2020 In Archosaur Paleontology
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This article records new
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
s of every kind that are scheduled described during the year 2020, as well as other significant discoveries and events related to
paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
of archosaurs that are scheduled to occur in the year 2020.


General research

* A study aiming to determine how mass properties and body proportions relate to each other and locomotor posture in archosaurs is published online by Bishop ''et al.'' (2020). * A study on the brain growth in the
American alligator The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
and in the
common ostrich The common ostrich (''Struthio camelus''), or simply ostrich, is a species of flightless bird native to certain large areas of Africa and is the largest living bird species. It is one of two extant species of ostriches, the only living members o ...
throughout their
ontogeny Ontogeny (also ontogenesis) is the origination and development of an organism (both physical and psychological, e.g., moral development), usually from the time of fertilization of the egg to adult. The term can also be used to refer to the stu ...
is published by Hu ''et al.'' (2020), who evaluate the implications of their findings for the knowledge of the development of the brain in non-avian dinosaurs. * A study on the evolution of metabolic rates along the bird
stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...
lineage is published by Rezende ''et al.'' (2020). * A review of the anatomy of the respiratory systems and mechanics of breathing in living and fossil archosaurs, evaluating their physiological implications, is published by Brocklehurst ''et al.'' (2020). * A study aiming to determine the relationship between atmospheric O2 and CO2 levels during the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
and the evolution of
skeletal pneumaticity Skeletal pneumaticity is the presence of air spaces within bones. It is generally produced during development by excavation of bone by pneumatic diverticula (air sacs) from an air-filled space, such as the lungs or nasal cavity. Pneumatization is h ...
and respiratory systems in
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
dinosaurs and in
paracrocodylomorphs Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha ...
is published by Hudgins, Uhen & Hinnov (2020). * A study on
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
in the skulls of extant
gharial The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males have a distinct b ...
s, and on its implications for the feasibility of detecting dimorphism in non-avian dinosaurs, is published by Hone ''et al.'' (2020). * A study evaluating the impact of different sampling effects on calculations of mean von Ebner line increment width (the measure of
dentin Dentin () (American English) or dentine ( or ) (British English) ( la, substantia eburnea) is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by ena ...
thickness divided by the mean width between von Ebner lines), tooth formation times and replacement rates in fossil archosaurs, based on data from extant
American alligator The American alligator (''Alligator mississippiensis''), sometimes referred to colloquially as a gator or common alligator, is a large crocodilian reptile native to the Southeastern United States. It is one of the two extant species in the g ...
, is published by Kosch & Zanno (2020). * A study on the microstructure of teeth of Mesozoic birds and non-avian
paravian Paraves are a widespread group of theropod dinosaurs that originated in the Middle Jurassic period. In addition to the extinct dromaeosaurids, troodontids, anchiornithids, and possibly the scansoriopterygids, the group also contains the aviala ...
theropods is published by Li ''et al.'' (2020), who evaluate the implications of their findings for the knowledge of differences in feeding ecology of early birds and closely related paravians. * A study on the phylogenetic distribution and structural diversity of
medullary bone The medullary cavity (''medulla'', innermost part) is the central cavity of bone shafts where red bone marrow and/or yellow bone marrow (adipose tissue) is stored; hence, the medullary cavity is also known as the marrow cavity. Located in the m ...
in extant birds, reevaluating the criteria proposed to allow the identification of medullary bone in fossils of
avemetatarsalia Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. ...
ns, is published by Canoville,
Schweitzer Schweitzer is a surname. Notable people with the surname include: * Albert Schweitzer, German theologian, musician, physician, and medical missionary, winner of the 1952 Nobel Peace Prize * Anton Schweitzer, opera composer * Brian Schweitzer, forme ...
& Zanno (2020). * An archosaur egg of uncertain affinities, with eggshell containing several parallel dark bands, is reported from the Upper Cretaceous of
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
by Choi ''et al.'' (2020), who investigate the origin of the dark bands, and name a new
ootaxon Egg fossils are the fossilized remains of eggs laid by ancient animals. As evidence of the physiological processes of an animal, egg fossils are considered a type of trace fossil. Under rare circumstances a fossil egg may preserve the remains of ...
'' Aenigmaoolithus vesicularis''. * A study on the relationship between the curvatures of
ungual An ungual (from Latin ''unguis'', i.e. ''nail'') is a highly modified distal toe bone which ends in a hoof, claw, or nail. Elephants and ungulates have ungual phalanges, as did the sauropods and horned dinosaurs. A claw is a highly modified ungual ...
bones and behaviour in extant birds and
squamates Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest order of reptiles, comprising lizards, snakes, and amphisbaenians (worm lizards), which are collectively known as squamates or scaled reptiles. With over 10,900 species, i ...
, evaluating its implications for the knowledge of the lifestyle of Mesozoic birds and non-avian theropods, is published by Cobb & Sellers (2020). * Xing, Cockx & McKellar (2020) describe a large sample set of 150 specimens of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
containing feathers most likely belonging to non-avian dinosaurs and
enantiornithean The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
birds.


