other members of Brachycera is controversial, they are usually considered close relatives of either Stratiomyomorpha or Muscomorpha. Internal relationships between the families are uncertain and the topology of the only cladistic analysis of the family was weakly supported, finding that Archisargidae was paraphyletic with respect to Eremochaetidae and miniaturized Tethepomyiidae. The sharp piercing oviscaps possessed by the group had alternatively been suggested to represent evidence of being parasitoids, injecting their eggs into hosts similar to parasitic wasps, or to inject eggs into plant material like rotting wood or fruit, similar to member ...
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Muscomorpha
The Brachyceran infraorder Muscomorpha is a large and diverse group of flies, containing the bulk of the Brachycera, and, most of the known flies. It includes a number of the most familiar flies, such as the housefly, the fruit fly, and the blow fly. The antennae are short, usually three-segmented, with a dorsal arista. Their bodies are often highly setose, and the pattern of setae is often taxonomically important. The larvae of muscomorphs (in the sense the name is used here; see below) have reduced head capsules, and the pupae are formed inside the exoskeleton of the last larval instar; exit from this puparium is by a circular line of weakness, and this pupal type is called "cyclorrhaphous"; this feature gives this group of flies their traditional name, Cyclorrhapha. Classification The name Cyclorrhapha is used, in various modern classifications, to represent either a subgroup within the infraorder Muscomorpha, or simply a rankless group within the Brachycera. In either ca ...
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Vermileonomorpha
The Brachyceran family Vermileonidae (the sole family in the infraorder Vermileonomorpha) is a small family of uncertain affinities and unusual biology. It includes fewer than 80 described species, most of them rare and with restricted distribution, in 10 genera. Historically the vermileonids had been regarded as belonging to the family Rhagionidae, possibly in a subfamily Vermileoninae. Their biology and morphology are so markedly distinct from the main Rhagionidae ''sensu stricto'' however, that the placement as a separate family has been widely accepted. Adult The adults are slender, fragile, long-legged flies, vaguely reminiscent of small crane flies. The adults generally visit flowers for nectar, but adults of some species may not feed at all. The mouthparts of the adult are hypognathous, used mainly for extracting nectar from flowers, long, and straight. This might have something to do with the common name "snipe-fly" for the family Rhagionidae, but it would be misleading to ...
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Asilomorpha
The Brachyceran infraorder Asilomorpha is a large and diverse group of flies, containing the bulk of the non muscoid Brachycera. The larvae of asilomorphs are extremely diverse in habits, as well. Classification In most modern classifications, the infraorder Asilomorpha is not recognized, as it is paraphyletic; the Empidoidea are the sister taxon to the Muscomorpha (used here in the same sense as the traditional Cyclorrhapha, not following the Tree of Life Web Project system in this one instance). However, these modern classifications are not Linnaean, and - while relatively easy to visualize on a phylogenetic tree diagram - do not lend themselves to any classification that uses Linnaean ranks (''see'' discussion ''for details''). Also, some classifications place the Nemestrinoidea Nemestrinoidea is a small, monophyletic superfamily of flies, whose relationship to the other Brachycera is uncertain; they are sometimes grouped with the Tabanomorpha rather than the Asilomorpha. Th ...
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Larger Brachycera
The larger brachycera is a name which refers to flies in the following families of the suborder Brachycera: * Acroceridae – hunch-back flies * Asilidae – robber flies * Athericidae – water snipe flies * Bombyliidae – bee flies * Rhagionidae – snipe flies * Scenopinidae – window flies * Stratiomyidae – soldier flies * Tabanidae – horse flies * Therevidae – stiletto flies * Xylomyidae – wood soldier flies * Xylophagidae The Brachyceran infraorder Xylophagomorpha is a small group that consists solely of the family Xylophagidae, which presently contains subfamilies that were sometimes considered to be two small related families (Coenomyiidae and Rachiceridae). ... – awl-flies References Brachycera {{Brachycera-stub ...
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Orthorrhapha
Orthorrhapha is a circumscriptional name which historically was used for an infraorder of Brachycera, one of the two suborders into which the order Diptera, the flies, are divided. As the group was paraphyletic, it has not been used in classifications in the last decade, and is effectively obsolete. However, many catalogs, checklists, and older works still contain the name. The taxa that used to be in the Orthorrhapha now comprise all of the infraorders in Brachycera excluding the Muscomorpha (= "Cyclorrhapha"). The recent revision of the taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ... of the order Diptera by Pape ''et al.'' revived the name Orthorrhapha. References *Cedric Gillott, 1980 ''Entomology'' Plenum Press, New York Brachycera Insect infraorders Obsolete ...
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Tabanomorpha
The Brachyceran infraorder Tabanomorpha is a small group that consists primarily of two large families, the Tabanidae (horse and deer flies) and Rhagionidae (snipe flies), and an assortment of very small affiliated families, most of which have been (or could be, or sometimes are) included within the Rhagionidae. Description Adult Tabanomorpha typically have a convex face and antenna bearing styli. The forewing has a costa along its entire perimeter (though its posterior portion may be weaker), while the tarsi have pulvilliform empodia. Males have eyes that are nearly or fully holoptic and have an endoaedeagal process which is usually quite long and distinct. Females have the cercus always flattened. Larval Tabanomorpha have a retractable head capsule and a brush of setae just under the fold of the integument. Ecology Adults of most species feed on nectar and pollen, but blood-feeding (hematophagy) occurs in the majority of female Tabanidae, some Rhagionidae and an Athericida ...
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Xylophagomorpha
The Brachyceran infraorder Xylophagomorpha is a small group that consists solely of the family Xylophagidae, which presently contains subfamilies that were sometimes considered to be two small related families (Coenomyiidae and Rachiceridae). Other obsolete names for members of this family include Exeretonevridae and Heterostomidae. The family is known by the English name awl-flies. The larvae are often predatory, consuming other insect larvae living in rotting wood. Description Flies in this family have elongated bodies and resemble ichneumon wasps in shape. The base of the abdomen is constricted. The antennae have three segments.Watson, L. and Dallwitz, M.J. 2003 onwardsXylophagidae. British Insects: The Families of Diptera. Version 1 January 2012. Genera These nine genera belong to the family Xylophagidae: *''Anacanthaspis'' Röder, 1889 *''Arthropeas'' Loew, 1850 *''Coenomyia'' Latreille, 1797 *''Dialysis'' Walker in Saunders, 1850 *''Exeretonevra'' Macquart, 1846 *' ...
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Wing Vein
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane (extreme examples include the dragonflies and lacewings). The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects. Physically, some insects move their flight muscles directly, others indirectly. In insects with direct flight, the wing muscles directly attach to the wing base, so that a small downward movement of the wing base lifts the wing itself upward. Those i ...
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Wing
A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expressed as its lift-to-drag ratio. The lift a wing generates at a given speed and angle of attack can be one to two orders of magnitude greater than the total drag on the wing. A high lift-to-drag ratio requires a significantly smaller thrust to propel the wings through the air at sufficient lift. Lifting structures used in water include various foils, such as hydrofoils. Hydrodynamics is the governing science, rather than aerodynamics. Applications of underwater foils occur in hydroplanes, sailboats and submarines. Etymology and usage For many centuries, the word "wing", from the Old Norse ''vængr'', referred mainly to the foremost limbs of birds (in addition to the architectural aisle). But in recent centuries the word's meaning has ...
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