Aquarism
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Fishkeeping is a popular
hobby A hobby is considered to be a regular activity that is done for enjoyment, typically during one's leisure time. Hobbies include collecting themed items and objects, engaging in creative and artistic pursuits, playing Sport, sports, or pursu ...
, practiced by
aquarist An aquarist is a person who manages aquariums, either professionally or as a hobby. They typically care for aquatic animals, including fish and marine invertebrates. Some may care for aquatic mammals. Aquarists often work at public aquariums. They ...
s, concerned with keeping
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
in a home
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
or
garden pond A garden pond is a water feature constructed in a garden or designed landscape, normally for aesthetic purposes, to provide wildlife habitat, or for swimming. Habitat Garden ponds can be excellent wildlife habitats, and can make a contribution ...
. There is also a piscicultural fishkeeping industry, serving as a branch of
agriculture Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to ...
.


Origins of fishkeeping

Fish have been raised as food in pools and ponds for thousands of years. Brightly colored or tame specimens of fish in these pools have sometimes been valued as pets rather than food. Many cultures, ancient and modern, have kept fish for both functional and decorative purposes. Ancient
Sumer Sumer () is the earliest known civilization in the historical region of southern Mesopotamia (south-central Iraq), emerging during the Chalcolithic and early Bronze Ages between the sixth and fifth millennium BC. It is one of the cradles of c ...
ians kept wild-caught fish in
pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from th ...
s, before preparing them for meals. Depictions of the sacred fish of
Oxyrhynchus Oxyrhynchus (; grc-gre, Ὀξύρρυγχος, Oxýrrhynchos, sharp-nosed; ancient Egyptian ''Pr-Medjed''; cop, or , ''Pemdje''; ar, البهنسا, ''Al-Bahnasa'') is a city in Middle Egypt located about 160 km south-southwest of Cairo ...
kept in captivity in rectangular temple pools have been found in
ancient Egyptian art Ancient Egyptian art refers to art produced in ancient Egypt between the 6th millennium BC and the 4th century AD, spanning from Prehistoric Egypt until the Christianization of Roman Egypt. It includes paintings, sculptu ...
. Similarly, Asia has experienced a long history of stocking
rice paddies A paddy field is a flooded field of arable land used for growing semiaquatic crops, most notably rice and taro. It originates from the Neolithic rice-farming cultures of the Yangtze River basin in southern China, associated with pre-Aust ...
with freshwater fish suitable for eating, including various types of
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
and
cyprinid Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...
.
Selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant mal ...
of
carp Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
into today's popular and completely domesticated
koi or more specifically , are colored varieties of the Amur carp ('' Cyprinus rubrofuscus'') that are kept for decorative purposes in outdoor koi ponds or water gardens. Koi is an informal name for the colored variants of ''C. rubrofuscus'' ke ...
and fancy
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have ...
began over 2,000 years ago in
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
and
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, respectively. The
Chinese Chinese can refer to: * Something related to China * Chinese people, people of Chinese nationality, citizenship, and/or ethnicity **''Zhonghua minzu'', the supra-ethnic concept of the Chinese nation ** List of ethnic groups in China, people of va ...
brought goldfish indoors during the
Song Dynasty The Song dynasty (; ; 960–1279) was an imperial dynasty of China that began in 960 and lasted until 1279. The dynasty was founded by Emperor Taizu of Song following his usurpation of the throne of the Later Zhou. The Song conquered the rest ...
to enjoy them in large ceramic vessels. In Medieval Europe, carp pools were a standard feature of estates and monasteries, providing an alternative to meat on
feast days The calendar of saints is the traditional Christian method of organizing a liturgical year by associating each day with one or more saints and referring to the day as the feast day or feast of said saint. The word "feast" in this context do ...
when meat could not be eaten for religious reasons. Marine fish have been similarly valued for centuries. Wealthy Romans kept
lamprey Lampreys (sometimes inaccurately called lamprey eels) are an ancient extant lineage of jawless fish of the order Petromyzontiformes , placed in the superclass Cyclostomata. The adult lamprey may be characterized by a toothed, funnel-like s ...
s and other fish in salt water pools.
Tertullian Tertullian (; la, Quintus Septimius Florens Tertullianus; 155 AD – 220 AD) was a prolific early Christian author from Carthage in the Roman province of Africa. He was the first Christian author to produce an extensive corpus of L ...
reports that
Asinius Celer Asinius is the nomen of the plebeian Gens Asinia of ancient Rome. Local tradition holds that the Italian town of Assignano derives its name from the gens, as well as the French town of Assignan. * Gaius Asinius Pollio, orator, poet, historian, con ...
paid 8000
sesterces The ''sestertius'' (plural ''sestertii''), or sesterce (plural sesterces), was an ancient Roman coin. During the Roman Republic it was a small, silver coin issued only on rare occasions. During the Roman Empire it was a large brass coin. The na ...
for a particularly fine mullet.
Cicero Marcus Tullius Cicero ( ; ; 3 January 106 BC – 7 December 43 BC) was a Roman statesman, lawyer, scholar, philosopher, and academic skeptic, who tried to uphold optimate principles during the political crises that led to the estab ...
reports that the
advocate An advocate is a professional in the field of law. Different countries' legal systems use the term with somewhat differing meanings. The broad equivalent in many English law–based jurisdictions could be a barrister or a solicitor. However, ...
Quintus Hortensius Quintus Hortensius Hortalus (114–50 BC) was a famous Roman lawyer, a renowned orator and a statesman. Politically he belonged to the Optimates. He was consul in 69 BC alongside Quintus Caecilius Metellus Creticus. His nickname was ''Dionysia'', ...
wept when a favored specimen died. Rather cynically, he referred to these ancient fishkeepers as the Piscinarii, the "fish-pond owners" or "fish breeders", for example when saying that "the rich (I mean your friends the fish-breeders) did not disguise their jealousy of me". The first person to breed a tropical fish in Europe was
Pierre Carbonnier Pierre Carbonnier, (7 August 1828 – 8 April 1883) was a French scientist, ichthyologist, fish breeder and public Aquarium director. Member of Imperial Society of acclimatization (Société Impériale d'Acclimatation). Biography Pierre Carb ...
, who founded one of the oldest
public aquaria A public aquarium (plural: ''public aquaria'' or ''public Water Zoo'') is the aquatic counterpart of a zoo, which houses living aquatic animal and plant specimens for public viewing. Most public aquariums feature tanks larger than those kept b ...
in Paris in 1850, and bred the first imported Macropods (
Paradise fish The paradise fish, paradise-fish, paradisefish, or paradise gourami (''Macropodus opercularis'') is a species of gourami found in most types of fresh water in East Asia, ranging from the Korean Peninsula to northern Vietnam. This species can ...
) in 1869, and later more species. A pioneer of tropical fish breeding, Carbonnier was awarded the Gold Medal of the Imperial French Acclimatization Society in 1875 for research and breeding of exotic freshwater aquarium fish, and for his success in introducing exotic fish species to France.