Pseudosuchians


Research

* A study on the skeletal anatomy and bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of ''
Gracilisuchus ''Gracilisuchus'' (meaning "slender crocodile") is an extinct genus of tiny pseudosuchian (a group which includes the ancestors of crocodilians) from the Late Triassic of Argentina. It contains a single species, ''G. stipanicicorum'', which is pl ...
stipanicicorum'', based on data from two new specimens, is published by Lecuona, Desojo & Cerda (2020). * Redescription of the anatomy of the postcranial skeleton of ''
Riojasuchus ''Riojasuchus'' is an extinct genus of Late Triassic ( Norian) quadrupedal archosaur. ''Riojasuchus'' is a member of Ornithosuchidae, a family of facultatively bipedal carnivores that were geographically widespread during the Late Triassic. Tw ...
tenuisceps'', and a study on the phylogenetic affinities of
ornithosuchids Ornithosuchidae is an extinct family of pseudosuchian archosaurs (distant relatives of modern crocodilians) from the Triassic period. Ornithosuchids were quadrupedal and facultatively bipedal (e.g. like chimpanzees), meaning that they had the a ...
, is published by von Baczko, Desojo & Ponce (2020). * Description of a new
erpetosuchid Erpetosuchidae is an extinct family of pseudosuchian archosaurs. Erpetosuchidae was named by D. M. S. Watson in 1917 to include ''Erpetosuchus''. It includes the type species ''Erpetosuchus granti'' from the Late Triassic of Scotland, ''Erpetosu ...
specimen from the Upper Triassic
Lossiemouth Sandstone The Lossiemouth Sandstone is a Middle to Late Triassic (Ladinian to Norian) age geological formation. It is exposed on the south side of the Moray Firth near Lossiemouth and near Golspie in Sutherland. Dinosaur remains are among the fossils th ...
(
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
,
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
) and a review of the anatomy, taxonomy and systematics of other erpetosuchid specimens from the Lossiemouth Sandstone (all previously referred to '' Erpetosuchus'') is published by Foffa ''et al.'' (2020). * Description of new fossil material of ''
Acaenasuchus ''Acaenasuchus'' (meaning "thorn crocodile") is an extinct genus of pseudosuchians, endemic to what would be Arizona during the Late Triassic, existing for approximately . Discovery and naming The holotype, UCMP 139576 (a left(?) dorsal paramedi ...
geoffreyi'' and a study on the phylogenetic relationships of this species is published by Marsh ''et al.'' (2020). * A three-dimensional reconstruction of the armour plates around the tail of ''
Stagonolepis ''Stagonolepis'' is an extinct genus of stagonolepidid aetosaur known from the Late Triassic (Carnian stage) Hassberge Formation of Germany, the Drawno Beds of Poland, and the Lossiemouth Sandstone of Scotland. Supposed fossils from North and S ...
robertsoni'' is presented by Keeble & Benton (2020). * Taxonomic revision, anatomical description, and a study on the phylogenetic relationships of the type and referred materials of ''
Prestosuchus ''Prestosuchus'' (meaning "Prestes crocodile") is an extinct genus of pseudosuchian in the group Loricata, which also includes '' Saurosuchus'' and '' Postosuchus''. It has historically been referred to as a " rauisuchian", and was the defining ...
'' from the original collections of Friedrich von Huene is published by Desojo, von Baczko & Rauhut (2020), who transfer the species ''
Stagonosuchus ''Stagonosuchus'' is an extinct genus of loricatan, or possibly a species of ''Prestosuchus''. Fossils have been found from the Late Triassic Manda Formation in Tanzania that are Anisian in age. Unlike other rauisuchians, which have a prominen ...
nyassicus'' to the genus ''Prestosuchus''. * A study on the skeletal anatomy and phylogenetic relationships of ''
Heptasuchus ''Heptasuchus'' is an extinct genus of loricatan pseudosuchian known from the Middle or Late Triassic upper Chugwater Group of Wyoming, United States. It contains a single species, ''Heptasuchus clarki'', the first formally recognized "rauisuch ...
clarki'' is published by Nesbitt, Zawiskie & Dawley (2020). * A review of the
crocodylomorph Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cro ...
Triassic record in South America is published by Leardi, Yáñez & Pol (2020), who report the occurrence of a large-bodied crocodylomorph in the
Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation da ...
and a putative new non-
crocodyliform Crocodyliformes is a clade of crurotarsan archosaurs, the group often traditionally referred to as "crocodilians". They are the first members of Crocodylomorpha to possess many of the features that define later relatives. They are the only pseudo ...
crocodylomorph taxon from
Los Colorados Formation The Los Colorados Formation is a sedimentary rock formation of the Ischigualasto-Villa Unión Basin, found in the provinces of San Juan and La Rioja in Argentina. The formation dates back to the Norian age of the Late Triassic. The up to thick f ...
(
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
). * A study on the anatomy of the braincase of ''
Almadasuchus ''Almadasuchus'' is an extinct genus of crocodylomorph known from the early Middle Jurassic (early Bajocian stage) Puesto Almada Member of the Cañadón Asfalto Formation of Patagonia, Argentina. It contains a single species, ''Almadasuchus fi ...
figarii'', and on early evolution of cranial pneumaticity in Crocodylomorpha, is published by Leardi, Pol & Clark (2020). * A study on the impact of the habitat on the evolution of body size in Crocodyliformes, based on data from extant and fossil taxa, is published by Gearty & Payne (2020). * New fossil material of crocodylomorphs from the
Birket Qarun Formation Birket is a small town in Lolland Municipality, Denmark. It had a population of 214 residents in 2014. It is located 8 km east of Horslunde, 17 km northeast of Nakskov, 10 km northwest of Stokkemarke and 21 km northwest of Maribo Maribo ...
in the
Fayum Depression The Faiyum Oasis ( ar, واحة الفيوم ''Waḥet El Fayyum'') is a depression or basin in the desert immediately to the west of the Nile, or just 62 miles south of Cairo in Egypt. The extent of the basin area is estimated at between 1,270 ...
(
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
), including the first record of a
sebecosuchia Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in th ...
n from the late
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of Africa, is described by Stefanic ''et al.'' (2020). * A study on the anatomy of the skull and on the phylogenetic relationships of ''
Araripesuchus ''Araripesuchus'' is a genus of extinct crocodyliform that existed during the Cretaceous period of the late Mesozoic era some 125 to 66 million years ago. Six species of ''Araripesuchus'' are currently known. They are generally considered to be n ...
buitreraensis'', based on data from new as well as previously reported specimens, is published by Fernandez Dumont ''et al.'' (2020). * A study on changes in the inner ear vestibular system, involved in sensing balance and equilibrium, throughout the evolutionary history of
thalattosuchia Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Cro ...
ns is published by Schwab ''et al.'' (2020). * A revision of the genus ''
Steneosaurus ''Steneosaurus'' (from el, στενός , 'narrow' and el, σαῦρος , 'lizard') is a dubious genus of teleosaurid crocodyliform from the Middle or Late Jurassic ( Callovian or early Oxfordian) of France. The genus has been used as a w ...
'' is published by Johnson, Young & Brusatte (2020), who designate ''S. rostromajor'' as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of this genus, consider ''S. rostromajor'' to be a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'' and propose that the genus ''Steneosaurus'' is undiagnostic. * Description of new fossil material of ''
Teleidosaurus ''Teleidosaurus'' is an extinct genus of carnivorous metriorhynchoid crocodyliform from Middle Jurassic (late Bajocian to early Bathonian stage) deposits of Normandy, France.Wilkinson LE, Young MT, Benton MJ. 2008. A new metriorhynchid crocodilia ...
calvadosii'' from the middle
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
of Ecouché (
Normandy Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
) and a redescription of the anatomy of this species is published by Hua (2020). * A study on the thermophysiology of metriorhynchids, as indicated by the oxygen isotope composition of the
tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...
phosphate, is published by Séon ''et al.'' (2020). * Fossil material of two large-bodied metriorhynchids is reported from lower
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
sediments in
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total lan ...
and
Baden-Württemberg Baden-Württemberg (; ), commonly shortened to BW or BaWü, is a German state () in Southwest Germany, east of the Rhine, which forms the southern part of Germany's western border with France. With more than 11.07 million inhabitants across a ...
(
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Abel, Sachs & Young (2020), who interpret these fossils as evidence of a new lineage of large-bodied geosaurines from the Kimmeridgian and
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
of Europe. * Redescription of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''
Enaliosuchus ''Enaliosuchus'' is a dubious genus of extinct marine crocodyliform within the family Metriorhynchidae that lived during the Valanginian stage of the Early Cretaceous. It is known from fossil remains found in France and Germany and it was firs ...
macrospondylus'', a revision of the fossil material assigned to this species, and a review of the current knowledge of metriorhynchid diversity during the Cretaceous is published by Sachs, Young & Hornung (2020). * A study aiming to determine whether
notosuchia Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group (see below); if Sebecosu ...
ns were warm-blooded, based on data from bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
, is published by Cubo ''et al.'' (2020), who interpret their findings as indicating that notosuchians were likely to be
ectotherm An ectotherm (from the Greek () "outside" and () "heat") is an organism in which internal physiological sources of heat are of relatively small or of quite negligible importance in controlling body temperature.Davenport, John. Animal Life a ...
s. * A study on the diversity of notosuchians, aiming to determine which factors are potentially distorting the interpretations of the diversity of this group, is published by de Celis ''et al.'' (2020). * A study on the anatomy and biomechanics of baurusuchid skulls, evaluating their implications for the knowledge of likely predatory behaviors of baurusuchids, is published by Montefeltro ''et al.'' (2020). * New information on the anatomy of the endocranial cavities of ''
Campinasuchus ''Campinasuchus'' is an extinct genus of baurusuchid mesoeucrocodylian from Minas Gerais State of Brazil. Description ''Campinasuchus'', like other baurusuchids, was a fully terrestrial predator. It had a deep and laterally compressed skull wi ...
dinizi'' is presented by Fonseca ''et al.'' (2020). * A study on the anatomy of the brain and inner ear of ''
Baurusuchus ''Baurusuchus'' is an extinct member of the ancestral crocodilian lineage, which lived in Brazil from 90 to 83.5 million years ago, in the Late Cretaceous period. Technically, it is a genus of baurusuchid mesoeucrocodylian. It was a terrest ...
'', based on data from reconstructed
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible sp ...
s, is published by Dumont ''et al.'' (2020). * Pholidosaurid fossil material, representing the most recent record of this group reported so far, is described from the
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
(
Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretaceous ...
) of
Ouled Abdoun Basin The Oulad Abdoun Basin (also known as the Ouled Abdoun Basin or Khouribga Basin) is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and a ...
(
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) by Jouve & Jalil (2020), who also reinterpret ''
Dakotasuchus ''Dakotasuchus'' (meaning "Dakota andstonecrocodile") is a genus of goniopholidid mesoeucrocodylian. Its fossils have been recovered from the Cenomanian-age Upper Cretaceous Dakota Sandstone of Kansas. The type specimen was found in an iron-ce ...
kingi'', ''
Woodbinesuchus ''Woodbinesuchus'' (meaning "Woodbine Formation crocodile") is an extinct genus of goniopholidid mesoeucrocodylian. Its fossils have been recovered from the Cenomanian-age Upper Cretaceous Woodbine Formation of Texas. Crocodyliform fossils are w ...
byersmauricei'' and '' Sabinosuchus coahuilensis'' as pholidosaurids, and study the diversity of tethysuchians from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
to the early
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
. * New specimen of ''
Susisuchus ''Susisuchus'' is an extinct genus of neosuchian mesoeucrocodylian crocodyliform from the Early Cretaceous of Brazil. Fossils have been found from the Nova Olinda Member of the Aptian-age Crato Formation in the Araripe and Lima Campos Basins ...
anatoceps'', displaying a non-
eusuchia Eusuchia is a clade of crocodylomorphs that first appeared in the Early Cretaceous with '' Hylaeochampsa''. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-T extinction. Since the other two clades ...
n type
palate The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly sepa ...
(i.e.
choana The choanae (singular choana), posterior nasal apertures or internal nostrils are two openings found at the back of the nasal passage between the nasal cavity and the throat in tetrapods, including humans and other mammals (as well as crocodilia ...
not entirely bounded by the pterygoids), is described by Montefeltro ''et al.'' (2020), who evaluate the implications of this finding for the knowledge of the anatomy of this taxon and the phylogenetic position of susisuchids. * A study on skull anatomy and phylogenetic relationships of ''
Bernissartia ''Bernissartia'' ('of Bernissart') is an extinct genus of neosuchian crocodyliform that lived in the Early Cretaceous, around 130 million years ago. At only in length, ''Bernissartia'' is one of the smallest crocodyliforms that ever lived. It ...
fagesii'' is published by Martin ''et al.'' (2020). * Reconstruction of the internal cavities of the skull of ''
Agaresuchus ''Agaresuchus'' is an extinct genus of allodaposuchid eusuchian crocodylomorph from the Late Cretaceous (Campanian-Maastrichtian) of Spain. It includes two species, the type species ''Agaresuchus fontisensis'', and ''Agaresuchus subjuniperus'', ...
fontisensis'', including the cavities that contained the brain, nerves and blood vessels, is presented by Serrano‐Martínez ''et al.'' (2020). * A study on the skeletal anatomy and phylogenetic relationships of ''
Eocaiman ''Eocaiman'' is an extinct genus of caiman containing species living from the Early Paleocene to Miocene in what is now Argentina (Salamanca Formation, Salamanca and Sarmiento Formations), Itaboraí Formation of Brazil and Colombia (Honda Group, ...
cavernensis'' is published by Godoy ''et al.'' (2020). * A
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
of the
mylodontid Mylodontidae is a family of extinct South American and North American ground sloths within the suborder Folivora of order Pilosa, living from around 23 million years ago (Mya) to 11,000 years ago. This family is most closely related to another fa ...
sloth ''
Pseudoprepotherium ''Pseudoprepotherium'' is an extinct genus of sloths of the family Mylodontidae. It was widespread across northern South America during the Early to Late Miocene epoch around 21 to 5.3 million years ago. Fossils of the animal have been found in B ...
'' bearing 46 predation tooth marks is described from the Miocene
Pebas Formation The Pebas Formation is a lithostratigraphic unit of Miocene age, found in western Amazonia. The formation extends over , including parts of Brazil, Peru, Ecuador and Colombia.Wesselingh et al., 2006 It is interpreted as representing the deposits ...
(
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
) by Pujos & Salas-Gismondi (2020), who interpret this finding as evidence of predation of a young or sub-adult ''
Purussaurus ''Purussaurus'' is an extinct genus of giant caiman that lived in South America during the Miocene epoch, from the Friasian to the Huayquerian in the SALMA classification. It is known from skull material found in the Brazilian and Peruvian Amazon ...
'' on a mylodontid ground sloth. * New fossil material of ''
Mourasuchus ''Mourasuchus'' is an extinct genus of giant, aberrant caiman from the Miocene of South America. Its skull has been described as duck-like, being broad, flat, and very elongate, superficially resembling '' Stomatosuchus'' from the Late Cretaceo ...
arendsi'' is described from the Miocene
Urumaco Formation The Urumaco Formation is a formation in Venezuela that includes deposits from the Late Miocene. It is the site of several "giant forms": the turtles, crocodiles, sloths and rodents of Urumaco are among the largest of their groups. Location Th ...
(
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
) by Cidade, Rincón & Solórzano (2020), who evaluate the implications of these fossils for the knowledge of the paleobiology of this species. * A study on the shape and biomechanical properties of the
humeri The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a round ...
of mekosuchines and extant Australian crocodiles, and on their implications for the knowledge of the locomotion of mekosuchines, is published by Stein ''et al.'' (2020). * Redescription of the anatomy and a study on the phylogenetic relationships of ''
Crocodylus checchiai ''Crocodylus checchiai'' is an extinct species of crocodile from the Miocene to Pliocene of Libya and Kenya. ''C. checchiai'' was named in 1947 based on a skull from the Sahabi Formation. Remains from the lower Nawata Formation in the Turkana Bas ...
'' is published by Delfino ''et al.'' (2020). * Fossil tracks produced by large crocodylomorphs, possibly moving
bipedally Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' a ...
, are described from the Lower Cretaceous
Jinju Formation The Jinju Formation () is an Early Cretaceous geologic Formation (geology), formation in South Korea.South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
) by Kim ''et al.'' (2020), who name a new ichnotaxon '' Batrachopus grandis''. * A study on the impact of recognition of
cryptic species In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
of extant crocodylians on interpretations of the crocodyliform fossil record is published by Brochu & Sumrall (2020).


