''Albertadromeus'' is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

orodromine Orodrominae is a subfamily of thescelosaurid dinosaurs known from the Cretaceous of North America and Asia. Distribution Orodromines were a mostly North American based group with fossils from Canada and United States only. ''Albertadromeus'', a ...

thescelosaurid Thescelosauridae is a clade of neornithischians from the Cretaceous of Asia, North America and possibly South America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

known from the upper part of the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

Oldman Formation The Oldman Formation is a stratigraphic unit of Late Cretaceous (Campanian stage) age that underlies much of southern Alberta, Canada. It consists primarily of sandstones that were deposited in fluvial channel and floodplain environments. It was ...

(middle

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...

stage) of

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

, Canada. It contains a single species, ''Albertadromeus syntarsus''.

Etymology

The composite term ''Albertadromeus'' is derived from the name of the Canadian province "Alberta", and the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

word ''dromeus'' (δρομεύς) meaning "runner", a reference to its inferred cursorial nature; thus "runner from Alberta". The specific name, ''syntarsus'' is derived from Greek words "syn" (συν) meaning "together" and "tarsus" (ταρσός) meaning "ankle", hence "together-ankle" a reference to the condition where its distal fibula is fused to its distal tibia. This dinosaur was described and named by Caleb Marshall Brown, David C. Evans, Michael J. Ryan & Anthony P. Russell in

2013 File:2013 Events Collage V2.png, From left, clockwise: Edward Snowden becomes internationally famous for leaking classified NSA wiretapping information; Typhoon Haiyan kills over 6,000 in the Philippines and Southeast Asia; The Dhaka garment fact ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is ''Albertadromeus syntarsus''.

Description

The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

specimen of ''Albertadromeus'' TMP 2009.037.0044 consists of two dorsal vertebrae, a caudal vertebra, cervical ribs, ossified tendons, the left

tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...

and

fibula The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is ...

, an incomplete right fibula, and a fragmentary

metatarsal The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...

and

ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropods and horned dinosaurs. A claw is a highly modified ungual ...

. The skull is unknown in this genus. The authors note that despite the few bones recovered, their moderate quality of preservation nonetheless provides enough morphological detail to allow for diagnosis on the species level. Its elongate tibia has been strongly correlated with cursorial habits. The tibia of ''Albertadromeus'' is slightly more elongate than that of ''

Orodromeus ''Orodromeus'' (meaning "Mountain Runner") is a genus of herbivorous orodromine thescelosaurid dinosaur from the Late Cretaceous of North America. Only one species is known, the type species ''Orodromeus makelai''. Discovery and naming The rem ...

'', and significantly more elongate than that of ''

Parksosaurus ''Parksosaurus'' (meaning " William Parks's lizard") is a genus of neornithischian dinosaur from the early Maastrichtian-age Upper Cretaceous Horseshoe Canyon Formation of Alberta, Canada. It is based on most of a partially articulated skeleton a ...

'' (ROM 804), ''

Thescelosaurus ''Thescelosaurus'' ( ; ancient Greek - (''-'') meaning "godlike", "marvellous", or "wondrous" and (') "lizard") was a genus of small neornithischian dinosaur that appeared at the very end of the Late Cretaceous period in North America. It was a ...

'' (USNM 7757 and RSM P 1225.1), ''

Hypsilophodon ''Hypsilophodon'' (; meaning "''Hypsilophus''-tooth") is a neornithischian dinosaur genus from the Early Cretaceous period of England. It has traditionally been considered an early member of the group Ornithopoda, but recent research has put this ...

'' (NMHUK R5830), ''

Dryosaurus ''Dryosaurus'' ( , meaning 'tree lizard', Greek ' () meaning 'tree, oak' and () meaning 'lizard'; the name reflects the forested habitat, not a vague oak-leaf shape of its cheek teeth as is sometimes assumed) is a genus of an ornithopod dinosaur ...

'' (YPM 1876), and ''

Stegoceras ''Stegoceras'' is a genus of pachycephalosaurid (dome-headed) dinosaur that lived in what is now North America during the Late Cretaceous period, about 77.5 to 74 million years ago (mya). The first specimens from Alberta, Canada, were describe ...

'' (UALVP 002). ''Albertadromeus'' was a small-bodied, bipedal, cursorially adapted (built to run) neornithischian.