Types of fishkeeping systems

Fishkeepers are often known as "aquarists" since many of them are not solely interested in keeping fish. The hobby can be broadly divided into three specific disciplines, depending on the type of water the fish originate from:
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
,
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
, and
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
(also called saltwater) fishkeeping.


Freshwater

Freshwater fishkeeping is by far the most popular branch of the hobby, with even small pet stores often selling a variety of
freshwater fish Freshwater fish are those that spend some or all of their lives in fresh water, such as rivers and lakes, with a salinity of less than 1.05%. These environments differ from marine conditions in many ways, especially the difference in levels of s ...
, such as
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have ...
,
guppies The guppy (), also known as millionfish and rainbow fish, is one of the world's most widely distributed tropical fish and one of the most popular List of freshwater aquarium fish species, freshwater aquarium fish species. It is a member of the ...
, and angelfish. While most freshwater aquaria are community tanks containing a variety of compatible
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, single-species breeding aquaria are also popular. Livebearing fish such as mollies and guppies are among those most easily raised in captivity, but aquarists also regularly breed many types of
cichlid Cichlids are fish from the family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses ( Labridae), in the order Perciformes, but molecular studies have contradicted this ...
,
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
, characins,
cyprinids Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest vert ...
, and
killifish A killifish is any of various oviparous (egg-laying) cyprinodontiform fish (including families Aplocheilidae, Cyprinodontidae, Fundulidae, Profundulidae and Valenciidae). All together, there are 1,270 species of killifish, the biggest family ...
. Many fishkeepers create freshwater aquascapes where the focus is on
aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...
s as well as fish. These aquaria include "Dutch Aquaria" that mass contrasting stem plants, named for European aquarists who first designed them. In recent years, one of the most active advocates of the heavily planted aquarium was the Japanese aquarist
Takashi Amano was a professional track cyclist, photographer, designer, and aquarist. His interest in aquaria led him to create the Japanese company Aqua Design Amano. Amano was the author of ''Nature Aquarium World'' (TFH Publications, 1994), a three-boo ...
. Garden ponds are in some ways similar to freshwater aquaria, but are usually much larger and exposed to ambient weather. In the tropics, tropical fish can be kept in garden ponds. In the
temperate zone In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
, species such as goldfish,
koi or more specifically , are colored varieties of the Amur carp ('' Cyprinus rubrofuscus'') that are kept for decorative purposes in outdoor koi ponds or water gardens. Koi is an informal name for the colored variants of ''C. rubrofuscus'' ke ...
, and
orfe The ide (''Leuciscus idus''), or orfe, is a freshwater fish of the family Cyprinidae found in larger rivers, ponds, and lakes across Northern Europe and Asia. It has been introduced outside its native range into Europe, North America, and New Z ...
work better.


Saltwater

Marine aquaria A marine aquarium is an aquarium that keeps marine plants and animals in a contained environment. Marine aquaria are further subdivided by hobbyists into fish only (FO), fish only with live rock (FOWLR), and reef aquaria. Fish only tanks often sh ...
have more specific needs and requirements to maintain, and the livestock is generally more expensive. As a result, this branch tends to attract more experienced fishkeepers. Marine aquaria can be exceedingly beautiful, due to the attractive colors and shapes of the
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
s and the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
they host. Temperate zone marine fish are not as commonly kept in home aquaria, primarily because they do not thrive at room temperature. Coldwater aquaria must provide cooler temperature via a cool room (such as an unheated basement) or using a
refrigeration The term refrigeration refers to the process of removing heat from an enclosed space or substance for the purpose of lowering the temperature.International Dictionary of Refrigeration, http://dictionary.iifiir.org/search.phpASHRAE Terminology, ht ...
device known as a 'chiller'. Marine aquarists often attempt to recreate a coral reef in their aquaria using large quantities of living rock, porous calcareous rocks encrusted with
coralline algae Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of re ...
,
sponge Sponges, the members of the phylum Porifera (; meaning 'pore bearer'), are a basal animal clade as a sister of the diploblasts. They are multicellular organisms that have bodies full of pores and channels allowing water to circulate through t ...
s,
worm Worms are many different distantly related bilateral animals that typically have a long cylindrical tube-like body, no limbs, and no eyes (though not always). Worms vary in size from microscopic to over in length for marine polychaete wor ...
s, and other small marine organisms. Larger corals, as well as
shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
s,
crab Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the ...
s,
echinoderm An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea ...
s, and
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s are added later on, once the aquarium has matured, as well as a variety of small fish. Such aquaria are sometimes called reef tanks.


Brackish water

Brackish water aquaria combine elements of the other types, with
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
that must stay between that of freshwater and seawater. Brackish water fish come from habitats with varying salinity, such as
mangrove A mangrove is a shrub or tree that grows in coastal saline water, saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evoluti ...
s and
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
, and do not thrive if kept permanently in freshwater. Although brackish water aquaria are not necessarily familiar to inexperienced aquarists, many species prefer brackish water, including some mollies, many
gobies Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising more than 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and the ...
, some
pufferfish Tetraodontidae is a family of primarily marine and estuarine fish of the order Tetraodontiformes. The family includes many familiar species variously called pufferfish, puffers, balloonfish, blowfish, blowies, bubblefish, globefish, swellfis ...
,
monos Monos is an island in the Republic of Trinidad and Tobago. It is one of the "Bocas Islands", which lie in the ''Bocas del Dragón'' (''Dragons' Mouth'') between Trinidad and Venezuela. It is so named as the island was once home to noisy red ho ...
, and
scats The Sydney Coordinated Adaptive Traffic System, abbreviated SCATS, is an intelligent transportation system that manages the dynamic (on-line, real-time) timing of signal phases at traffic signals, meaning that it tries to find the best phasing (i ...
.