New taxa


Non-avialan dinosaurs


Research


General

* A study on the
palynological Palynology is the "study of dust" (from grc-gre, παλύνω, palynō, "strew, sprinkle" and ''-logy'') or of "particles that are strewn". A classic palynologist analyses particulate samples collected from the air, from water, or from deposit ...
record from the Carnian–Norian transition in the western
Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian territo ...
region is published by Klausen, Paterson & Benton (2020), who interpret their findings as indicating that major sea‐level changes across the vast delta plains situated in the northern
Pangaea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...
might have triggered terrestrial turnovers during the Carnian–Norian transition and facilitated the gradual rise of the dinosaurs to ecosystem dominance. * A study comparing and testing for correlation between rates of morphological evolution and extinction at the species level in non-avian dinosaurs is published by Crouch (2020). * A study on the
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
of the Cretaceous
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
n dinosaur fauna is published by Kubo (2020). * A study assessing the accuracy and precision of two major approaches to body mass estimation in non-avian dinosaurs is published by Campione &
Evans Evans may refer to: People *Evans (surname) *List of people with surname Evans Places United States *Evans Island, an island of Alaska *Evans, Colorado *Evans, Georgia *Evans County, Georgia *Evans, New York *Evans Mills, New York *Evans City, ...
(2020). * A study on the relationships between
trabecular bone A trabecula (plural trabeculae, from Latin for "small beam") is a small, often microscopic, tissue element in the form of a small beam, strut or rod that supports or anchors a framework of parts within a body or organ. A trabecula generally has ...
architecture and its mechanical properties in dinosaurs is published by Aguirre ''et al.'' (2020). * A study on small dinosaur tracks from the Lower Jurassic
Portland Formation The Portland Formation is a geological formation in Connecticut and Massachusetts in the northeastern United States.Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York to the west, and Long Island Sound to the south. Its cap ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), aiming to reconstruct the foot motions of the trackmaker, is published by Falkingham, Turner & Gatesy (2020). * A review of the Late Cretaceous dinosaur tracksites of
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
is published by Meyer ''et al.'' (2020), who describe new dinosaur tracksites from the Chuquisaca and Potosi departments, and report parallel trackways of subadult ankylosaurs interpreted as evidence of social behavior amongst these dinosaurs. * A study on the evolutionary history of dinosaur integument, aiming to determine the most likely ancestral integumentary condition in dinosaurs, is published by Campione, Barrett & Evans (2020). * A study aiming to determine dinosaur body temperatures on the basis of data from fossil eggshells, comparing them with paleoenvironmental temperatures, and evaluating their implications for the knowledge of dinosaur thermoregulation, is published by Dawson ''et al.'' (2020). * A study on body temperatures of Late Cretaceous sauropods and theropods from western and central
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
, based on data from fossil eggshells, is published by Laskar ''et al.'' (2020). * Evidence for an originally non-biomineralized, soft-shelled nature of eggs of ''
Mussaurus ''Mussaurus'' (meaning " mouse lizard") is a genus of herbivorous sauropodomorph dinosaur that lived in southern Argentina during the Early Jurassic, with a maximum age of 192.78 ± 0.14 Ma. It receives its name from the small size of the sk ...
'' and ''
Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
'' is presented by
Norell Norell is a surname. Notable people with the surname include: * Arne Norell (1917–1971), Swedish furniture designer *Ingeborg Norell (born 1727), the first Finnish female to have received an official decoration * Mark Norell (born 1957), American ...
''et al.'' (2020), who argue that the first dinosaur egg was soft-shelled, and that the calcified, hard-shelled dinosaur egg evolved independently at least three times throughout the Mesozoic era; their interpretation of a soft ''Mussaurus'' eggshell is subsequently contested by Choi ''et al.'' (2022). * A study on the trace elements and isotopic compositions of eggshells of dinosaur eggs from the Cretaceous Zhaoying Formation (
Henan Henan (; or ; ; alternatively Honan) is a landlocked province of China, in the central part of the country. Henan is often referred to as Zhongyuan or Zhongzhou (), which literally means "central plain" or "midland", although the name is al ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), evaluating their implications for reconstructions of local paleoenvironment, is published by He ''et al.'' (2020). * A study on the affinities of putative
gekkota Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates throughout the world. They range from . Geckos ar ...
n eggshells from the Late Cretaceous of Europe is published by Choi ''et al.'' (2020), who interpret the fossil material of '' Pseudogeckoolithus'' as theropod eggshells. * Remains of small theropod eggs, providing new information on the diversity of small dinosaurs in the Hyogo region (
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
), are reported from the Cretaceous (
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
) of the Kamitaki Egg Quarry ( Ohyamashimo Formation) by Tanaka ''et al.'' (2020), who name new ootaxa '' Himeoolithus murakamii'' (the smallest non-avian theropod egg known to date), '' Nipponoolithus ramosus'' and ''
Subtiliolithus ''Subtiliolithus'' is an oogenus of fossil egg from the Nemegt Formation of Mongolia and the Ohyamashimo Formation of Japan. The eggs are notable for a very thin eggshell. It contains three oospecies: ''S. hyogoensis'', ''S. kachchhensis'' and ' ...
hyogoensis''. * Chapelle, Fernandez & Choiniere (2020) evaluate the possibility of estimating the developmental stage of dinosaur embryos, on the basis of a study of skull ossification sequences in embryos of ''
Massospondylus ''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
carinatus'' and extant
sauria Sauria is the clade containing the most recent common ancestor of archosaurs (such as crocodilians, dinosaurs, etc.) and lepidosaurs ( lizards and kin), and all its descendants. Since most molecular phylogenies recover turtles as more closely re ...
ns. * Fossil remains of a member or a relative of the genus ''
Scelidosaurus ''Scelidosaurus'' (; with the intended meaning of "limb lizard", from Greek / meaning 'rib of beef' and ''sauros''/ meaning 'lizard')Liddell & Scott (1980). Greek-English Lexicon, Abridged Edition. Oxford University Press, Oxford, UK. is a gen ...
'' and of an indeterminate
neotheropod Neotheropoda (meaning "new theropods") is a clade that includes coelophysoids and more advanced theropod dinosaurs, and is the only group of theropods that survived the Triassic–Jurassic extinction event. All neotheropods became extinct by the e ...
are described from the Lower Jurassic
Lias Group The Lias Group or Lias is a lithostratigraphic unit (a sequence of rock strata) found in a large area of western Europe, including the British Isles, the North Sea, the Low Countries and the north of Germany. It consists of marine limestones, sh ...
(
Northern Ireland Northern Ireland ( ga, Tuaisceart Éireann ; sco, label= Ulster-Scots, Norlin Airlann) is a part of the United Kingdom, situated in the north-east of the island of Ireland, that is variously described as a country, province or region. Nort ...
) by Simms ''et al.'' (2020), representing the first non-avian dinosaur remains reported from
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
. * Prasad & Parmar (2020) describe fossil teeth of
ornithischia Ornithischia () is an extinct order of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek s ...
n and theropod dinosaurs (including five
morphotype In biology, polymorphism is the occurrence of two or more clearly different morphs or forms, also referred to as alternative ''phenotypes'', in the population of a species. To be classified as such, morphs must occupy the same habitat at the s ...
s of putative
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
teeth) from the
Middle Jurassic The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations co ...
Kota Formation The Kota Formation is a geological formation in India. The precise age of Kota Formation are uncertain, but it dates from the Early to Middle Jurassic, and is split into a Lower Member and Upper Member.Prasad GVR, and Manhas BK. 2007A new docodont ...
, providing new information on the Jurassic dinosaur fauna of
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
. * Two
sacral vertebra The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
e representing the oldest record of fusion of these vertebrae among dinosaurs are described from the Upper Triassic Candelária Sequence (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Moro ''et al.'' (2020), who also review the occurrence of sacral fusion in dinosaurs and their close relatives. *A study aiming to test whether non-avian dinosaurs were in long-term decline prior to the
Cretaceous–Paleogene extinction event The Cretaceous–Paleogene (K–Pg) extinction event (also known as the Cretaceous–Tertiary extinction) was a sudden mass extinction of three-quarters of the plant and animal species on Earth, approximately 66 million years ago. With the ...
is published by Bonsor ''et al.'' (2020); the study is subsequently criticized by Sakamoto, Benton & Venditti (2021). * A study on the causes of extinction of non-avian dinosaurs at the end of the Cretaceous, evaluating dinosaur habitability in the wake of climatic perturbations caused by various asteroid impact and Deccan volcanism scenarios, is published by Chiarenza ''et al.'' (2020).


