Symplastic Pathway
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Symplastic Pathway
The symplast of a plant is the inner side of a cell membrane in which water and low-molecular-weight solutes can freely diffuse. Symplast cells have more than one nucleus. ''Symplast'' could also refer to the connection of the inner contents (cytoplasm) of neighbouring cells made by the microscopic channels that traverse the cell walls. These channels, which are called plasmodesmata, allow the direct flow of small molecules such as sugars, amino acids, and ions between cells (from the inner part of one cell to the inner partof the next cell). Larger molecules, including transcription factors and plant viruses, can also be transported through with the help of actin structures. This allows direct cytoplasm-to-cytoplasm flow of water and other nutrients along concentration gradients. In particular, symplastic flow is used in the root systems to bring in nutrients from soil. Nutrient solutes move in this way through three skin layers of the roots: from cells of the ''epidermis'' ...
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Apoplast And Symplast Pathways
Inside a plant, the apoplast can mean the space outside of cell membranes, where material can diffuse freely; that is, the extracellular spaces. ''Apoplast '' can also refer especially to the continuum of cell walls of adjacent cells; fluid and material flows occurring there or in any extacellular space are called ''apoplastic'' flow or apoplastic transport. The apoplastic route is one way by which water and solutes are transported and distributed to different places through tissues and organs; another way is symplastic flow. To prevent uncontrolled leakage to unwanted places, in certain areas there are barriers to the apoplastic flow: in roots the Casparian strip has this function larification needed On the outside of the skin of certain plant parts is a protective waxy film called plant cuticle to achieve this (protection against e.g. drying out, but also waterproofing against soaking). Air bubbles occupying extracellular spaces can also hinder apoplastic transport. The ap ...
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Apoplast
Inside a plant, the apoplast can mean the space outside of cell membranes, where material can diffuse freely; that is, the extracellular spaces. ''Apoplast '' can also refer especially to the continuum of cell walls of adjacent cells; fluid and material flows occurring there or in any extacellular space are called ''apoplastic'' flow or apoplastic transport. The apoplastic route is one way by which water and solutes are transported and distributed to different places through tissues and organs; another way is symplastic flow. To prevent uncontrolled leakage to unwanted places, in certain areas there are barriers to the apoplastic flow: in roots the Casparian strip has this function larification needed On the outside of the skin of certain plant parts is a protective waxy film called plant cuticle to achieve this (protection against e.g. drying out, but also waterproofing against soaking). Air bubbles occupying extracellular spaces can also hinder apoplastic transport. The a ...
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Protoplast
Protoplast (), is a biological term coined by Hanstein in 1880 to refer to the entire cell, excluding the cell wall. Protoplasts can be generated by stripping the cell wall from plant, bacterial, or fungal cells by mechanical, chemical or enzymatic means. Protoplasts differ from spheroplasts in that their cell wall has been completely removed. Spheroplasts retain part of their cell wall. In the case of Gram-negative bacterial spheroplasts, for example, the peptidoglycan component of the cell wall has been removed but the outer membrane component has not. Enzymes for the preparation of protoplasts Cell walls are made of a variety of polysaccharides. Protoplasts can be made by degrading cell walls with a mixture of the appropriate polysaccharide-degrading enzymes: During and subsequent to digestion of the cell wall, the protoplast becomes very sensitive to osmotic stress. This means cell wall digestion and protoplast storage must be done in an isotonic solution to prevent ...
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Polar Auxin Transport
Polar auxin transport is the regulated transport of the plant hormone auxin in plants. It is an active process, the hormone is transported in cell-to-cell manner and one of the main features of the transport is its asymmetry and directionality (polarity). The polar auxin transport functions to coordinate plant development; the following spatial auxin distribution underpins most of plant growth responses to its environment and plant growth and developmental changes in general. In other words, the flow and relative concentrations of auxin informs each plant cell where it is located and therefore what it should do or become. Chemiosmotic model Polar auxin transport (PAT) is directional and active flow of auxin molecules through the plant tissues. The flow of auxin molecules through the neighboring cells is driven by carriers (''type of membrane transport protein'') in the cell-to-cell fashion (from one cell to other cell and then to the next one) and the direction of the flow is de ...
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Plant Sap
Sap is a fluid transported in xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a separate substance, separately produced, and with different components and functions. Insect honeydew is called sap, particularly when it falls from trees, but is only the remains of eaten sap and other plant parts. Types of sap Saps may be broadly divided into two types: xylem sap and phloem sap. Xylem sap Xylem sap (pronounced ) consists primarily of a watery solution of hormones, mineral elements and other nutrients. Transport of sap in xylem is characterized by movement from the roots toward the leaves. Over the past century, there has been some controversy regarding the mechanism of xylem sap transport; today, most plant scientists agree that the cohesion-tension theory best explains this process, but multiforce theories that hy ...