Aquarium maintenance

Ideal aquarium
ecology Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
reproduces the balance found in nature in the
closed system A closed system is a natural physical system that does not allow transfer of matter in or out of the system, although — in contexts such as physics, chemistry or engineering — the transfer of energy (''e.g.'' as work or heat) is allowed. In ...
of an aquarium. In practice, it is virtually impossible to maintain a perfect balance. As an example, a balanced
predator-prey relationship Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
is nearly impossible to maintain in even the largest aquaria. Typically, an aquarium keeper must actively maintain balance in the small
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
s that aquaria provide. Basic aquarium maintenance should generally be performed weekly to maintain optimum conditions for fish and plants. Balance is facilitated by larger volumes of water which dilute the effects of a
systemic shock A systemic shock is a shock to any system that perturbs a system enough to drive it out of equilibrium. Systemic shocks occur in a wide range of fields, ranging from medicine (see shock), ecology, economics to engineering. Designers of systems ...
. For example, the death of the only fish in a tank causes dramatic changes in the system, while the death of that same fish in a tank that holds many fish may create only a minor imbalance. For this reason, hobbyists often favor larger tanks whenever possible, as they require less intensive attention. This same concept extends to the filtration system as well, external (outside of the tank) systems in particular. Generally speaking, the larger the filtration system depending on its configuration, the more capable it will be of properly maintaining an aquatic environment. External filtration systems provide the added benefit of increasing the overall volume of water and its dilution effect. For example, a aquarium with an external filter that holds creates a aquatic system, an increase of over twenty percent. A variety of
nutrient cycles A nutrient cycle (or ecological recycling) is the movement and exchange of inorganic and organic matter back into the production of matter. Energy flow is a unidirectional and noncyclic pathway, whereas the movement of mineral nutrients is cycli ...
is important in the aquarium.
Dissolved oxygen Oxygen saturation (symbol SO2) is a relative measure of the concentration of oxygen that is dissolved or carried in a given medium as a proportion of the maximal concentration that can be dissolved in that medium at the given temperature. It ca ...
enters at the surface water-air interface through agitation or what would be observed as
waves Waves most often refers to: *Waves, oscillations accompanied by a transfer of energy that travel through space or mass. * Wind waves, surface waves that occur on the free surface of bodies of water. Waves may also refer to: Music * Waves (ban ...
in a natural environment, and
carbon dioxide Carbon dioxide (chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is transpar ...
escapes into the air. The
phosphate In chemistry, a phosphate is an anion, salt, functional group or ester derived from a phosphoric acid. It most commonly means orthophosphate, a derivative of orthophosphoric acid . The phosphate or orthophosphate ion is derived from phospho ...
cycle is an important, although often overlooked, nutrient cycle.
Sulfur Sulfur (or sulphur in British English) is a chemical element with the symbol S and atomic number 16. It is abundant, multivalent and nonmetallic. Under normal conditions, sulfur atoms form cyclic octatomic molecules with a chemical formula ...
,
iron Iron () is a chemical element with symbol Fe (from la, ferrum) and atomic number 26. It is a metal that belongs to the first transition series and group 8 of the periodic table. It is, by mass, the most common element on Earth, right in f ...
, and
micronutrient Micronutrients are nutrient, essential dietary elements required by organisms in varying quantities throughout life to orchestrate a range of physiological functions to maintain health. Micronutrient requirements differ between organisms; for exam ...
s enter the system as food and exit as waste. Appropriate handling of the
nitrogen cycle The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into multiple chemical forms as it circulates among atmospheric, terrestrial, and marine ecosystems. The conversion of nitrogen can be carried out through both biologi ...
, along with a balanced food supply and consideration of biological loading, is usually enough to keep these nutrient cycles in adequate equilibrium.


Water conditions

The
solute In chemistry, a solution is a special type of homogeneous mixture composed of two or more substances. In such a mixture, a solute is a substance dissolved in another substance, known as a solvent. If the attractive forces between the solvent ...
content of water is perhaps the most important aspect of water conditions, as
total dissolved solids Total dissolved solids (TDS) is a measure of the dissolved combined content of all inorganic and organic substances present in a liquid in molecular, ionized, or micro-granular ( colloidal sol) suspended form. TDS concentrations are often repor ...
and other constituents can dramatically impact basic water chemistry, and therefore how organisms interact with their environment. Salt content, or
salinity Salinity () is the saltiness or amount of salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensionless and equal ...
, is the most basic classification of water conditions. An aquarium may have
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
(salinity below 0.5 PPT), simulating a lake or river environment;
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estua ...
(a salt level of 0.5 to 30 PPT), simulating environments lying between fresh and salt, such as
estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
; and salt water or
seawater Seawater, or salt water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has appro ...
(a salt level of 30 to 40 PPT), simulating an ocean or sea environment. Even higher salt concentrations are maintained in specialized tanks for raising brine organisms. Several other water characteristics result from dissolved materials in the water and are important to the proper simulation of natural environments. Saltwater is typically
alkaline In chemistry, an alkali (; from ar, القلوي, al-qaly, lit=ashes of the saltwort) is a base (chemistry), basic, ionic compound, ionic salt (chemistry), salt of an alkali metal or an alkaline earth metal. An alkali can also be defined as ...
, while the pH of fresh water varies. "Hardness" measures overall dissolved mineral content; hard or soft water may be preferred. Hard water is usually alkaline, while soft water is usually neutral to acidic. Dissolved organic content and dissolved gases content are also important factors. Home aquarists typically use modified tap water supplied through their local
water supply network A water supply network or water supply system is a system of engineered hydrologic and hydraulic components that provide water supply. A water supply system typically includes the following: # A drainage basin (see water purification – source ...
. Because of the
chlorine Chlorine is a chemical element with the Symbol (chemistry), symbol Cl and atomic number 17. The second-lightest of the halogens, it appears between fluorine and bromine in the periodic table and its properties are mostly intermediate betwee ...
used to
disinfect A disinfectant is a chemical substance or compound used to inactivate or destroy microorganisms on inert surfaces. Disinfection does not necessarily kill all microorganisms, especially resistant bacterial spores; it is less effective than st ...
drinking water supplies for human consumption, tap water cannot be immediately used. In the past, it was possible to "condition" the water by simply letting the water stand for a day or two, which allows the chlorine to dissipate. However,
monochloramine Monochloramine, often called chloramine, is the chemical compound with the formula NH2Cl. Together with dichloramine (NHCl2) and nitrogen trichloride (NCl3), it is one of the three chloramines of ammonia. It is a colorless liquid at its melting p ...
became popular in
water treatment Water treatment is any process that improves the Water quality, quality of water to make it appropriate for a specific end-use. The end use may be drinking water, drinking, industrial water supply, irrigation, river flow maintenance, water recrea ...
because it stays longer in the water. Additives are available to remove chlorine or chloramine and suffice to make the water ready. Brackish or saltwater aquaria require the addition of a mixture of salts and other minerals. More sophisticated aquarists may modify the water's alkalinity, hardness, or dissolved content of organics and gases. This can be accomplished by additives such as
sodium bicarbonate Sodium bicarbonate (IUPAC name: sodium hydrogencarbonate), commonly known as baking soda or bicarbonate of soda, is a chemical compound with the formula NaHCO3. It is a salt composed of a sodium cation ( Na+) and a bicarbonate anion ( HCO3−) ...
to raise pH. Some aquarists
filter Filter, filtering or filters may refer to: Science and technology Computing * Filter (higher-order function), in functional programming * Filter (software), a computer program to process a data stream * Filter (video), a software component tha ...
or purify their water using one of two processes:
deionization Purified water is water that has been mechanically filtered or processed to remove impurities and make it suitable for use. Distilled water was, formerly, the most common form of purified water, but, in recent years, water is more frequently puri ...
or
reverse osmosis Reverse osmosis (RO) is a water purification process that uses a partially permeable membrane to separate ions, unwanted molecules and larger particles from drinking water. In reverse osmosis, an applied pressure is used to overcome osmotic pre ...
. In contrast, public aquaria with large water needs often locate themselves near a natural water source (such as a river, lake, or ocean) in order to have easy access to water that requires only minimal treatment. Water
temperature Temperature is a physical quantity that expresses quantitatively the perceptions of hotness and coldness. Temperature is measured with a thermometer. Thermometers are calibrated in various temperature scales that historically have relied o ...
forms the basis of one of the two most basic aquarium classifications:
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
vs. cold water. Most fish and plant species tolerate only a limited range of water temperatures: Tropical or warm water aquaria maintain an average temperature of about are much more common, and
tropical fish Tropical fish are generally those fish found in aquatic tropical environments around the world. Fishkeepers often keep tropical fish in freshwater and saltwater aquariums. The term "tropical fish" is not a taxonomic group, but rather is a gene ...
are among the most popular aquarium denizens. Cold water aquaria maintain temperatures below the room temperature. More important than the range is temperature consistency; most organisms are not accustomed to sudden changes in temperatures, which can cause
shock Shock may refer to: Common uses Collective noun *Shock, a historic commercial term for a group of 60, see English numerals#Special names * Stook, or shock of grain, stacked sheaves Healthcare * Shock (circulatory), circulatory medical emerge ...
and lead to disease. Water temperature can be regulated with a combined
thermometer A thermometer is a device that temperature measurement, measures temperature or a temperature gradient (the degree of hotness or coldness of an object). A thermometer has two important elements: (1) a temperature sensor (e.g. the bulb of a merc ...
and heating or cooling unit. Water movement can also be important in accurately simulating a natural ecosystem. Fish may prefer anything from nearly still water up to swift, simulated
currents Currents, Current or The Current may refer to: Science and technology * Current (fluid), the flow of a liquid or a gas ** Air current, a flow of air ** Ocean current, a current in the ocean *** Rip current, a kind of water current ** Current (stre ...
. Water movement can be controlled through the use of aeration from air pumps, powerhead pumps, and careful design of water flow (such as the location of filtration system points of inflow and outflow).