Saurischians

* A study on the skeletal anatomy and phylogenetic relationships of ''Daemonosaurus chauliodus'' is published by Nesbitt & Hans-Dieter Sues, Sues (2020). * A study on the evolutionary trends and functional relationships between giant body size and hip anatomy in saurischians is published by Tsai ''et al.'' (2020). * A study on the metabolism of ''Coelophysis'' and ''Plateosaurus'', aiming to determine whether the absence of large sauropodomorph dinosaurs in the tropical to subtropical latitudes during the Late Triassic (e.g. the Chinle Formation) was caused by physiological limitations, is published by Lovelace ''et al.'' (2020). * A study on locomotion in non-avian theropods, aiming to determine the selective pressures that influenced evolution of limb length and proportions of limb components in theropods, is published by Dececchi ''et al.'' (2020). * A study on the growth strategies of theropod dinosaurs, with a focus on gigantic Tyrannosauridae, tyrannosaurids and Carcharodontosauridae, carcharodontosaurids, is published by Cullen ''et al.'' (2020). * The discovery of sternal plates of ''Tawa hallae'' from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
of New Mexico and Arizona, representing the oldest known dinosaur sternal plates described so far, is reported by Bradley ''et al.'' (2020), who note the presence of morphological features similar to sternal traits in Avialae, avialans. * A study on the anatomy and phylogenetic relationships of ''Dilophosaurus wetherilli'', based on data from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, referred, and previously undescribed specimens from the Kayenta Formation, is published by Marsh & Rowe (2020). * Redescription of the anatomy, revision of the taxonomy and a study on the phylogenetic relationships of the genus ''Sarcosaurus'' is published by Martín Ezcurra, Ezcurra ''et al.'' (2020). * New fossil material of theropod dinosaurs representing a wide taxonomic range is reported from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
of the Langenberg Quarry (Lower Saxony,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Evers & Wings (2020), who interpret these fossils as evidence of the presence of several taxa of theropods in the Late Jurassic archipelago in the area of Central Europe. * Teeth attributed to the genus ''Ceratosaurus'' are described from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
Tacuarembó Formation (Uruguay) by Matías ''et al.'', (2020). * A vertebra of an Elaphrosaurinae, elaphrosaurine theropod is described from the Lower Cretaceous (
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
) Eumeralla Formation (Victoria (Australia), Victoria, Australia) by Poropat ''et al.'' (2020), representing the first record of Elaphrosaurinae from Australia reported so far. * New theropod fossil material is reported from the Griman Creek Formation by Brougham, Smith & Bell (2020), who interpret it as evidence of the presence of Noasauridae, noasaurids in Australia during the Cretaceous. * A study on the bone microstructure and growth dynamics of ''Vespersaurus paranaensis'' is published by Souza ''et al.'' (2020). * A study on a row of large foramina on the external surface of the skull of ''Skorpiovenator bustingorryi'' is published by Cerroni ''et al.'' (2020), who report evidence indicating that these foramina were linked to an internal canal that ran across the nasal bones, which they interpret as indicative of the presence of blood vessels and nerves, and attempt to determine possible biological significance of this neurovascular system. * A study on the anatomy of the skull of ''Carnotaurus sastrei'' is published by Cerroni, Canale & Fernando Novas, Novas (2020). * Almost complete skeleton of ''Majungasaurus crenatissimus'' preserving evidence of multiple pre-mortem pathologies is described from the Upper Cretaceous Maevarano Formation (Madagascar) by Gutherz ''et al.'' (2020), who interpret these pathologies as most likely to be the result of multiple non-fatal events experienced during the life of the individual, rather than a single traumatic incident. * Hornung (2020) interprets the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''"Ornithocheirus" hilsensis'' as a partial Phalanx bone, phalanx of a large-sized theropod, making it one of the earliest dinosaur discoveries in
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
and one of the few records of large-sized theropods near the Valanginian/Hauterivian boundary of Central Europe. * Pereira ''et al.'' (2020) describe theropod fossil material from the
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
-Cenomanian Açu Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), and evaluate the diversity of theropods from this formation. * Fragmentary maxilla of a member of the genus ''Torvosaurus'' is described from the Middle Jurassic (Callovian) Ornatenton Formation (
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Rauhut ''et al.'' (2020), representing the first occurrence of this genus from Germany and the oldest record of ''Torvosaurus'' reported so far. * A study on the formation time and replacement rates of Spinosauridae, spinosaurid teeth from the Kem Kem Group (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
), comparing them to those of other archosaurs and evaluating their palaeoecological implications, is published by Heckeberg & Rauhut (2020). * A study on the anatomy of the braincase of ''Irritator challengeri'', and on its implications for the knowledge of the neuroanatomy and ecology of this dinosaur, is published by Schade, Rauhut & Evers (2020). * A study on the anatomy of the tail of ''Spinosaurus aegyptiacus'' is published by Nizar Ibrahim, Ibrahim ''et al.'' (2020), who present evidence of tall neural spines and elongate Chevron (anatomy), chevrons forming a large, flexible fin-like organ, interpreted by the authors as evidence of adaptation to tail-propelled aquatic locomotion. * A study on the taxonomic status of spinosaurs from the Kem Kem Group (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) is published by Smyth, Ibrahim & Martill (2020), who consider ''Oxalaia quilombensis'', ''Spinosaurus maroccanus,'' and ''Sigilmassasaurus brevicollis'' to be Synonym (taxonomy), junior synonyms of ''Spinosaurus aegyptiacus''. * Beevor ''et al.'' (2020) report a new locality near Tarda on the northern margin of the Tafilalt (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) dominated by dental remains of ''Spinosaurus'', and interpret the high abundance of spinosaur teeth compared to remains of terrestrial dinosaurs as evidence supporting the interpretation of ''Spinosaurus'' as an aquatic animal. * A study on the anatomy of teeth of ''Sinraptor dongi'', comparing it with dentition of other theropods and evaluating its implications for the knowledge of the feeding ecology of ''S. dongi'', is published by Hendrickx ''et al.'' (2020). * A study on theropod bite marks on Late Jurassic vertebrate fossils from the Mygatt-Moore Quarry (Colorado,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), the identification of the trace makers and their feeding ecology is published by Drumheller ''et al.'' (2020), who report possible evidence of cannibalism in ''Allosaurus''. * A revision of putative carcharodontosaurid teeth from the Upper Cretaceous Bauru Group (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) is published by Delcourt ''et al.'' (2020), who interpret the studied fossil material as more likely to belong to Abelisauridae, abelisaurid theropods. * A study on an indeterminate megaraptoran specimen from the Winton Formation (Australia) is published by White ''et al.'' (2020), who interpret this finding as evidence of either ontogenetic or intraspecific variation in ''Australovenator'', or the presence of a second megaraptorid taxon in the Winton Formation. * Two partial skeletons of large-bodied megaraptorid theropods, representing the most ancient unquestionable records of Megaraptoridae from South America reported so far, are described from the Upper Cretaceous (Cenomanian–Turonian) Bajo Barreal Formation (
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
) by Matt Lamanna, Lamanna ''et al.'' (2020). * A study on the pneumaticity of the sacrum and tail of ''Aoniraptor libertatem'', and on its implications for the knowledge of the evolution of pneumaticity through Theropoda, is published by Rolando, Marsà & Fernando Novas, Novas (2020). * Pol & Goloboff (2020) present a protocol that identifies unstable taxa that decrease support measures in the phylogenetic analyses, and explore a dataset of coelurosaurian relationships published by Pei ''et al.'' (2020) using this protocol. * A study on the
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
of coelurosaurian theropods is published by Ding ''et al.'' (2020). * A study on the endocranial anatomy of ''Bistahieversor sealeyi'', evaluating its implications for the knowledge of the evolution of the brains and sinuses of Tyrannosauroidea, tyrannosauroids, is published by McKeown ''et al.'' (2020). * A metatarsal bone of a young tyrannosaurid theropod, assigned to a very small juvenile ''Gorgosaurus'' is described from the Campanian Dinosaur Park Formation (Alberta, Canada) by Yun (2020). * A frontal bone of a subadult ''Daspletosaurus torosus'' is described from the Campanian Dinosaur Park Formation (Alberta, Canada) by Yun (2020). * A study on the proposed autapomorphies of ''Dynamoterror dynastes'' is published by Yun (2020), who determined a taxonomic name to be a ''nomen dubium''. * A study on the bone microstructure of two half-grown specimens of ''Tyrannosaurus rex'', evaluating its implications for the knowledge of the early life history of members of this species and the taxonomic validity of ''Nanotyrannus lancensis'', is published by Woodward ''et al.'' (2020). * A study on changes in skeleton of ''Tyrannosaurus rex'' during its growth, aiming to assign known specimens of this taxon to specific growth categories, is published by Thomas Carr (paleontologist), Carr (2020). * A study on the pathologies observed in the caudal vertebrae and left fibula of the ''Tyrannosaurus rex'' specimen Field Museum of Natural History, FMNH PR2081 ("Sue (dinosaur), Sue") is published by Hamm ''et al.'' (2020), who diagnose this specimen as affected by osteomyelitis. * A study on the anatomy of the integumentary structures of ''Juravenator starki'' and ''Sciurumimus albersdoerferi'' from the Kimmeridgian Torleite Formation of southern Germany is published by Foth ''et al.'' (2020). * A unique scale type with distinctive circular nodes, interpreted as integumentary sense organs analogous to those in modern crocodylians, is reported from the tail of ''Juravenator starki'' by Bell & Hendrickx (2020). * A study on diversity and possible functions of the Epidermis, epidermal covering of ''Juravenator starki'' is published online by Bell & Hendrickx (2020). * A Ubirajara jubatus, compsognathid specimen preserved with elaborate integumentary structures was described from the Lower Cretaceous Crato Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