Diagnosis

According to Brown et al. (2013) ''Albertadromeus'' can be distinguished based on the following characteristics: * The distal fibula is reduced to a thin sheet of bone that is fused to the anterior surface of tibia for the distal one-third of its length (shared with heterodontosaurids) * The lateral condyle of proximal tibia is strongly bilobed (shared with ''Changchunsaurus'', ''Eocursor'', ''Hypsilophodon'', ''Jeholosaurus'', ''Lesothosaurus'', ''Orodromeus'', ''Oryctodromeus'') * A prominent

cnemial crest The cnemial crest is a crestlike prominence located at the front side of the head of the tibiotarsus or tibia in the legs of many mammals and reptiles (including birds and other dinosaurs). The main extensor muscle of the thigh In human anatomy ...

that projects dorsally (and anterolaterally), resulting in a posterior sloped dorsal extremity of the tibia (also present in ''Gasparinisaura'' and ''Micropachycephalosaurus'').

Classification

Brown et al. (2013) erected a new taxon ''Orodrominae'' to differentiate those species that are more closely related to ''

'' from those more closely related to ''

''. The validity of this new taxon is supported by the following: (a) the

foramen In anatomy and osteology, a foramen (;Entry "foramen"
in

magnum is between 20% and 30% of the width of the

occipital condyle The occipital condyles are undersurface protuberances of the occipital bone in vertebrates, which function in articulation with the superior facets of the atlas vertebra. The condyles are oval or reniform (kidney-shaped) in shape, and their anteri ...

; (b) the pubis is articulated with a sacral rib; (c) there is a sharp and pronounced scapular spine; and (d) the fibular shaft is 'D'-shaped in cross-section. The new taxon ''Orodrominae'' includes the newly discovered dinosaur ''Albertadromeus'', ''

Zephyrosaurus ''Zephyrosaurus'' (meaning "westward wind lizard") is a genus of orodromine ornithischian dinosaur. It is based on a partial skull and postcranial fragments discovered in the Aptian-Albian-age Lower Cretaceous Cloverly Formation of Carbon County ...

'', ''Orodromeus'' and ''

Oryctodromeus ''Oryctodromeus'' (meaning "digging runner") was a genus of small orodromine thescelosaurid dinosaur. Fossils are known from the Late Cretaceous Blackleaf Formation of southwestern Montana and the Wayan Formation of southeastern Idaho, USA, bot ...

''. Phylogenetic analyses suggest that ''Albertadromeus'' forms a clade with ''Zephyrosaurus'' and ''Orodromeus'', with ''Oryctodromeus'' being less closely related. The following is a

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

based on the phylogenetic analysis conducted by Brown et al. in 2013, showing the relationships of ''Albertadromeus'':

Paleoecology

The remains of the type specimen of ''Albertadromeus'' was recovered in the Canal Creek locality in the Upper

, which is part of the

Belly River Group The Belly River Group is a stratigraphical unit of Late Cretaceous age in the Western Canadian Sedimentary Basin. It takes the name from the Belly River, a tributary of the Oldman River in southern Alberta, and was first described in outcrop o ...

Alberta, Canada Alberta ( ) is one of the thirteen provinces and territories of Canada, provinces and territories of Canada. It is part of Western Canada and is one of the three Canadian Prairies, prairie provinces. Alberta is bordered by British Columbia to t ...

. The specimen was collected in a two-meter-thick fine to very fine sandstone unit that was deposited during the

stage of the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...

period, approximately 77 to 76 million years ago. This specimen is housed in the collection of the

Royal Tyrrell Museum of Palaeontology The Royal Tyrrell Museum of Palaeontology (RTMP, and often referred to as the Royal Tyrrell Museum) is a palaeontology museum and research facility in Drumheller, Alberta, Canada. The museum was named in honour of Joseph Burr Tyrrell, and is situ ...

Drumheller Drumheller is a town on the Red Deer River in the badlands of Central Alberta, east-central Alberta, Canada. It is northeast of Calgary and south of Stettler, Alberta, Stettler. The Drumheller portion of the Red Deer River valley, often ref ...

. Studies suggest that the

paleoenvironment Paleoecology (also spelled palaeoecology) is the study of interactions between organisms and/or interactions between organisms and their environments across geologic timescales. As a discipline, paleoecology interacts with, depends on and informs ...

of the

was an ancient

coastal plain A coastal plain is flat, low-lying land adjacent to a sea coast. A fall line commonly marks the border between a coastal plain and a piedmont area. Some of the largest coastal plains are in Alaska and the southeastern United States. The Gulf Coa ...