Nitrogen cycle

Fish are animals and generate waste as they metabolize food, which aquarists must manage. Fish, invertebrates,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, and some bacteria excrete
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
in the form of
ammonia Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogenous was ...
(which converts to
ammonium The ammonium cation is a positively-charged polyatomic ion with the chemical formula or . It is formed by the protonation of ammonia (). Ammonium is also a general name for positively charged or protonated substituted amines and quaternary a ...
in acidic water) and must then pass through the
nitrogen cycle The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into multiple chemical forms as it circulates among atmospheric, terrestrial, and marine ecosystems. The conversion of nitrogen can be carried out through both biologi ...
. Ammonia is also produced through the
decomposition Decomposition or rot is the process by which dead organic substances are broken down into simpler organic or inorganic matter such as carbon dioxide, water, simple sugars and mineral salts. The process is a part of the nutrient cycle and is e ...
of plant and animal matter, including
fecal Feces ( or faeces), known colloquially and in slang as poo and poop, are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the large intestine. Feces contain a relati ...
matter and other
detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...
. Nitrogen waste products become
toxic Toxicity is the degree to which a chemical substance or a particular mixture of substances can damage an organism. Toxicity can refer to the effect on a whole organism, such as an animal, bacterium, or plant, as well as the effect on a subst ...
to fish and other aquarium inhabitants above a certain concentration. Ammonia is toxic to fish and other aquatic life in large quantities and that is why many fish keepers purchase testing kits to monitor the levels of ammonia in their water as well as monitoring nitrites and nitrates which are also part of the nitrogen cycle. Ammonia is produced from fish waste and uneaten food, after this is created it is broken down into nitrites by beneficial bacteria that is present in a properly cycled aquarium. Nitrites are then further broken down into the less toxic nitrates which can be absorbed by aquarium plants and nitrates absorbing filter media. Aquarists also use water changes as a way to keep these toxins under control by removing water from the aquarium and vacuuming fish waste and food from the gravel and replacing it with fresh, treated water.


The process

A well-balanced tank contains organisms that
metabolize Metabolism (, from el, μεταβολή ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run cell ...
the waste products of other inhabitants.
Nitrogenous waste Metabolic wastes or excrements are substances left over from metabolic processes (such as cellular respiration) which cannot be used by the organism (they are surplus or toxic), and must therefore be excreted. This includes nitrogen compounds, w ...
is metabolized in aquaria by a group of
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
known as nitrifiers (genus ''
Nitrosomonas ''Nitrosomonas'' is a genus of Gram-negative bacteria, belonging to the Betaproteobacteria. It is one of the five genera of ammonia-oxidizing bacteria and, as an obligate chemolithoautotroph, uses ammonia (NH3) as an energy source and carbon diox ...
''). Nitrifying bacteria metabolize ammonia into
nitrite The nitrite polyatomic ion, ion has the chemical formula . Nitrite (mostly sodium nitrite) is widely used throughout chemical and pharmaceutical industries. The nitrite anion is a pervasive intermediate in the nitrogen cycle in nature. The name ...
, which is highly toxic to fish, even at low concentrations. Another type of bacteria (genus ''
Nitrospira ''Nitrospira'' (from Latin: nitro, meaning "nitrate" and Greek: spira, meaning "spiral") translate into “a nitrate spiral” is a genus of bacteria within the monophyletic clade of the Nitrospirota phylum. The first member of this genus was d ...
''), converts this nitrite into the less toxic compound,
nitrate Nitrate is a polyatomic ion A polyatomic ion, also known as a molecular ion, is a covalent bonded set of two or more atoms, or of a metal complex, that can be considered to behave as a single unit and that has a net charge that is not zer ...
. (Members of genus ''
Nitrobacter ''Nitrobacter'' is a genus comprising rod-shaped, gram-negative, and chemoautotrophic bacteria. The name ''Nitrobacter'' derives from the Latin neuter gender noun ''nitrum, nitri'', alkalis; the Ancient Greek noun βακτηρία'','' βακτ ...
'' were once thought to fill the role of ''Nitrospira'', and they may appear in "jump start" kits. Although ''Nitrobacter'' species could theoretically fill the same
niche Niche may refer to: Science *Developmental niche, a concept for understanding the cultural context of child development *Ecological niche, a term describing the relational position of an organism's species *Niche differentiation, in ecology, the ...
as ''Nitrospira'', it has recently been found that ''Nitrobacter'' are not present in detectable levels in established aquaria, while ''Nitrospira'' is plentiful.) This process represents a portion of the
nitrogen cycle The nitrogen cycle is the biogeochemical cycle by which nitrogen is converted into multiple chemical forms as it circulates among atmospheric, terrestrial, and marine ecosystems. The conversion of nitrogen can be carried out through both biologi ...
. In a planted aquarium, aquatic plants also metabolize
ammonium The ammonium cation is a positively-charged polyatomic ion with the chemical formula or . It is formed by the protonation of ammonia (). Ammonium is also a general name for positively charged or protonated substituted amines and quaternary a ...
and nitrate as
nutrient A nutrient is a substance used by an organism to survive, grow, and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi, and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
s, removing them from the
water column A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
primarily through leaf surfaces. Plants remove some nutrients through their roots, either in or at the substrate level or via
aerial roots Aerial roots are roots above the ground. They are almost always adventitious. They are found in diverse plant species, including epiphytes such as orchids (''Orchidaceae''), tropical coastal swamp trees such as mangroves, banyan figs ('' Fi ...
floating in the water. Additional nitrogen and other nutrients are also made available for root uptake by decomposing organic matter in the substrate as well as the breakdown of mulm. While very small amounts of rotting foliage may be allowed to decompose and cycle nitrogen back into a planted aquarium, in practice aquarists will prune and remove substantial amounts of plant litter.