); the announcement of its discovery sparked a legal and ethical controversy regarding the circumstances of the fossil's export from Brazil, and the publication describing the specimen was subsequently withdrawn. * A study on the pneumatic chambers in the vertebrae of ''Nothronychus mckinleyi'' is published by Smith, Sanders & Wolfe (2020). * A review of the research on the phylogenetic relationships, morphology and locomotory (including aerial) capabilities of Scansoriopterygidae, scansoriopterygids, and on their implications for the knowledge of the origin of Oviraptorosauria, oviraptorosaurs, is published by Sorkin (2020). * Partial skeleton of an oviraptorosaur theropod closely associated with two eggs (one within the pelvic canal and the other just posterior to it) is described from the Upper Cretaceous Nanxiong Formation (China) by Jin ''et al.'' (2020), who note the complete absence of medullary bone in this egg-bearing specimen. * New fossil material of ''Chirostenotes pergracilis'', representing the first associated Mandible, mandibular and postcranial material of a Caenagnathidae, caenagnathid from the Dinosaur Park Formation (Alberta, Canada), is described by Funston & Philip J. Currie, Currie (2020), who evaluate the implications of these fossils for the knowledge of taxonomy and diversity of caenagnathids from the Dinosaur Park Formation and the growth patterns of ''Chirostenotes pergracilis''. * Description of new caenagnathid fossil material from the Dinosaur Park Formation (Alberta, Canada), providing new information on pelvic anatomy of caenagnathids, is published by Rhodes, Funston & Currie (2020). * Description of a partial skeleton of a caenagnathid theropod from the Upper Cretaceous Hell Creek Formation (Montana,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) and study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of this specimen is published by Cullen ''et al.'' (2020), who evaluate the implications of their findings for the knowledge of the utility of size as a determinant for referral of incomplete or fragmentary skeletal remains to specific or new Coelurosauria, coelurosaur taxa. * An adult Oviraptoridae, oviraptorid specimen preserved atop an egg clutch that contains embryonic remains, representing the first such finding among non-avialan dinosaurs, is described by Bi ''et al.'' (2020). * The first probable Deinonychosauria, deinonychosaur (likely Troodontidae, troodontid) tracks from Canada are described from the Campanian Wapiti Formation (Alberta) by Enriquez ''et al.'' (2020). * New theropod teeth, possibly belonging to members of the family Dromaeosauridae and representing the first record of that group from the southern Junggar Basin, are reported from the Upper Jurassic Qigu Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Maisch & Matzke (2020). * A study on the facial Skeletal pneumaticity, pneumatic features of members of the family Dromaeosauridae, and on the evolutionary history of these features, is published by Brownstein (2020). * A study on the differences in the locomotor and predatory specializations of Eudromaeosauria, eudromaeosaurs and Unenlagiinae, unenlagiines, as indicated by the anatomy of their hindlimbs, is published by Gianechini, Ercoli & Díaz‐Martínez (2020). * A study on eudromaeosaurian maxillae, aiming to determine the extent to which maxillae can be used to draw ecological and phylogenetic inferences about dromaeosaurids, is published by Powers, Sullivan & Currie (2020). * Evidence of sequential wing feather molt in a specimen of ''Microraptor'' is presented by Kiat ''et al.'' (2020), who evaluate the implications of this finding for the knowledge of the ecology and locomotion of this theropod. * Partial dentary of a juvenile Saurornitholestinae, saurornitholestine dromaeosaurid is described from the Upper Cretaceous Prince Creek Formation (Alaska, United States) by Chiarenza ''et al.'' (2020), representing the first confirmed non-dental fossil specimen of a member of Dromaeosauridae in the Arctic. * The first cranial material of ''Saurornitholestes'' is described from the Judith River Formation (Montana, United States) by Wilson & Fowler (2020), representing the easternmost occurrence of this genus reported so far. * A study testing for dietary changes through growth in ''Deinonychus antirrhopus'' is published by Frederickson, Engel & Cifelli (2020). * A study on the anatomy of the hindbrain and inner ear of ''Velociraptor mongoliensis'', evaluating its implications for the knowledge of the trophic ecology and sensory aptitude of this theropod, is published by King ''et al.'' (2020). * A study aiming to determine the thermoregulatory efficiency of contact Egg incubation, incubation of partially buried eggs by ''Troodon formosus'' is published by Hogan & Varricchio (2020). * Description of the anatomy of the skeleton of ''Rahonavis ostromi'' is published by Catherine Forster, Forster ''et al.'' (2020). * A study on the flight potential and gliding capabilities of ''Yi (dinosaur), Yi qi'' and ''Ambopteryx longibrachium'' is published by Dececchi ''et al.'' (2020). * A study on the chemical preservation of fossil feathers preserved in association with the skeleton of ''Anchiornis huxleyi'' is published by Cincotta ''et al.'' (2020). * A study on the quality of the Sauropodomorpha, sauropodomorph fossil record is published by Cashmore ''et al.'' (2020). * A study on the anatomy of the endocranial cavity and the probable anatomy of the brain of ''Buriolestes schultzi'' is published by Müller ''et al.'' (2020). * Description of new fossil material of ''Thecodontosaurus antiquus'', providing new information on the skeletal anatomy of this species, is published by Ballell, Emily Rayfield, Rayfield & Benton (2020), who evaluate the implications of these fossils for the knowledge of the paleoecology of ''Thecodontosaurus'' and the taxonomy of Late Triassic British sauropodomorphs. * A study on the anatomy of the braincase of ''Thecodontosaurus antiquus'' is published by Ballell ''et al.'' (2020), who also reconstruct the anatomy of the brain of this dinosaur, and evaluate its implications for the knowledge of the paleobiology of ''Thecodontosaurus''. *Greenfield ''et al.'' (2020) reviewed the nomenclature of ''Coloradisaurus'' and determined that the authorship should be attributed to Peter Galton and not David Lambert. * A study on the morphological variation of ''Plateosaurus'' occurring at the genus level, as indicated by data on the shape variation of a sample of limb long bones, is published by Lefebvre ''et al.'' (2020). * New skeleton of ''Plateosaurus'', representing the first substantially complete specimen of a juvenile ''Plateosaurus'' and the first such specimen with a body size significantly below the known adult size range of this taxon, is described from the Norian Klettgau Formation (Switzerland) by Nau ''et al.'' (2020). * The second-known specimen of ''Ignavusaurus rachelis'', extending known geographic range of this species, is described from the Likhoele Mountain near Mafeteng (Upper Elliot Formation, Lesotho) by Bodenham & Barrett (2020). * A study on teeth development in embryos of ''Lufengosaurus'' is published by Robert R. Reisz, Reisz ''et al.'' (2020). * A study on the
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
of the
humeri The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a round ...
of two Basal (phylogenetics), basal sauropod specimens from the Jurassic of Niger and Thailand, reporting evidence of a layer of the radial fibrolamellar bone buried in the outer cortex of these bones, is published by Jentgen-Ceschino, Stein & Fischer (2020), who interpret their findings as evidence of these sauropods being affected by pathologies similar to Ewing's sarcoma and avian osteopetrosis or haemangioma. * A study comparing articulation and range of motion of necks of extant giraffes and ''Spinophorosaurus nigerensis'' is published by Vidal ''et al.'' (2020). * A study on the body plan, functional morphology of the neck and feeding capabilities of ''Spinophorosaurus nigerensis'' is published by Vidal ''et al.'' (2020). * A study on the skeletal anatomy and phylogenetic relationships of ''Klamelisaurus gobiensis'' is published by Moore ''et al.'' (2020). * Two vertebrae of Diplodocoidea, diplodocoid sauropods are described from the Middle Jurassic (Callovian) Podosinki Formation (Russia) by Averianov & Zverkov (2020), who evaluate the implications of this finding for the knowledge of the initial radiation of Diplodocoidea. * Fossils of a member of Flagellicaudata are described from the Middle Jurassic Otlaltepec Formation (Mexico) by Rivera-Sylva & Espinosa-Arrubarrena (2020), representing the first conclusive evidence of the occurrence of Flagellicaudata in this part of North America throughout the
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
-Callovian. * Baron (2020) argues that the elongate tails of Diplodocidae, diplodocid sauropods were used for herding co-ordination. * A review of the distribution of the Cretaceous fossils of Rebbachisauridae, rebbachisaurid sauropods is published by Pereira ''et al.'' (2020), who report the first occurrence of a rebbachisaurid from the Açu Formation (Potiguar Basin,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
), and discuss its Biogeography#Paleobiogeography, paleobiogeographic implications. * A reconstruction of the Epaxial and hypaxial muscles, epaxial and hypaxial musculature of the tail of ''Giraffatitan brancai'' is published by Díez Díaz ''et al.'' (2020). * A humerus of a Titanosauriformes, titanosauriform sauropod, likely belonging to a member or a relative of the genus ''Duriatitan'', is described from the
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
-Berriasian Rupelo Formation (Province of Burgos, Burgos, Spain) by Torcida Fernández-Baldor, Canudo & Huerta (2020). * A large sauropod humerus, probably belonging to a member of the species ''Fusuisaurus zhaoi'', is described from the Lower Cretaceous Xinlong Formation (Guangxi,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Mo ''et al.'' (2020). * A study on sauropod teeth from the Cenomanian Griman Creek Formation (
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
), evaluating their implications for the knowledge of the diversity and palaeoecology of the sauropods from this formation, is published by Frauenfelder ''et al.'' (2020), who report evidence of the presence of at least two taxa of non-Titanosauria, titanosaur titanosauriforms and a possible titanosaur. * A study on
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
and affinities of two bone fragments from the Upper Cretaceous (lower Santonian to/or lower Campanian) of the Western Srednogorie (Bulgaria) is published by Nikolov ''et al.'' (2020), who interpret these fossils as bones of a titanosaur sauropod, coming from a time interval when sauropods are rare in the fossil record of Europe. * An almost intact embryonic skull of a titanosaur sauropod is described from the Upper Cretaceous Allen Formation (
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
) by Kundrát ''et al.'' (2020), who interpret this specimen as indicating that titanosaurs hatched with a temporary monocerotid (single-horned) face, retracted narial openings, and early binocular vision. * Evidence of aggressive case of osteomyelitis affecting a titanosaur specimen from the Upper Cretaceous Adamantina Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) is reported by Aureliano ''et al.'' (2020), who also report the preservation of tens of parasites throughout the specimen’s vascular canals. * Description of the skeletal anatomy of ''Savannasaurus elliottorum'' is published by Poropat ''et al.'' (2020). * A study on the anatomy of the brain and inner ear of ''Narambuenatitan palomoi'' is published by Paulina-Carabajal, Filippi & Knoll (2020). * Voegele ''et al.'' (2020) reconstruct the forelimb and shoulder girdle musculature of ''Dreadnoughtus schrani'', as well as the pelvic girdle and hindlimb musculature of this sauropod. * A study on the anatomy of the appendicular skeleton of ''Patagotitan mayorum'' is published by Otero, Carballido & Moreno (2020), who also provide a new body mass estimate of this species.