.Eberth, D.A., and Hamblin, A.P., 1993 , Tectonic, stratigraphic, and sedimentologic significance of a regional discontinuity in the upper Judith River Group (Belly River wedge) of southern Alberta, Saskatchewan, and northern Montana: Canadian Journal of Earth Sciences, 30, 174‒200. This formation has produced the remains of the theropods ''

Saurornitholestes ''Saurornitholestes'' ("lizard-bird thief") is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States (Montana, New Mexico, Alabama, North Carolina, and South Carolina). Two spec ...

'', ''

Daspletosaurus ''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three species ...

'', ''

Troodon ''Troodon'' ( ; ''Troödon'' in older sources) is a wastebasket taxon and a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Late Cretaceous period (about 77 mya). It includes at least ...

'', and ''

Dromaeosaurus ''Dromaeosaurus'' (, "running lizard") is a genus of dromaeosaurid theropod dinosaur which lived during the Late Cretaceous period (middle late Campanian and Maastrichtian), sometime between 80 and 69 million years ago, in Alberta, Canada and the ...

'', the ceratopsids ''

Albertaceratops ''Albertaceratops'' (meaning "Alberta horned face") was a genus of centrosaurine horned dinosaur from the middle Campanian-age Upper Cretaceous Oldman Formation of Alberta, Canada. Description ''Albertaceratops'' is unusual in combining long ...

'', ''

Chasmosaurus ''Chasmosaurus'' ( ) is a genus of ceratopsid dinosaur from the Upper Cretaceous Period of North America. Its name means 'opening lizard', referring to the large openings ( fenestrae) in its frill (Greek ''chasma'' meaning 'opening' or 'hollow' ...

'', ''

Anchiceratops ''Anchiceratops'' ( ) is an extinct genus of chasmosaurine ceratopsid dinosaur that lived approximately 72 to 71 million years ago during the latter part of the Cretaceous Period in what is now Alberta, Canada. ''Anchiceratops'' was a medium-size ...

'', and ''

Coronosaurus ''Coronosaurus'' is a genus of centrosaurine ceratopsian dinosaurs which lived in the Late Cretaceous, in the middle Campanian stage. Its remains, two bone beds, were discovered by Phillip J. Currie in the Oldman Formation of Alberta, Canada, and ...

'', the hadrosaurids ''

Brachylophosaurus ''Brachylophosaurus'' ( or ; meaning "short-crested lizard", Greek ''brachys'' = short + ''lophos'' = crest + ''sauros'' = lizard, referring to its small crest) was a mid-sized member of the hadrosaurid family of dinosaurs. It is known from sev ...

'', ''

Gryposaurus ''Gryposaurus'' (meaning "hooked-nosed (Ancient Greek, Greek ''grypos'') lizard"; sometimes incorrectly translated as "griffin (Latin ''gryphus'') lizard") was a genus of hadrosaur, duckbilled dinosaur that lived about 80 to 75 million years ...

'', ''

Parasaurolophus ''Parasaurolophus'' (; meaning "near crested lizard" in reference to '' Saurolophus)'' is a genus of herbivorous hadrosaurid ornithopod dinosaur that lived in what is now North America and possibly Asia during the Late Cretaceous Period, abou ...

'', ''

Maiasaura ''Maiasaura'' (from the Greek ''μαῖα'', meaning "good mother" and ''σαύρα'', the feminine form of ''saurus'', meaning "reptile") is a large herbivorous saurolophine hadrosaurid ("duck-billed") dinosaur genus that lived in the area curre ...

'', and ''

Corythosaurus ''Corythosaurus'' (; ) is a genus of hadrosaurid "duck-billed" dinosaur from the Upper Cretaceous Period (geology), Period, about 77–75.7 million years ago. It lived in what is now North America. Its name means "helmet lizard", derived fr ...

'', as well as other dinosaurs that shared that their paleoenvironment with ''Albertadromeus''.

References

{{Taxonbar, from=Q12973101 Fossils of Canada Late Cretaceous dinosaurs of North America Ornithischian genera Fossil taxa described in 2013 Oldman fauna Paleontology in Alberta Campanian genus first appearances Campanian genus extinctions