Maintaining the nitrogen cycle

Although called the nitrogen "cycle" by hobbyists, in aquaria the cycle is not complete: nitrogen must be added (usually indirectly through food) and nitrates must be removed at the end. Nitrogen bound up in plant matter is removed when the plant grows too large. Hobbyist aquaria typically do not have the requisite bacteria needed to detoxify nitrogen waste. This problem is most often addressed through
filtration Filtration is a physical separation process that separates solid matter and fluid from a mixture using a ''filter medium'' that has a complex structure through which only the fluid can pass. Solid particles that cannot pass through the filter ...
.
Activated carbon Activated carbon, also called activated charcoal, is a form of carbon commonly used to filter contaminants from water and air, among many other uses. It is processed (activated) to have small, low-volume pores that increase the surface area avail ...
filters absorb nitrogen compounds and other
toxin A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849– ...
s from the water. Biological filters provide a medium specially designed for
colonization Colonization, or colonisation, constitutes large-scale population movements wherein migrants maintain strong links with their, or their ancestors', former country – by such links, gain advantage over other inhabitants of the territory. When ...
by the desired nitrifying bacteria. Activated carbon and other substances, such as ammonia absorbing
resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on natu ...
s, stop working when their
pore Pore may refer to: Biology Animal biology and microbiology * Sweat pore, an anatomical structure of the skin of humans (and other mammals) used for secretion of sweat * Hair follicle, an anatomical structure of the skin of humans (and other m ...
s fill, so these components have to be replaced with fresh stocks periodically. New aquaria often have problems associated with the nitrogen cycle due to insufficient beneficial bacteria, which is known as "New Tank Syndrome". Therefore, new tanks have to mature before stocking them with fish. There are three basic approaches to this: the ''fishless cycle'', the ''silent cycle'', and ''slow growth''. *Tanks undergoing a "fishless cycle" have no fish. Instead, the keeper adds ammonia to feed the bacteria. During this process,
ammonia Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogenous was ...
,
nitrite The nitrite polyatomic ion, ion has the chemical formula . Nitrite (mostly sodium nitrite) is widely used throughout chemical and pharmaceutical industries. The nitrite anion is a pervasive intermediate in the nitrogen cycle in nature. The name ...
, and
nitrate Nitrate is a polyatomic ion A polyatomic ion, also known as a molecular ion, is a covalent bonded set of two or more atoms, or of a metal complex, that can be considered to behave as a single unit and that has a net charge that is not zer ...
levels measure progress. *The "silent cycle" involves adding fast-growing
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
s and relying on them to consume the
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
, filling in for the bacteria work until their number increases. Anecdotal reports indicate that such plants can consume nitrogenous waste so efficiently that the ammonia and nitrite spikes that occur in more traditional cycling methods are greatly reduced or undetectable. *"Slow growth" entails slowly increasing the fish population over 6 to 8 weeks, giving bacteria time to grow and reach a balance with the increasing waste production. Adding too many fish too quickly or failing to allow enough time for the bacteria colony to establish itself in the filter media can lead to ammonia stress. This is not always fatal but can result in the death of aquarium fish. A few days after adding hardy fish for the cycling process, it is essential to look out for the key signs of ammonia stress. These include a lack of movement and appetite, inflammation and redness of the gills, fins, and body, and occasionally gasping for air at the water's surface. The latter can also be attributed to poor aeration, which can be negated by the inclusion of an air pump or spray bar in the setup. The largest bacterial populations inhabit the filter; efficient filtration is vital. Sometimes, simply cleaning the filter is enough to seriously disturb the aquarium's balance.
Best practice A best practice is a method or technique that has been generally accepted as superior to other known alternatives because it often produces results that are superior to those achieved by other means or because it has become a standard way of doing ...
is to flush mechanical filters using compatible water to dislodge organic materials while preserving bacteria populations. Another safe practice involves cleaning only one-half of the filter media every time the filter or filters are serviced to allow the remaining bacteria to repopulate the cleaned half.


Tank capacity

Biological loading is a measure of the burden placed on the aquarium ecosystem by its living inhabitants. Higher biological loading represents a more complicated ecology, which makes equilibrium easier to imbalance. The
surface area The surface area of a solid object is a measure of the total area that the surface of the object occupies. The mathematical definition of surface area in the presence of curved surfaces is considerably more involved than the definition of arc ...
of water exposed to air limits
dissolved oxygen Oxygen saturation (symbol SO2) is a relative measure of the concentration of oxygen that is dissolved or carried in a given medium as a proportion of the maximal concentration that can be dissolved in that medium at the given temperature. It ca ...
. The population of nitrifying bacteria is limited by the available physical space which includes all surfaces in the aquarium such as the inner facing sides and the surface of rock substrate and any objects such as large rocks or pieces of wood.


Tank size

Fish capacity is a function of aquarium size.
Limiting factor A limiting factor is a variable of a system that causes a noticeable change in output or another measure of a type of system. The limiting factor is in a pyramid shape of organisms going up from the producers to consumers and so on. A factor not l ...
s include the availability of
oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements as wel ...
in the water and the rate at which the
filter Filter, filtering or filters may refer to: Science and technology Computing * Filter (higher-order function), in functional programming * Filter (software), a computer program to process a data stream * Filter (video), a software component tha ...
can process waste. Aquarists apply rules of thumb estimating appropriate population size; the examples below are for small freshwater fish. Larger freshwater fish and most marine fishes need much more generous allowances. Some aquarists claim that increasing water depth beyond some relatively shallow minimum does not affect capacity. * 1.5 litres of water for each centimetre of fish length (1 US gallon per inch).Ulrich Baensch, Tropical Aquarium Fish, Tetra, 1983 * 30 square centimetres of surface area per centimetre of fish length (12 square inches per inch).Peter Scott, The Complete Aquarium, Dorling Kindersley, 1996, Experienced aquarists warn against mechanically applying these rules because they do not consider other important issues such as growth rate, activity level, social behavior, and such. Once the tank nears capacity, the best practice is to add the remaining fish over a period of time while monitoring water quality. The capacity can be improved by surface movement and water circulation such as through aeration, which not only improves oxygen exchange but also the decomposition of waste materials. Capacity can also be increased with the addition of external filtration which increases the total volume of water in the aquatic system.


Other factors

Other variables affect tank capacity. Smaller fish consume more oxygen per unit of body weight than larger fish.
Labyrinth fish The Anabantoidei are a suborder of anabantiform ray-finned freshwater fish distinguished by their possession of a lung-like labyrinth organ, which enables them to breathe air. The fish in the Anabantoidei suborder are known as anabantoids or ...
can breathe atmospheric oxygen and need less surface area (however, some are territorial, and do not tolerate crowding). Barbs require more surface area than
tetra Terrestrial Trunked Radio (TETRA; formerly known as Trans-European Trunked Radio), a European standard for a trunked radio system, is a professional mobile radio and two-way transceiver specification. TETRA was specifically designed for use by go ...
s of comparable size. The presence of waste materials presents itself as a variable as well. Decomposition consumes oxygen, reducing the amount available for fish. Oxygen dissolves less readily in warmer water, while warmer water temperature increase fish activity levels, which in turn consume more oxygen.