Ornithischians

* A study on the phylogenetic relationships of the silesaurids is published by Müller & Garcia (2020), who recover silesaurids as an evolutionary grade of early ornithischian dinosaurs. * A study on the microstructure of the
tooth enamel Tooth enamel is one of the four major Tissue (biology), tissues that make up the tooth in humans and many other animals, including some species of fish. It makes up the normally visible part of the tooth, covering the Crown (tooth), crown. The ...
of ''Manidens condorensis'', evaluating its implications for the knowledge of the evolution of tooth enamel in Ornithischia, is published by Becerra & Pol (2020). * A study on tooth replacement in ''Manidens condorensis'' is published by Becerra ''et al.'' (2020). * New specimens of ''Scutellosaurus lawleri'', providing new information on the anatomy of this species, are described from the Lower Jurassic Kayenta Formation (Arizona, United States), by Breeden & Rowe (2020). * Studies on the structure and development of the dermal skeleton of ''
Scelidosaurus ''Scelidosaurus'' (; with the intended meaning of "limb lizard", from Greek / meaning 'rib of beef' and ''sauros''/ meaning 'lizard')Liddell & Scott (1980). Greek-English Lexicon, Abridged Edition. Oxford University Press, Oxford, UK. is a gen ...
harrisonii'', the neurocranium and the associated principal sensory systems of this dinosaur, its locomotor abilities, breathing, and on its phylogenetic relationships, are published by David B. Norman, Norman (2020). * Description of the dermal armor of ''Stegosaurus'', revision of the various reconstructions of the dermal armor of ''S. ungulatus'' and ''S. stenops'', and a summary of the evidence for and against the different functions proposed for the plates and spines of ''Stegosaurus'' is published by Peter Galton, Galton (2020). * An isolated caudal vertebra representing the first evidence of the presence of an ankylosaur in the Upper Jurassic Qigu Formation (China) is described by Augustin ''et al.'' (2020). * A study aiming to determine the social lifestyle of ankylosaurs, as indicated by anatomy, taphonomic history, ontogenetic composition of the mass death assemblages and inferred habitat characteristics, is published by Botfalvai, Prondvai & Ősi (2020). * Redescription of the anatomy of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimens of ''Hylaeosaurus armatus'' and ''Polacanthus foxii'', and a study on the taxonomy of all ankylosaur specimens from the British Wealden Group, Wealden Supergroup, is published by Raven ''et al.'' (2020). * Fossil stomach contents preserved within the abdominal cavity of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Borealopelta markmitchelli'' are described by Brown ''et al.'' (2020). * Description of the anatomy of braincases of three specimens of ''Bissektipelta archibaldi'' is published by Kuzmin ''et al.'' (2020). * A study on the phylogenetic relationships of cerapodan ornithischians is published by Dieudonné ''et al.'' (2020). * A study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
and probable life history of ''Jeholosaurus shangyuanensis'' is published by Han ''et al.'' (2020). * A study on the bone histology and growth patterns of ''Trinisaura santamartaensis'' and ''Morrosaurus antarcticus'' is published by Garcia-Marsà ''et al.'' (2020). * Redescription of ''Eucercosaurus tanyspondylus'' and ''Syngonosaurus macrocercus'' from the Cenomanian Cambridge Greensand (
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) is published by Barrett & Bonsor (2020), who interpret both these taxa as described on the basis of fossils of iguanodontian dinosaurs possessing no clear diagnostic features. * A study on the bone microstructure of Mongolian Hadrosauroidea, hadrosauroid dinosaurs, evaluating its implications for the knowledge of growth strategies and evolution of gigantism in hadrosauroids, is published by Słowiak ''et al.'' (2020). * Brownstein (2020) describes new fossil material of Hadrosauromorpha, hadrosauromorphs from the Maastrichtian New Egypt Formation (New Jersey,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), including a skeleton of a specimen which was probably a small-bodied adult hadrosauromorph from a lineage outside Hadrosauridae and fossils of juvenile hadrosauromorphs. * A study on the anatomy of the tail of ''Tethyshadros insularis'' is published by Dalla Vecchia (2020). * A study on pathologies affecting two hadrosaurid vertebrae from the Dinosaur Provincial Park (Alberta, Canada) is published by Rothschild ''et al.'' (2020), who consider Langerhans cell histiocytosis to be the most likely diagnosis, making it the first case of LCH recognized in a dinosaur so far. * A study on a set of fused hadrosaur vertebrae with fragments of a tooth of ''Tyrannosaurus rex'' scattered through the intervertebral space is published by Rothschild ''et al.'' (2020), who interpret this findings as evidence indicating that the space between the vertebrae was not occupied by intervertebral discs, but rather by an articular space similar to that in modern reptiles. * A study on the migratory behaviours of hadrosaurs, as indicated by strontium isotope data from hadrosaur teeth from the Late Cretaceous of Alberta (Canada), is published by Terrill, Henderson & Anderson (2020). * A study aiming to determine whether body size and Ontogeny, ontogenetic age were strongly correlated in hadrosaurid dinosaurs from the Dinosaur Park Formation (Alberta, Canada), and to test the hypothesis of a rapid growth rate of hadrosaurids from the Dinosaur Park Formation relative to those from the Two Medicine Formation, is published by Wosik ''et al.'' (2020). * Partial forelimb of a large hadrosaurid with similarities to forelimbs of Lambeosaurinae, lambeosaurines is described from the Maastrichtian New Egypt Formation (New Jersey, United States) by Brownstein & Bissell (2020), who interpret this findings as evidence of the presence of a morphotype of large hadrosauromorph with elongate forelimbs in the latest Maastrichtian of eastern North America. * A study on the anatomy of fossils of ''Ugrunaaluk kuukpikensis'' and on the taxonomic status of this species is published by Takasaki ''et al.'' (2020), who consider ''Ugrunaaluk'' to be a Synonym (taxonomy), junior synonym of the genus ''Edmontosaurus''. * Evidence of pre-mortem traumatic injuries in multiple skeletal elements (especially in tail vertebrae) of ''Edmontosaurus annectens'' from the Lance Formation (Wyoming, United States) is presented by Siviero ''et al.'' (2020). * A study on the taphonomy and depositional history of an extensive Maastrichtian bonebed in the Lance Formation of eastern Wyoming dominated by fossils of ''Edmontosaurus annectens'' is published by Snyder ''et al.'' (2020). * A study on the interior structure of the nasal spine of ''Tsintaosaurus spinorhinus'' is published by Zhang ''et al.'' (2020). * Description of new fossil material of ''Pararhabdodon isonensis'', and a study on the bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
and life history of this taxon, is published by Serrano ''et al.'' (2020). * A study on the morphology and likely causes of the injuries in the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
specimen of ''Parasaurolophus walkeri'' is published by Bertozzo ''et al.'' (2020). * Evidence of preservation of proteins, chromosomes and chemical markers of DNA in the cartilage of a nestling of ''Hypacrosaurus stebingeri'' from the Campanian Two Medicine Formation (Montana,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) is presented by Bailleul ''et al.'' (2020). * A study on patterns of morphological variation of the ceratopsian frill, and on its implications for the knowledge of the ontogeny and evolution of this structure, is published by Prieto‐Márquez ''et al.'' (2020). * New Protoceratopsidae, protoceratopsid specimens are described from the Üüden Sair and Zamyn Khond localities (Mongolia) by Czepiński (2020), who evaluates the implications of these specimens for correlation of fossil sites of the Djadochta Formation, and interprets one of these specimens as probable evidence of an Anagenesis, anagenetic transition from ''
Protoceratops ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
andrewsi'' to ''Bagaceratops rozhdestvenskyi''. * Evidence of osteosarcoma affecting a specimen of ''Centrosaurus apertus'', representing the first case of osteosarcoma in a dinosaur reported so far, is presented by Ekhtiari ''et al.'' (2020). * Description of an immature specimen of ''Styracosaurus albertensis'' (the smallest known for this species) from the Campanian Dinosaur Park Formation (Alberta, Canada), and a study comparing the ontogeny and individual variation of the skulls in ''Styracosaurus'' and ''Centrosaurus'', is published by Brown, Holmes & Currie (2020). * A study on the braincases of two specimens of ''Triceratops'' is published by Sakagami & Kawabe (2020), who present three-dimensional virtual renderings of the endocasts of the cranial cavities and bony labyrinths, and compare the endocranial endocasts of specimens of ''Triceratops'' and other ceratopsians.


