Robustness (morphology)
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Robustness (morphology)
In biology, robustness is used to describe a taxon with a stronger and heavier build (morphology) when compared to a related gracile taxon. The terms are used in contrast to one another. The term is used by physical anthropologists and paleoanthropologists to refer to a big-boned and muscular body. For example, members of the genus ''Sapajus'' have robust body types and are called the robust capuchin monkeys while members of the genus ''Cebus'' have gracile body types and are called the gracile capuchin monkeys. Male and female members of the same species may display sexual dimorphism and have robust and gracile morphologies. The terms "robust" vs. "gracile" are used in the context of human evolution, to distinguish: *"robust" vs. "gracile" australopithecines, see '' Paranthropus'' *"robust" archaic humans vs. "gracile" anatomically modern humans *"robust" early modern humans (Cro-Magnon) vs. "gracile" Epipaleolithic In archaeology, the Epipalaeolithic or Epipaleolithic (some ...
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Biology
Biology is the scientific study of life. It is a natural science with a broad scope but has several unifying themes that tie it together as a single, coherent field. For instance, all organisms are made up of cells that process hereditary information encoded in genes, which can be transmitted to future generations. Another major theme is evolution, which explains the unity and diversity of life. Energy processing is also important to life as it allows organisms to move, grow, and reproduce. Finally, all organisms are able to regulate their own internal environments. Biologists are able to study life at multiple levels of organization, from the molecular biology of a cell to the anatomy and physiology of plants and animals, and evolution of populations.Based on definition from: Hence, there are multiple subdisciplines within biology, each defined by the nature of their research questions and the tools that they use. Like other scientists, biologists use the sc ...
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Sexual Dimorphism
Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most animals and some plants. Differences may include secondary sex characteristics, size, weight, colour, markings, or behavioural or cognitive traits. These differences may be subtle or exaggerated and may be subjected to sexual selection and natural selection. The opposite of dimorphism is ''monomorphism'', which is when both biological sexes are phenotypically indistinguishable from each other. Overview Ornamentation and coloration Common and easily identified types of dimorphism consist of ornamentation and coloration, though not always apparent. A difference in coloration of sexes within a given species is called sexual dichromatism, which is commonly seen in many species of birds and reptiles. Sexual selection leads to the exaggerated dim ...
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Combe-Capelle
Combe-Capelle is a Paleolithic and Epipaleolithic site situated in the Couze valley in the Périgord region of Southern France. Henri-Marc Ami carried out excavations in the area from the late 1920s until his death in 1931. The famous ''Homo sapiens'' fossil from Combe-Capelle, discovered in 1909 was sold to the Museum für Völkerkunde, Berlin, in 1910. It was for a long time considered to be 30,000 years old, an Upper Paleolithic Cro-Magnon man and one of the oldest finds of modern humans in Europe, formerly classified as '' Homo aurignaciensis hauseri''. This was revised in a 2011 study, which dated collagen from a tooth of the skull in Berlin with accelerator mass spectrometry. The fossil was found to date to the early Holocene (Mesolithic Europe The Mesolithic (Greek: μέσος, ''mesos'' 'middle' + λίθος, ''lithos'' 'stone') or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic is oft ...
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Epipaleolithic
In archaeology, the Epipalaeolithic or Epipaleolithic (sometimes Epi-paleolithic etc.) is a period occurring between the Upper Paleolithic and Neolithic during the Stone Age. Mesolithic also falls between these two periods, and the two are sometimes confused or used as synonyms. More often, they are distinct, referring to approximately the same period of time in different geographic areas. Epipaleolithic always includes this period in the Levant and, often, the rest of the Near East. It sometimes includes parts of Southeast Europe, where Mesolithic is much more commonly used. Mesolithic very rarely includes the Levant or the Near East; in Europe, Epipalaeolithic is used, though not very often, to refer to the early Mesolithic. The Epipalaeolithic has been defined as the "final Upper Palaeolithic industries occurring at the end of the final glaciation which appear to merge technologically into the Mesolithic". The period is generally dated from   BP to 10,000 BP in t ...
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Cro-Magnon
Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They interacted and interbred with the indigenous Neanderthals (''H. neanderthalensis'') of Europe and Western Asia, who went extinct 40,000 to 35,000 years ago; and from 37,000 years ago onwards all EEMH descended from a single founder population which contributes ancestry to present-day Europeans. Early European modern humans (EEMH) produced Upper Palaeolithic cultures, the first major one being the Aurignacian, which was succeeded by the Gravettian by 30,000 years ago. The Gravettian split into the Epi-Gravettian in the east and Solutrean in the west, due to major climate degradation during the Last Glacial Maximum (LGM), peaking 21,000 years ago. As Europe warmed, the Solutrean evolved into the Magdalenian by 20,000 years ago, and these p ...
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Early Modern Humans
Early modern human (EMH) or anatomically modern human (AMH) are terms used to distinguish ''Homo sapiens'' (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans from extinct archaic human species. This distinction is useful especially for times and regions where anatomically modern and archaic humans co-existed, for example, in Paleolithic Europe. Among the oldest known remains of ''Homo sapiens'' are those found at the Omo-Kibish I archaeological site in south-western Ethiopia, dating to about 233,000 to 196,000 years ago, the Florisbad site in South Africa, dating to about 259,000 years ago, and the Jebel Irhoud site in Morocco, dated about 315,000 years ago. Extinct species of the genus ''Homo'' include ''Homo erectus'' (extant from roughly 2 to 0.1 million years ago) and a number of other species (by some authors considered subspecies of either ''H. sapiens'' or ''H. erectus''). The divergence of ...