Fishkeeping industry

Worldwide, the fishkeeping hobby is a multibillion-dollar industry. The United States is the largest market, followed by Europe and Japan. In 1993 the
United States Census Bureau The United States Census Bureau (USCB), officially the Bureau of the Census, is a principal agency of the U.S. Federal Statistical System, responsible for producing data about the American people and economy. The Census Bureau is part of the ...
found that 10.6% of U.S. households owned ornamental freshwater or saltwater fish, with an average of 8.8 fish per household. In 2002, census data indicated that aquarium products and fishing accounted for US$684 million.


Aquatic suppliers

From 1989 to 1992, almost 79% of all U.S. ornamental fish imports came from
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
and Japan. Singapore, Thailand, the Philippines, Hong Kong, and Indonesia were the top five exporting nations. South America was the second largest exporting region, accounting for 14% of the total annual value. Colombia, Brazil, and Peru were the major suppliers. Approximately 200 million fish worth $44.7 million were imported into the United States in 1992. These fish comprised 1,539 different
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
; 730
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
species, and 809
saltwater Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
species. Freshwater fish accounted for approximately 96% of the total volume and 80% of the total import value. Only 32 species had import values over $10,000. The top species were freshwater and accounted for 58% of the total imported value. The top imported species are the
guppy The guppy (), also known as millionfish and rainbow fish, is one of the world's most widely distributed tropical fish and one of the most popular freshwater aquarium fish species. It is a member of the family Poeciliidae and, like almost all ...
,
neon tetra The neon tetra (''Paracheirodon innesi'') is a Freshwater fish of the characin family (family Characidae) of order Characiformes. The type species of its genus, it is native to blackwater and clearwater streams in the Amazon basin of South Am ...
, platy,
betta ''Betta'' is a large genus of small, active, often colorful, freshwater ray-finned fishes, in the gourami family (Osphronemidae). The best known ''Betta'' species is ''B. splendens,'' commonly known as the Siamese fighting fish and often kept ...
,
Chinese algae eater ''Gyrinocheilus aymonieri'' is a freshwater fish native to large parts of Southeast Asia. It is of interest as a local food source and for the aquarium trade. Its common names include honey sucker, sucking loach and Chinese algae eater. ''G. ay ...
, and
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have ...
. Given 91.9 million total US households in 1990, 9.7 million are fishkeepers. 8.8 fish per household implies a total aquarium fish population of approximately 85.7 million, suggesting that the US aquarium fish population turns over more than 2.3 times per year, counting only imported fish. Historically, fish and plants for the first modern aquaria were gathered from the wild and transported (usually by ship) to Europea and America. During the early 20th century many species of small colorful
tropical fish Tropical fish are generally those fish found in aquatic tropical environments around the world. Fishkeepers often keep tropical fish in freshwater and saltwater aquariums. The term "tropical fish" is not a taxonomic group, but rather is a gene ...
were exported from Manaus, Brazil; Bangkok, Thailand; Jakarta, Indonesia; the Netherlands Antilles; Kolkata, India; and other tropical countries. Import of wild fish, plants, and invertebrates for aquaria continues today around the world. Many species have not been successfully bred in captivity. In many developing countries, locals survive by collecting specimens for the aquarium trade and continue to introduce new species to the market.


Animal welfare

Fish are sometimes maintained in inadequate conditions by inexperienced aquarists who often keep too many fish in one tank, or add fish too quickly into an immature aquarium, killing many of them. This has given the hobby a bad reputation among some animal welfare groups, such as
PETA Peta or PETA may refer to: Acronym * Pembela Tanah Air, a militia established by the occupying Japanese in Indonesia in 1943 * People for the Ethical Treatment of Animals, an American animal rights organization * People Eating Tasty Animals, an ...
, who accuse aquarists of treating aquarium fish as cheap toys to be replaced when they die. Goldfish and bettas, in particular, have often been kept in small bowls or aquaria that are too small for their needs. In some cases, fish have been installed in all sorts of inappropriate objects such as the "AquaBabies Micro Aquaria", "Bubble Gear Bubble Bag", and "Betta in a Vase", all of which house live fish in unfiltered and insufficient water. The last is sometimes marketed as a complete ecosystem because a plant is included in the neck of the vase. Some sellers say the fish eat the plant roots. However,
betta ''Betta'' is a large genus of small, active, often colorful, freshwater ray-finned fishes, in the gourami family (Osphronemidae). The best known ''Betta'' species is ''B. splendens,'' commonly known as the Siamese fighting fish and often kept ...
s are
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
and need live food or pellet foods. They cannot survive on plant roots. Another problem is that the plant sometimes blocks the betta's passage to the water surface. They are
labyrinth fish The Anabantoidei are a suborder of anabantiform ray-finned freshwater fish distinguished by their possession of a lung-like labyrinth organ, which enables them to breathe air. The fish in the Anabantoidei suborder are known as anabantoids or ...
and need to breathe at the surface to avoid suffocation. Such products are aimed at people looking for a novelty gift. Aquarists actively condemn them. Similarly, the awarding of goldfish as prizes at
funfair A fair (archaic: faire or fayre) is a gathering of people for a variety of entertainment or commercial activities. Fairs are typically temporary with scheduled times lasting from an afternoon to several weeks. Types Variations of fairs incl ...
s is traditional in many parts of the world but has been criticized by aquarists and activists as cruel and irresponsible. The United Kingdom outlawed live-animal prizes such as goldfish in 2004. The use of live prey to feed carnivorous fish such as
piranha A piranha or piraña (, , or ; or , ) is one of a number of freshwater fish in the family Serrasalmidae, or the subfamily Serrasalminae within the tetra family, Characidae in order Characiformes. These fish inhabit South American rivers, ...
s also draws criticism. ;Fish modification Modifying fish to make them more attractive as pets is increasingly controversial. Historically, artificially dyeing fish was common.
Glassfish GlassFish is an open-source Jakarta EE platform application server project started by Sun Microsystems, then sponsored by Oracle Corporation, and now living at the Eclipse Foundation and supported by Payara, Oracle and Red Hat. The supported v ...
, in particular, were often injected with fluorescent dyes. The British fishkeeping magazine ''
Practical Fishkeeping ''Practical Fishkeeping'' (also known as ''PFK'') is a United Kingdom-based aquarium magazine. It is published every four weeks by Warners Publications Plc. The title covers the entire aquatic market from tropical freshwater and tropical marin ...
'' has campaigned to remove these fish from the market by educating retailers and aquarists to the cruelty and health risks involved. In 2006, ''Practical Fishkeeping'' published an article exposing the techniques for performing
cosmetic surgery Plastic surgery is a surgical specialty involving the restoration, reconstruction or alteration of the human body. It can be divided into two main categories: reconstructive surgery and cosmetic surgery. Reconstructive surgery includes craniofa ...
on aquarium fish, without
anesthesia Anesthesia is a state of controlled, temporary loss of sensation or awareness that is induced for medical or veterinary purposes. It may include some or all of analgesia (relief from or prevention of pain), paralysis (muscle relaxation), ...
, as described by
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...
an fishkeeping magazine ''Fish Love Magazine''. The tail is cut off and dye is injected into the body. The piece also included the first documented evidence to demonstrate that parrot cichlids are injected with coloured dye. Hong Kong suppliers were offering a service in which fish could be
tattoo A tattoo is a form of body modification made by inserting tattoo ink, dyes, and/or pigments, either indelible or temporary, into the dermis layer of the skin to form a design. Tattoo artists create these designs using several Process of tatt ...
ed with company logos or messages using a
dye laser A dye laser is a laser that uses an organic dye as the lasing medium, usually as a liquid solution. Compared to gases and most solid state lasing media, a dye can usually be used for a much wider range of wavelengths, often spanning 50 to 100 na ...
; such fishes have been sold in the UK under the name of kaleidoscope gourami and striped parrot cichlid. Some people give their fish
body piercing Body piercing, which is a form of body modification, is the practice of puncturing or cutting a part of the human body, creating an opening in which jewelry may be worn, or where an implant could be inserted. The word ''piercing'' can refer to ...
s. Hybrid fish such as
flowerhorn cichlid Flowerhorn cichlids are ornamental aquarium fish noted for their vivid colors and the distinctively shaped heads for which they are named. Their head protuberance is formally called a Nuchal lines, nuchal hump. Like blood parrot cichlids, they ar ...
s and
blood parrot cichlid The blood parrot cichlid (''Amphilophus citrinellus'' × ''Vieja melanurus''), or parrot cichlid, is a hybrid species of fish in your the family Cichlidae. The fish was first bred in Taiwan around 1986. Blood parrots should not be confused wit ...
s are controversial. Blood parrot cichlids in particular have a very unnatural shape that prevents them from swimming properly and makes it difficult for them to engage in normal feeding and social behaviors. The biggest concern with hybrids is that they may be bred with native species, making it difficult for hobbyists to identify and breed particular species. This is especially important to hobbyists who shelter species that are rare or
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
in the wild. Extreme mutations have been selected for by some breeders; some fancy
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have ...
varieties in particular have features that prevent the fish from swimming, seeing, or feeding properly.
Genetically modified Genetic engineering, also called genetic modification or genetic manipulation, is the modification and manipulation of an organism's genes using technology. It is a set of Genetic engineering techniques, technologies used to change the gene ...
fish such as the
GloFish The GloFish is a patented and trademarked brand of genetically engineered fluorescent fish. They have been created from several different species of fish: zebrafish (''Danio rerio'') were the first GloFish available in pet stores, and recently te ...
are likely to become increasingly available, particularly in the United States and Asia. Although GloFish are unharmed by their genetic modifications, they remain illegal in many places, including the European Union, though at least some have been
smuggled Smuggling is the illegal transportation of objects, substances, information or people, such as out of a house or buildings, into a prison, or across an international border, in violation of applicable laws or other regulations. There are various ...
into the EU, most likely from Taiwan, via the Czech Republic.