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Ernst Münch
Ernst Münch (26 November 1876 – 9 October 1946) was a German plant physiologist who proposed the Pressure Flow Hypothesis in 1930. He studied in Aschaffenburg, and then in Munich with Robert Hartig. He worked in a number of fields including forest pathology, resin production, and fungi. He is best known for the phloem pressure flow hypothesis. Works * ''Untersuchungen über Immunität und Krankheitsempfänglichkeit der Holzpflanzen'', Dissertation, Ludwigsburg 1909 (doctorate thesis) * ''Die Stoffbewegungen in der Pflanze'', Jena 1930 * ''Beiträge zur Forstpflanzenzüchtung. Versuche einer Auslesezüchtung durch Einzelstamm-Absaaten bei Fichte. Weitere Beiträge zur Forstpflanzenzüchtung'' us dem wissenschaftlichen Nachlass herausgegeben von Bruno Huber München 1949 Further reading * E. Höxtermann: ''Ernst Münch''. In: Ilse Jahn (Hrsg.): ''Geschichte der Biologie. Theorien, Methoden, Institutionen, Kurzbiografien''. 3. Auflage, Spektrum Akademischer Verlag, Heid ...
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Johannes Von Hanstein
Johannes Ludwig Emil Robert von Hanstein (15 May 1822 – 27 August 1880) was a German botanist who was a native of Potsdam. He attended classes at the ''Gärtnerlehranstalt'' (Institute of Horticulture) in Potsdam, and later studied sciences in Berlin, obtaining his doctorate in 1848. In 1855 he was a lecturer of botany at the University of Berlin, and six years later became curator of the royal herbarium. In 1865 he was appointed professor of botany at the University of Bonn and director of the botanical garden. Hanstein is remembered for studies in plant anatomy and morphology. In 1868 he introduced the " histogen theory" to explain shoot apex behaviour in plants. With his close friend, Nathanael Pringsheim (1823–1894), he conducted pioneer research on the fertilization process in ferns. The plant genus '' Hansteinia'' of the family Acanthaceae is named after him. Gallery File:Capsella bursa-pastoris, kiembol (Hanstein).jpg, ''Capsella bursa-pastoris'' drawing by Joha ...
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Eduard Strasburger
Eduard Adolf Strasburger (1 February 1844 – 18 May 1912) was a Polish-German professor and one of the most famous botanists of the 19th century. He discovered mitosis in plants. Life Eduard Strasburger was born in Warsaw, Congress Poland, the son of Krystyna Anna (von Schütz) and Edward Bogumił Strasburger (1803–1874).Klaus Oskar Leyde: Strasburger. In: Deutsches Geschlechterbuch Band 207 (56. Allgemeiner Band), C. A. Starke Verlag, Limburg 1998, S. 227–242. In 1870, he married Aleksandra Julia Wertheim (1847–1902), they had two children: Anna (1870–1942) and Julius (1871–1934). Strasburger studied biological sciences in Paris, Bonn and Jena, receiving a PhD in 1866 after working with Nathanael Pringsheim. In 1868 he taught at the University of Warsaw. In 1869 he was appointed professor of botany at the University of Jena. From 1881 he was head of the ''Botanisches Institut'' at the University of Bonn. Strasburger died in Bonn, Germany. Achievements Stras ...
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Huntia (journal)
''Huntia'' is a peer-reviewed scientific journal published by the Hunt Institute for Botanical Documentation, a research division of the Carnegie Mellon University. In continuous publication since 1964, this journal is the institute's scholarly journal of botanical history. The journal is published irregularly in one or more numbers per volume of approximately 200 pages by Hunt Institute. Huntia was established by the American botanist George H.M. Lawrence, who was the founding director of the Hunt Institute. Volumes 1 to 7, was issued in 14 volumes, large octavo (folded paper) they were paperback apart from Volume 2 which was clothbound. Vol.2 was reviewed by botanist William C. Steere in June 1966. He noted "outstanding typography, high quality of illustrations and paper, the beautiful style and manufacture all indicate the high standards we have come to anticipate in publications of the Hunt Botanical Library". There was a big gap in publication of the journal when Lawrence ...
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Plasmodesmata
Plasmodesmata (singular: plasmodesma) are microscopic channels which traverse the cell walls of plant cells and some algal cells, enabling transport and communication between them. Plasmodesmata evolved independently in several lineages, and species that have these structures include members of the Charophyceae, Charales, Coleochaetales and Phaeophyceae (which are all algae), as well as all embryophytes, better known as land plants. Unlike animal cells, almost every plant cell is surrounded by a polysaccharide cell wall. Neighbouring plant cells are therefore separated by a pair of cell walls and the intervening middle lamella, forming an extracellular domain known as the apoplast. Although cell walls are permeable to small soluble proteins and other solutes, plasmodesmata enable direct, regulated, symplastic transport of substances between cells. There are two forms of plasmodesmata: primary plasmodesmata, which are formed during cell division, and secondary plasmodesmata, which c ...
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Xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from the Ancient Greek word (''xylon''), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Xylem also contains two other type of cells: parenchyma and fibers. Xylem can be found: * in vascular bundles, present in non-woody plants and non-woody parts of woody plants * in secondary xylem, laid down by a meristem called the vascular cambium in woody plants * as part of a stelar arrangement n ...
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Pericycle
The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants. Although it is composed of non-vascular parenchyma cells, it's still considered part of the vascular cylinder because it arises from the procambium as do the vascular tissues it surrounds. In eudicots, it also has the capacity to produce lateral roots. Branch roots arise from this primary meristem tissue. In plants undergoing secondary growth, the pericycle contributes to the vascular cambium often diverging into a cork cambium. In angiosperms certain molecules within the endodermis and the surrounding vasculature are sent to the pericycle which promotes the growth of the root meristems. Location The pericycle is located between the endodermis and phloem in plant roots. In dicot stems, it is situated around the ring of vascular bundles in the stele. Function In dicot roots, the pericycle strengthens the roots and provides ...
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