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Anatomically Modern Humans
Early modern human (EMH) or anatomically modern human (AMH) are terms used to distinguish ''Homo sapiens'' (the only extant Hominina species) that are anatomically consistent with the range of phenotypes seen in contemporary humans from extinct archaic human species. This distinction is useful especially for times and regions where anatomically modern and archaic humans co-existed, for example, in Paleolithic Europe. Among the oldest known remains of ''Homo sapiens'' are those found at the Omo-Kibish I archaeological site in south-western Ethiopia, dating to about 233,000 to 196,000 years ago, the Florisbad site in South Africa, dating to about 259,000 years ago, and the Jebel Irhoud site in Morocco, dated about 315,000 years ago. Extinct species of the genus ''Homo'' include ''Homo erectus'' (extant from roughly 2 to 0.1 million years ago) and a number of other species (by some authors considered subspecies of either ''H. sapiens'' or ''H. erectus''). The divergence of ...
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Archaic Humans
A number of varieties of ''Homo'' are grouped into the broad category of archaic humans in the period that precedes and is contemporary to the emergence of the earliest early modern humans (''Homo sapiens'') around 300 ka. Omo-Kibish I (Omo I) from southern Ethiopia ( 195 or 233 ka), the remains from Jebel Irhoud in Morocco (about 315 ka) and Florisbad in South Africa (259 ka) are among the earliest remains of ''Homo sapiens''. The term typically includes ''Homo neanderthalensis'' (430 ± 25 ka), Denisovans, ''Homo rhodesiensis'' (300–125 ka), ''Homo heidelbergensis'' (600–200 ka), ''Homo naledi'', ''Homo ergaster'', ''Homo antecessor'', and '' Homo habilis''. There is no universal consensus on this terminology, and varieties of "archaic humans" are included under the binomial name of either ''Homo sapiens'' or ''Homo erectus'' by some authors. Archaic humans had a brain size averaging 1,200 to 1,400 cubic centimeters, which overlaps with the range of modern ...
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Paranthropus
''Paranthropus'' is a genus of extinct hominin which contains two widely accepted species: ''Paranthropus robustus, P. robustus'' and ''P. boisei''. However, the validity of ''Paranthropus'' is contested, and it is sometimes considered to be synonym (taxonomy), synonymous with ''Australopithecus''. They are also referred to as the robust australopithecines. They lived between approximately 2.6 and 1.2 million years ago (mya) from the end of the Pliocene to the Middle Pleistocene. ''Paranthropus'' is characterised by Robustness (morphology), robust skulls, with a prominent gorilla-like sagittal crest along the midline—which suggest strong chewing muscles—and broad, herbivorous teeth used for grinding. However, they likely preferred soft food over tough and hard food. ''Paranthropus'' species were generalist feeders, but ''P. robustus'' was likely an omnivore, whereas ''P. boisei'' was likely herbivorous and mainly ate bulbotubers. They were bipeds. Despite their robust heads, ...
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Australopithecines
Australopithecina or Hominina is a subtribe in the tribe Hominini. The members of the subtribe are generally ''Australopithecus'' (cladistically including the genera ''Homo'', '' Paranthropus'', and ''Kenyanthropus''), and it typically includes the earlier ''Ardipithecus'', ''Orrorin'', ''Sahelanthropus'', and ''Graecopithecus''. All these closely related species are now sometimes collectively termed australopiths or homininians. They are the extinct, close relatives of humans and, with the extant genus ''Homo'', comprise the human clade. Members of the human clade, i.e. the Hominini after the split from the chimpanzees, are now called Hominina (''see Hominidae; terms "hominids" and hominins''). While none of the groups normally directly assigned to this group survived, the australopiths do not appear to be literally extinct (in the sense of having no living descendants) as the genera ''Kenyanthropus'', ''Paranthropus'' and ''Homo'' probably emerged as sister of a late ''Aus ...
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Human Evolution
Human evolution is the evolutionary process within the history of primates that led to the emergence of ''Homo sapiens'' as a distinct species of the hominid family, which includes the great apes. This process involved the gradual development of traits such as human bipedalism and language, as well as interbreeding with other hominins, which indicate that human evolution was not linear but a web.Human Hybrids
(PDF). Michael F. Hammer. ''Scientific American'', May 2013.
The study of human evolution involves scientific disciplines, including
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Gracile Capuchin Monkey
Gracile capuchin monkeys are capuchin monkeys in the genus ''Cebus''. At one time all capuchin monkeys were included within the genus ''Cebus''. In 2011, Jessica Lynch Alfaro ''et al.'' proposed splitting the genus between the robust capuchin monkeys, such as the tufted capuchin, and the gracile capuchins. The gracile capuchins retain the genus name ''Cebus'', while the robust species have been transferred to ''Sapajus''. Taxonomy Following Groves (2005), taxa within the genus ''Cebus'' include: * White-fronted capuchin, ''Cebus albifrons'' ** Ecuadorian capuchin, ''Cebus albifrons aequatorialis'' ** Humboldt's white-fronted capuchin, ''Cebus albifrons albifrons'' ** Shock-headed capuchin, ''Cebus albifrons cuscinus'' ** Trinidad white-fronted capuchin, ''Cebus albifrons trinitatis'' ** Spix's white-fronted capuchin, ''Cebus albifrons unicolor'' ** Varied capuchin, ''Cebus albifrons versicolor'' * White-headed capuchin or white-faced capuchin, ''Cebus capucinus'' * Kaapor ...
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