Fish breeding

Fish breeding is a challenge that attracts many aquarists. While some species reproduce freely in community tanks, most require special conditions, known as
spawning triggers Spawning triggers'' are environmental cues that cause marine animals to breed. Most commonly they involve sudden changes in the environment, such as changes in temperature, salinity, and/or the abundance of food. Catfish of the genus ''Corydoras'', ...
before they will breed. The majority of fish lay eggs, known as
spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquati ...
, and the juvenile fish that emerge are very small and need tiny live food or substitutes to survive. A fair number of popular aquarium fish are
livebearers Livebearers are aquarium fish that retain the eggs inside the body and give birth to live, free-swimming young. Among aquarium fish, livebearers are nearly all members of the family Poeciliidae and include guppies, Sailfin molly, mollies, platies ...
which produce a small number of relatively large offspring. These usually eat fish flakes that are ground into an appropriate size.


Conservation

The two main sources of fish for aquaria are from capture in the wild or captive breeding. United Nations studies show that more than 90% of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
aquarium fish are captive–bred, while virtually all
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
fish and invertebrates are wild-caught. Freshwater fish habitats are threatened by habitat erosion from pollution, dams and hydropower, commercial fishing, climate change, the introduction of invasive species and disease. The few marine species bred in captivity supplement but rarely replace the trade in wild-caught specimens. Wild-caught animals provide valuable income for people in regions lacking other high-value
export An export in international trade is a good produced in one country that is sold into another country or a service provided in one country for a national or resident of another country. The seller of such goods or the service provider is an ...
s. Marine fish are typically less resilient during transport than freshwater fish with relatively large numbers of them dying before they reach the aquarist. Although the aquarium trade is viewed as a minor threat to coral reefs compared to habitat destruction, fishing for food, and climate change, it is a booming trade and may be a serious problem in specific locations such as the Philippines and Indonesia where most collecting is done. Catching fish in the wild can reduce their population sizes, placing them in danger of extinction in collecting areas, as has been observed with the
dragonet Dragonets are small, percomorph, marine fish of the diverse family Callionymidae (from the Greek ''kallis'', "beautiful" and ', "name") found mainly in the tropical waters of the western Indo-Pacific. They are benthic organisms, spending most ...
''
Synchiropus splendidus ''Synchiropus splendidus'', the mandarinfish or mandarin dragonet, is a small, brightly colored member of the dragonet family, which is popular in the saltwater aquarium trade. The mandarinfish is native to the Pacific, ranging approximately from ...
''.


Collecting

In theory, reef fish should be a good example of a
renewable resource A renewable resource, also known as a flow resource, is a natural resource which will replenish to replace the portion resource depletion, depleted by usage and consumption, either through natural reproduction or other recurring processes in a ...
that encourages fishermen to maintain the integrity and diversity of the natural habitat: more and better fish can be exported from pristine habitats than those that have been polluted or over-harvested. However, this has not been the case in similar industries such as
fur trapping Animal trapping, or simply trapping or gin, is the use of a device to remotely catch an animal. Animals may be trapped for a variety of purposes, including food, the fur trade, hunting, pest control, and wildlife management. History Neolithic ...
,
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidding, on-site processing, and loading of trees or logs onto trucks or skeleton cars. Logging is the beginning of a supply chain ...
, or
fishing Fishing is the activity of trying to catch fish. Fish are often caught as wildlife from the natural environment, but may also be caught from stocked bodies of water such as ponds, canals, park wetlands and reservoirs. Fishing techniques inclu ...
that experience the ''
tragedy of the commons Tragedy (from the grc-gre, τραγῳδία, ''tragōidia'', ''tragōidia'') is a genre of drama based on human suffering and, mainly, the terrible or sorrowful events that befall a main character. Traditionally, the intention of tragedy ...
''. Fish are caught by net, trap, or cyanide. Collecting expeditions can be lengthy and costly, and are not always successful. Fish can also be injured during collection and/or shipping;
mortality rate Mortality rate, or death rate, is a measure of the number of deaths (in general, or due to a specific cause) in a particular population, scaled to the size of that population, per unit of time. Mortality rate is typically expressed in units of de ...
s during shipping are high. Many others are weakened by
stress Stress may refer to: Science and medicine * Stress (biology), an organism's response to a stressor such as an environmental condition * Stress (linguistics), relative emphasis or prominence given to a syllable in a word, or to a word in a phrase ...
and become diseased. Other problems include the poisoning of
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
s and non-target species, the depletion of rare species from their natural habitat, and ecosystem degradation from large scale removal of key species. Additionally, destructive fishing techniques concern environmentalists and hobbyists alike. There has been a concerted movement to captive breeding and certification programs for wild-caught fish. Two thirds of American marine aquarists surveyed in 1997 preferred farmed coral and over 80% think that only sustainably caught or farmed fish should be traded. Annually, 30 million fish from over 1,400 species are traded, with approximately 16 million imported into the United States. This trade has an estimated global value of over $800 million in 2002.