New taxa


Birds


Research

* A study on the phylogenetic relationships and powered flight potential of early birds and their closest relatives is published by Pei ''et al.'' (2020), who argue that the potential for powered flight evolved at least three times (once in birds and twice in Dromaeosauridae, dromaeosaurids). * A study aiming to determine whether the origin of birds was marked with a distinct shift in cranial evolutionary dynamics, based on data from birds and non-avian dinosaurs, is published by Felice ''et al.'' (2020). * A study on patterns of evolution of avian brain size and its relationship with body size evolution, based on data from extant and fossil birds and from non-avian theropod dinosaurs, is published by Ksepka ''et al.'' (2020). * A study aiming to determine the volumes of the brain structures used to infer behavior or functional capabilities in ''Archaeopteryx lithographica'', ''Lithornis plebius'', ''Dinornis robustus'', ''Paraptenodytes antarcticus'', ''Psilopterus lemoinei'', ''Llallawavis scagliai'' and an unnamed Miocene Galliformes, galliform is published by Early, Ridgely & Lawrence Witmer, Witmer (2020). * The study on the identity of the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
feather of ''Archaeopteryx lithographica'' published by Kaye ''et al.'' (2019) is criticized by Carney, Tischlinger & Shawkey (2020). * Evidence of Moulting#In birds, feather moulting in the Specimens of Archaeopteryx#The Thermopolis specimen, Thermopolis specimen of ''Archaeopteryx'' is presented by Kaye, Pittman & Wahl (2020), who evaluate the implications of this finding for the knowledge of the origins of flight-related molting and flight; the study is subsequently criticized by Kiat ''et al.'' (2021). * A study on the structure and possible function of the paddle-shaped skeletal elements preserved in the thoracic region of the skeleton of ''Jeholornis'' is published by Zheng ''et al.'' (2020), who interpret these elements as anomalously expanded sternal ribs. * Traces of the soft tissue of the beak preserved with two specimens of ''Confuciusornis'' are described by Zheng ''et al.'' (2020). * Miller ''et al.'' (2020) describe a new specimen of ''Confuciusornis sanctus'' preserving a disassociated Beak#Rhamphotheca, rhamphotheca, and evaluate the differences in the keratinous and bony beak anatomy between confuciusornithids and modern birds. * A study on the furcula-coracoid articulation in ''Confuciusornis'' and in extant birds that utilize different flight styles is published by Wu ''et al.'' (2020), who report the first evidence of fossilized secondary cartilage on the furcula of a fossil bird, and evaluate the implications of their findings for the knowledge of flight styles in Mesozoic birds. * A study on the anatomy of the skull of ''Sapeornis chaoyangensis'' is published by Hu ''et al.'' (2020). * New specimen of ''Longusunguis kurochkini'', providing new information on the anatomy of this taxon and indicating that the Plesiomorphy and symplesiomorphy, plesiomorphic diapsid skull was retained by at least some Basal (phylogenetics), basal Enantiornithes, enantiornithines, is described from the Lower Cretaceous Jiufotang Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Hu ''et al.'' (2020). * An isolated foot of an enantiornithine consisting of complete Metatarsal bones, metatarsals and Digit (anatomy), digits, including the claws, is described from the Cretaceous
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
by Xing ''et al.'' (2020). * New enantiornithine specimen preserving portions of two forelimbs and two feet, as well as associated feathers, is described from the Cretaceous Burmese amber by Xing ''et al.'' (2020), providing new evidence of a diversity of limb proportions and plumage patterns in the enantiornithine fauna from Myanmar. * Bailleul ''et al.'' (2020) confirm the presence of ovarian follicles in an enantiornithine specimen Shandong Tianyu Museum of Nature, STM10–12 from the Lower Cretaceous of China; the conclusions of this study are subsequently contested by Mayr ''et al.'' (2020), who interpret putative ovarian follicles of this specimen and other birds from the Jehol Biota as more likely to be ingested food items. * New specimen of ''Protopteryx fengningensis'', providing additional information on the plumage of this species, is described by O’Connor ''et al.'' (2020). * Wang & Zhou Zhonghe, Zhou (2020) describe a new specimen of ''Piscivorenantiornis inusitatus'' from the Lower Cretaceous Jiufotang Formation (China), providing new information on the anatomy and phylogenetic relationships of this taxon. * A feather fragment from an aquatic bird is reported from amber recovered from the Pipestone Creek bonebed from the Campanian Wapiti Formation (Alberta, Canada) by Cockx ''et al.'' (2020). * A tibiotarsus of a non-hesperornithid Hesperornithes, hesperornithiform is described from the Upper Cretaceous (Maastrichtian) Kita-ama Formation (
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
) by Tanaka ''et al.'' (2020), representing the first hesperornithiform record from marine Maastrichtian deposits in Asia reported so far, and indicating that the habitat of hesperornithiforms during the Maastrichtian extended to both terrestrial and marine environments in Asia and North America. * A study on the anatomy of the skeleton of ''Parahesperornis alexi'' is published by Bell & Luis M. Chiappe, Chiappe (2020), who report that this taxon possessed a mosaic of Basal (phylogenetics), basal and derived traits found among other hesperornithiform taxa. * A study on melanosome Morphology (biology), morphologies in two Lithornithidae, lithornithid specimens from the Eocene Green River Formation (
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
), evaluating their implications for reconstructions of coloration in lithornithids and for the knowledge of color evolution in Palaeognathae, palaeognaths, is published by Eliason & Julia Clarke, Clarke (2020). * Evidence of the presence of the bill tip organ in lithornithids is presented by du Toit, Anusuya Chinsamy-Turan, Chinsamy & Cunningham (2020), who interpret their findings as indicating that remote-touch probe foraging evolved very early among the Neornithes and may even have predated the palaeognathous–Neognathae, neognathous divergence. * A study on the life history of the elephant birds, as indicated by bone
histology Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures vis ...
, is published by Anusuya Chinsamy-Turan, Chinsamy ''et al.'' (2020). * A study on the evolutionary history of the ostriches in Africa and Eurasia during the Miocene, Pliocene and Pleistocene, as indicated by data from eggshell and bone fossil record, is published by Mikhailov & Zelenkov (2020). * A dentary fragment of a Pelagornithidae, pelagornithid bird with an estimated body size on par with the largest known members of Pelagornithidae is described from the Eocene Submeseta Formation (Seymour Island, Antarctica) by Kloess, Poust & Stidham (2020), who interpret this finding as evidence of early evolution of giant body size in Pelagornithidae. * A study on the flight capacity of ''Pelagornis chilensis'' is published by Meza-Vélez (2020). * Volkova & Zelenkov (2020) describe new fossil material of geese from the late Miocene locality Khyargas Nuur 2 in western Mongolia, and evaluate the implications of these fossils for the knowledge of the late Miocene evolution and paleogeography of geese. * A coracoid of a small-bodied Paraortygidae, paraortygid is reported from the Uinta Formation (Utah,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
) by Stidham, Townsend & Holroyd (2020), representing the only known Pangalliformes, pangalliform from the middle
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
of North America, occurring in a temporal gap in their history between the early Eocene ''Gallinuloides wyomingensis'' and late Eocene ''Nanortyx inexpectus''. * A revision of the fossil material from Ukraine attributed to ''Plioperdix pontica'', and a study on the taxonomy of small Phasianidae, phasianids of the Neogene-Pleistocene in the Northern Black Sea region and Eastern Europe, is published by Zelenkov & Gorobets (2020). * Barton ''et al.'' (2020) reinterpret purported chicken specimens from the Neolithic site at Dadiwan culture, Dadiwan as remains of pheasants, and argue that these remains provide evidence of exploitation of grain-fed pheasants by early farmers in arid northwest
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. * Lawal ''et al.'' (2020) report that chicken was domesticated 8,000 years ago from its primary ancestor, Red junglefowl and that the genome of chicken was subsequently enhanced through introgression with the other three junglefowls i.e. Grey junglefowl, Sri Lankan junglefowl, and Green junglefowl. * A study on the origin and history of domestication of chickens, as indicated by data from domestic chicken and wild jungle fowl genomes, is published by Wang ''et al.'' (2020), who interpret their findings as indicating that domestic chickens were initially derived from the red junglefowl subspecies ''Gallus gallus spadiceus'', and that they interbred locally with other subspecies of the red junglefowl and with other jungle fowl species after their domestication. * Partial skull of a large-bodied Crane (bird), crane with a long beak, representing the oldest record of the subfamily Gruinae reported so far and expanding the known temporal range for the occurrence of large-sized cranes in Europe, is described from the Miocene (Tortonian) locality Hammerschmiede clay pit, Hammerschmiede (
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total lan ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Mayr, Lechner & Böhme (2020). * A study comparing the osteology of Plotopteridae, plotopterids and
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
Crown group#Stem groups, stem group penguins is published by Gerald Mayr, Mayr ''et al.'' (2020). * New fossil material of ''Macranhinga ranzii'' is described from the Miocene Solimões Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Guilherme ''et al.'' (2020), who also study the phylogenetic relationships of this species. * A study on the morphological diversity of bills of extant and fossil penguins, and its relationship with feeding habits, is published by Chávez-Hoffmeister (2020). * Partial skeleton of an early penguin (possibly belonging to the species ''Muriwaimanu tuatahi''), preserving the first complete wing of a
Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...
penguin reported so far and providing new information on the skeletal anatomy of this taxon, is described from the Waipara Greensand (New Zealand) by Mayr ''et al.'' (2020). * An articulated wing of ''Palaeeudyptes gunnari'', preserving mineralized skin, is described from the
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
(Lutetian) of Seymour Island (Antarctica) by Acosta Hospitaleche ''et al.'' (2020). * A study on the swimming capabilities of ''Inkayacu paracasensis'' is published by Meza-Vélez (2020). * New fossil material of ''Anhinga pannonica'' is described from the Miocene (Tortonian) of the Hammerschmiede clay pit (
Bavaria Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total lan ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Mayr, Lechner & Madelaine Böhme, Böhme (2020), who also reinterpret the putative Miocene cormorant ''Phalacrocorax brunhuberi'' as another, previously misclassified, record of ''A. pannonica''. * New fossil material of penguins and a member of Gruiformes is reported from the Eocene La Meseta Formation, La Meseta and Submeseta Formations of the Seymour Island by Davis ''et al.'' (2020), supporting previously controversial reports of Gruiformes from Antarctica. * Partial tibiotarsus of an owl (possibly a member of Selenornithinae) is described from the Oligocene of the Jebel Qatrani Formation (
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
) by Smith, Stidham & Mitchell (2020), representing the first occurrence of a fossil owl from the Paleogene of Africa reported so far. * Part of a maxilla of a member of the genus ''Tockus'' is described from the early Miocene of Napak (Uganda) by Riamon ''et al.'' (2020), representing the earliest fossil of a hornbill reported so far. * A nearly complete passerine specimen is described from the early Oligocene of Revest-des-Brousses (Luberon, Alpes-de-Haute-Provence,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
) by Riamon, Tourment & Louchart (2020), who interpret this specimen as a member of Tyranni, most likely belonging to the Crown group#Stem groups, stem group of Tyrannida. * New fossil material of larks is reported from the late Pliocene localities in Transbaikalia (Russia) and Mongolia by Palastrova & Zelenkov (2020), who transfer the species ''Pliocalcarius orkhonensis'' to the genus ''Eremophila (bird), Eremophila'', and evaluate the implications of their findings for the knowledge of the evolutionary history of larks. * An exceptionally well-preserved bird carcass found in the Siberian permafrost and dated to approximately 44–49 Year#SI prefix multipliers, ka Before Present, BP is described by Dussex ''et al.'' (2020), who identify this specimen as a female horned lark, and evaluate the implications of this specimen for the knowledge of the evolution and biogeography of its species during the Pleistocene. * New fossil material of ''Emberiza shaamarica, Rhodospiza shaamarica'' is described from the Pliocene localities of Shaamar (northern Mongolia) and Beregovaya (south Transbaikalia, Russia) by Palastrova & Zelenkov (2020), who transfer this species to the genus ''Bunting (bird), Emberiza''. * A study on Pleistocene bird tracks from the Cape south coast of South Africa is published by Helm ''et al.'' (2020), who report six tracksites with tracks produced by large birds, possibly indicating the existence of large Pleistocene forms of extant bird taxa. * A study on the impact of the climate changes of the last 35,000 years on small birds from the La Brea Tar Pits is published by Long, Donald Prothero, Prothero & Syverson (2020). * A study comparing predicted breeding and wintering distributions for landbird species identified from the La Brea Tar Pits during the Last Glacial Maximum, aiming to determine if niche models successfully predict species’ presence, to estimate the degree of species turnover, to evaluate the fluidity of life history strategies of birds from La Brea, and to compare niche breadths of bark-foraging birds from La Brea between the Last Glacial Maximum and the present, is published by Zink ''et al.'' (2020). * New fossil material of seabirds, including remains of the little auk or a related species, is reported from the Pleistocene Kazusa and Shimosa groups (
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
) by Watanabe ''et al.'' (2020), who interpret this finding as possible evidence that the little auk more widespread in the North Pacific in the middle Pleistocene than it is today. * A study comparing the bird communities of the Bahamian Archipelago (The Bahamas and Turks and Caicos Islands) during the Late Pleistocene, Holocene prior to human arrival and Late Holocene subsequent to human arrival, aiming to determine what drove changes in biodiversity of Bahamian birds throughout the Late Quaternary, is published by David Steadman, Steadman & Janet Franklin, Franklin (2020). * A study on Flightless bird, flightlessness in bird species known to have gone extinct since the rise of humans (i.e. in the Late Pleistocene and Holocene), aiming to determine the extent to which inferences about evolutionary transitions and rates of evolution of flightlessness in birds are biased by anthropogenic extinctions, is published by Sayol ''et al.'' (2020).