Cyanide

Cyanide Cyanide is a naturally occurring, rapidly acting, toxic chemical that can exist in many different forms. In chemistry, a cyanide () is a chemical compound that contains a functional group. This group, known as the cyano group, consists of ...
is a poison that stuns and immobilizes fish. Fishermen put cyanide in the ocean to ease the process of netting fish. It can irreversibly damage or kill the target fish, as well as other fish, mammals, reptiles or invertebrates that are left behind. Some wholesalers advertise that they avoid cyanide-caught animals. In the Philippines,
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in th ...
and cyanide caused a drastic decline in aquarium fish, leading the country away from cyanide and towards netting healthy animals.


Captive breeding and aquaculture

Since the
Siamese fighting fish The Siamese fighting fish (''Betta splendens''), commonly known as the betta, is a freshwater fish native to Southeast Asia, namely Cambodia, Laos, Myanmar, Malaysia, Indonesia, Thailand, and Vietnam. It is one of 73 species of the genus ''Bett ...
(''
Betta splendens The Siamese fighting fish (''Betta splendens''), commonly known as the betta, is a freshwater fish native to Southeast Asia, namely Cambodia, Laos, Myanmar, Malaysia, Indonesia, Thailand, and Vietnam. It is one of 73 species of the genus ''Bett ...
'') was first successfully bred in France in 1893, captive spawning and brooding techniques used in
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
have slowly improved. Captive breeding for aquaria is concentrated in southern Florida, Singapore, Hong Kong and Bangkok, with smaller industries in Hawaii and Sri Lanka. Captive breeding of marine organisms has been in development since the mid-1990s. Breeding for freshwater species is more advanced than for
saltwater Saline water (more commonly known as salt water) is water that contains a high concentration of dissolved salts (mainly sodium chloride). On the United States Geological Survey (USGS) salinity scale, saline water is saltier than brackish water, ...
species. Currently, only a few captive-bred marine species are in the trade, including
clownfish Clownfish or anemonefish are fishes from the subfamily Amphiprioninae in the family Pomacentridae. Thirty species of clownfish are recognized: one in the genus ''Premnas'', while the remaining are in the genus ''Amphiprion''. In the wild, th ...
,
damselfish Damselfish are those within the subfamilies Abudefdufinae, Chrominae, Lepidozyginae, Pomacentrinae, and Stegastenae within the family Pomacentridae. Most species within this group are relatively small, with the largest species being about 30 ...
, and dwarf angelfish. Aquaculture can help in lessening the impacts on wild stocks, either by using cultivated organisms for sale or by releasing them to replenish wild stocks. Breeding programs help preserve species that have become rare or extinct in the wild, most notably the
Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after ...
cichlids. Some species have also become important as laboratory animals.
Cichlids Cichlids are fish from the family (biology), family Cichlidae in the order Cichliformes. Cichlids were traditionally classed in a suborder, the Labroidei, along with the wrasses (Labridae), in the order Perciformes, but molecular studies have c ...
, poecilids and zebra danios are especially important for studies on learning, mating, and social behavior. Hobbyists also keep and observe many fishes not otherwise studied, and thereby provide valuable ecological and behavioral data. Captive breeding has reduced prices for hobbyists, but cultured animals remain more expensive. Selective breeding has also led to wider intra–species variation, creating more diverse commercial stocks.


Invasive species

Serious problems can occur when fish originally raised in ponds or aquaria are released into the wild. While
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
fish do not survive in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
climates, they can thrive in waters that are similar to their native habitat. Non–native species that become established are called
exotic species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
.Azevedo-Santos, V.M.; Pelicice, F.M.; Lima-Junior, D.P.; Magalhães, A.L.B.; Orsi, M.L.; Vitule, J. R. S. & A.A. Agostinho, 2015. How to avoid fish introductions in Brazil: education and information as alternatives. Natureza & Conservação, in press. Freshwater examples include various cichlids in Florida,
goldfish The goldfish (''Carassius auratus'') is a freshwater fish in the family Cyprinidae of order Cypriniformes. It is commonly kept as a pet in indoor aquariums, and is one of the most popular aquarium fish. Goldfish released into the wild have ...
in temperate waters, and
South American South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
suckermouth catfishes in warm waters around the world.
Invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
can seriously disrupt their new homes by preying on, or competing with, native species. Many marine fish have also been introduced into non-native waters, disrupting the local habitat.


Humane treatment

In January 2011 the
Maui The island of Maui (; Hawaiian: ) is the second-largest of the islands of the state of Hawaii at 727.2 square miles (1,883 km2) and is the 17th largest island in the United States. Maui is the largest of Maui County's four islands, which ...
County Council passed a measure requiring
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
fisheries Fishery can mean either the enterprise of raising or harvesting fish and other aquatic life; or more commonly, the site where such enterprise takes place ( a.k.a. fishing ground). Commercial fisheries include wild fisheries and fish farms, both ...
to adopt humane practices for preparing fish for transport to market. The regulations control harvesting and shipping practices, including prohibiting clipping the fins on fish to protect the plastic shipping bags, outlawing puncturing
swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled Organ (anatomy), organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their curren ...
s that fish use to regulate their buoyancy, which enabled divers to rapidly surface and prohibiting "starving" the fish which permitted smaller shipping bags without killing the fish with their own waste. The measure also requires that shippers file mortality reports on the animals they ship.


See also

*
Fish farming upright=1.3, Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye">mariculture.html" ;"title="Salmon farming in the sea (mariculture">Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye, Scotland Fish farming or ...
* List of aquarium fish by scientific name


References


Further reading

* ''Advanced Marine Aquarium Techniques'', by Jay Hemdal * ''Aquarium Atlas, vol. 1'', by Hans A. Baensch and Rudiger Riehl * ''Brackish Water Fishes'', by Frank Schäfer * ''The Conscientious Marine Aquarist'', by Robert Fenner (2001) * ''The New Marine Aquarium'', by Michael S. Paletta (2001) *


External links

* {{Authority control Ichthyology Fish diseases