New taxa


Pterosaurs


Research

* A study on the ingroup relationships within Pterosauria is published by Baron (2020), who names new clades Zambellisauria and Caviramidae. * A study on the fusion process of the Notarium, notarial elements of pterosaurs is published by Aires ''et al.'' (2020). * A study on the evolution of the flight efficiency of pterosaurs is published by Venditti ''et al.'' (2020). * A study aiming to determine diets of 17 pterosaur genera, as indicated by data from dental microwear texture analysis, is published by Bestwick ''et al.'' (2020). * Mazin & Pouech (2020) describe non-Pterodactyloidea, pterodactyloid pterosaur tracks from the ichnological site known as "the Pterosaur Beach of Crayssac" (
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
; south-western
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
), evaluate the implications of these tracks for the knowledge of the terrestrial capabilities of non-pterodactyloid pterosaurs, and name a new Ichnotaxon, ichnogenus ''Rhamphichnus''. * The record of Pterosaur#Pycnofibers, pycnofibers in two specimens of Anurognathidae, anurognathid pterosaurs from the Jurassic of
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
interpreted by Yang ''et al.'' (2019) as showing diagnostic features of feathers is reexamined and challenged by Unwin & Martill (2020). * A Coleoidea, coleoid cephalopod specimen preserved with an associated tooth of a pterosaur (probably ''Rhamphorhynchus'') is reported from the Upper Jurassic Altmühltal Formation (
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
) by Hoffmann ''et al.'' (2020), who evaluate the implications of this finding for the knowledge of feeding behaviours of ''Rhamphorhynchus''. * A study on changes in the skeletal anatomy during growth in ''Rhamphorhynchus muensteri'' is published by Hone ''et al.'' (2020), who consider it likely that ''R. muensteri'' was able to fly soon after hatching. * A well-preserved basihyal is reported for the first time in a pterosaur specimen (possibly belonging to the species ''Gladocephaloideus jingangshanensis'') from the Lower Cretaceous Yixian Formation (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
) by Jiang ''et al.'' (2020). * Jacobs ''et al.'' (2020) describe new fossil material of pterosaurs from the Kem Kem Beds (
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
), bringing the Kem Kem pterosaur fauna up to at least nine species (of which three are Ornithocheiridae, ornithocheirids), and confirming that toothed pterosaurs remained diverse during the mid-Cretaceous. * Fossil material of pterosaurs (including a large non-pteranodontian Ornithocheiroidea, ornithocheiroid) is described from the Valanginian Rosablanca Formation by Cadena, Unwin & Martill (2020), representing the first record of pterosaurs from Colombia. * New forelimb of a Pteranodontoidea, pteranodontoid pterosaur is described from the Lower Cretaceous Yixian Formation (China) by Jiang ''et al.'' (2020), who also revise the species ''Yixianopterus jingangshanensis''. * Averianov (2020) reassesses the taxonomy of the Lonchodectidae, transferring the species "''Lonchodraco''" ''machaerorhynchus'' (including ''L. microdon'' and ''Pterodactylus oweni'') to the genus ''Ikrandraco''. * Martill ''et al.'' (2020) report evidence of the presence of clusters of circular foramina on the tip of the beak of ''Lonchodraco giganteus'', interpret this finding as evidence of enhanced sensitivity of the rostrum tip, and argue that this pattern implies tactile feeding in ''L. giganteus''. * An ornithocheirid Metacarpal bones, metacarpus, representing one of the geologically youngest ornithocheirid remains reported worldwide, is described from the Upper Cretaceous (Cenomanian) of the Crema Bonfil quarry (Coahuila, Mexico) by Frey ''et al.'' (2020), who evaluate the implications of this finding for the knowledge of the extinction of the toothed pterosaurs during the Late Cretaceous. * Fossil material of a large Pteranodontidae, pteranodontid pterosaur is described from the Campanian Beloe Ozero locality (Rybushka Formation; Saratov Oblast, Russia) by Averianov & Arkhangelsky (2020), who also review other putative records of pteranodontids in the Late Cretaceous of North America, Europe and Asia, and argue that ''Volgadraco bogolubovi'' is more likely to be a pteranodontid than Azhdarchidae, azhdarchid. * New Tapejaridae, tapejarid specimen, providing new information on the anatomy of the vertebral column of tapejarids, is described from the Aptian Crato Formation (
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
) by Cheng ''et al.'' (2020). * New information on the anatomy of ''Dsungaripterus weii'' (especially on the Palate, palatal region), based on the study of new and previously collected specimens, is published by Chen ''et al.'' (2020). * New fossil material of ''Tethydraco regalis'' is described from the Maastrichtian deposits of the
Ouled Abdoun Basin The Oulad Abdoun Basin (also known as the Ouled Abdoun Basin or Khouribga Basin) is a phosphate sedimentary basin located in Morocco, near the city of Khouribga. It is the largest in Morocco, comprising 44% of Morocco's phosphate reserves, and a ...
(
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
) by Labita & Martill (2020), who consider it more likely that ''T. regalis'' was an azhdarchid rather than a pteranodontian. * New specimens of edentulous pterosaur jaw fragments are described from the Cambridge Greensand (eastern England, United Kingdom) by Smith ''et al.'' (2020), who also revise the fossil material of ''Ornithostoma sedgwicki'' and assign this taxon to the group Azhdarchoidea rather than to the family Pteranodontidae.


New taxa


Other archosaurs


Research

* A study on the anatomy, locomotion and phylogenetic relationships of ''Scleromochlus taylori'' is published by Bennett (2020). * An Otischalkian assemblage of Lagerpetidae, lagerpetid and Silesauridae, silesaurid fossils, including lagerpetid material of unusually large size assignable to ''Dromomeron'', is described from the Los Esteros Member of the Santa Rosa Formation, New Mexico, Santa Rosa Formation (New Mexico, United States) by Beyl, Nesbitt & Stocker (2020), who interpret this finding as evidence that lagerpetids achieved large body size earlier than previously recognized. * A study on the phylogenetic relationships of lagerpetids is published by Ezcurra ''et al.'' (2020), who interpret lagerpetids as the sister group of pterosaurs. * New Dinosauromorpha, dinosauromorph fossil material is described from the Upper Triassic Chinle Formation (Petrified Forest National Park, Arizona, United States) by Marsh & Parker (2020), who also review the global fossil record of Late Triassic dinosauromorphs. * A study on the musculoskeletal apparatus and posture of ''Silesaurus opolensis'', evaluating its implications for the knowledge of the evolution of the fully erect limb posture in archosaurs, is published by Piechowski & Tałanda (2020).


New taxa


References

{{reflist Archosaurs 2020 in paleontology