Procolophonia
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Procolophonia
The Procolophonia are a suborder of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll 1988) but are now considered a clade of Parareptilia. They are closely related to other generally lizard-like Permian reptiles such as the Millerettidae, Bolosauridae, Acleistorhinidae, and Lanthanosuchidae, all of which are included under the Anapsida or "Parareptiles" (as opposed to the Eureptilia). Classification There are two main groups of Procolophonia, the small, lizard-like Procolophonoidea, and the Pareiasauroidea, which include the large, armoured Pareiasauridae. According to the traditional classification of Carroll 1988 as well as recent phylogenetic analyses, smaller groups like Rhipaeosauridae (now a synonym of Nycteroleteridae) and Sclerosauridae are classified with the pareiasaurs and with the procolophonids, respectively. T ...
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Owenettidae
Owenettidae is an extinct family of procolophonian parareptiles. Fossils have been found primarily from Africa and Madagascar, with one genus present from South America. It is the sister taxon to the family Procolophonidae. The family was constructed in 1939 for the type genus ''Owenetta''. Since then several other genera have been assigned to Owenettidae, including '' Barasaurus'' and '' Saurodektes''. The oldest known owenettid, ''Owenetta rubidgei'', dates back to the Wuchiapingian stage of the Late Permian. It and ''Saurodektes'', have been found from the Beaufort Group in the Karoo Basin of South Africa, while '' Barasaurus'' is known from the Late Permian and the Early Triassic of Sakamena Group in Madagascar. These strata span the Permo-Triassic boundary and provide evidence for the great faunal change that occurred as a result of the Permian–Triassic extinction event. Although most owenettids lived during the Permian, remains of some taxa such as ''"Owenetta" kitchingoru ...
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Nycteroleteridae
Nycteroleteridae is a family of procolophonian parareptilians (extinct early reptiles) from the Middle to Late Permian of Russia and North America. They are sometimes classified as a sister group to pareiasaurids (but see ''Classification)''. The group includes the genera ''Macroleter'', ''Bashkyroleter'', ''"Bashkyroleter" mesensis'', ''Nycteroleter'', ''Emeroleter'', and probably ''Rhipaeosaurus''. They were carnivorous, and occasionally ate insects. The group was most common in European Russia, with only a few fossils in North America. One fossil has also been found in Africa, but this is the only one from Gondwana. Classification Nycteroleteridae is sometimes considered a sister group to the Pareiasauridae, but Bayesian inference suggests that it was in fact paraphyletic, with ''Rhipaeosaurus'' a basal member of the Pareiasauridae and other members of the Nycteroleteridae as outgroups. This is supported by the appearance of ''Rhipaeosaurus skull and teeth - it had tricuspid ...
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Procolophonoid
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa. Taxonomy Procolophonoidea includes the families Owenettidae and Procolophonidae. '' Sclerosaurus'', which is placed within its own family Sclerosauridae, may be a member of the superfamily as well. In 1997, De Braga and Rieppel defined this same taxon (the oldest common ancestor of Procolophonidae and Owenettidae and all its descendants) using the name Procolophoniformes.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. When the superfamily was constructed in 1956, it was thought to be within the anthracosaur suborder Diadectomorpha. Since then it has b ...
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Rhipaeosauridae
Nycteroleteridae is a family of procolophonian parareptilians (extinct early reptiles) from the Middle to Late Permian of Russia and North America. They are sometimes classified as a sister group to pareiasaurids (but see ''Classification)''. The group includes the genera ''Macroleter'', '' Bashkyroleter'', ''"Bashkyroleter" mesensis'', ''Nycteroleter'', ''Emeroleter'', and probably ''Rhipaeosaurus''. They were carnivorous, and occasionally ate insects. The group was most common in European Russia, with only a few fossils in North America. One fossil has also been found in Africa, but this is the only one from Gondwana. Classification Nycteroleteridae is sometimes considered a sister group to the Pareiasauridae, but Bayesian inference suggests that it was in fact paraphyletic, with ''Rhipaeosaurus'' a basal member of the Pareiasauridae and other members of the Nycteroleteridae as outgroups. This is supported by the appearance of ''Rhipaeosaurus skull and teeth - it had tricu ...
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Acleistorhinidae
Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged (Moscovian to Kungurian stage) parareptiles. Acleistorhinids are most diverse from the Richards Spur locality of the Early Permian of Oklahoma. Richards Spur acleistorhinids include '' Acleistorhinus'', '' Colobomycter'', and possibly '' Delorhynchus'' and ''Feeserpeton''. Other taxa include '' Carbonodraco'' from the Late Carboniferous of Ohio and ''Karutia'' from the Early Permian of Brazil. Acleistorhinidae is commonly considered a subgroup of lanthanosuchoids, related to taxa such as '' Chalcosaurus'', '' Lanthaniscus'' and '' Lanthanosuchus''. However, a re-examination of parareptile phylogeny conducted by Cisneros ''et al.'' (2021) argued that lanthanosuchids were not closely related to acleistorhinids. The phylogenetic analysis conducted by these authors recovered acleistorhinids as the sister group of the clade Procolophonia, while lanthanosuchids were recovered within the procolopho ...
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Procolophonoidea
Procolophonoidea is an extinct superfamily of procolophonian parareptiles. Members were characteristically small, stocky, and lizard-like in appearance. Fossils have been found worldwide from many continents including Antarctica. The first members appeared during the Late Permian in the Karoo Basin of South Africa. Taxonomy Procolophonoidea includes the families Owenettidae and Procolophonidae. '' Sclerosaurus'', which is placed within its own family Sclerosauridae, may be a member of the superfamily as well. In 1997, De Braga and Rieppel defined this same taxon (the oldest common ancestor of Procolophonidae and Owenettidae and all its descendants) using the name Procolophoniformes.Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132. When the superfamily was constructed in 1956, it was thought to be within the anthracosaur suborder Diadectomorpha. Since then it has b ...
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Bolosauridae
Bolosauridae is an extinct family of ankyramorph parareptiles known from the latest Carboniferous (Gzhelian) or earliest Permian (Asselian) to the early Guadalupian epoch (latest Roadian stage) of North America, China, Germany, Russia and France. The bolosaurids were unusual for their time period by being bipedal, the oldest known tetrapods to have been so. Their teeth suggest that they were herbivores. The bolosaurids were a rare group and died out without any known descendants. The following cladogram shows the phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... position of the Bolosauridae, from Johannes Müller, Jin-Ling Li and Robert R. Reisz, 2008. References Procolophonomorphs Permian reptiles Paleozoic reptiles of Asia Paleozoic reptiles of Europe Pa ...
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Lanthanosuchidae
Lanthanosuchidae is a family of procolophonomorph parareptiles that lived 268–255 million years ago. The group was named in 1946 by Ivan Antonovich Efremov.Ivakhnenko, M.F. (1980). "Lanthanosuchids from the Permian of the East European Platform". ''Paleontologica Journal'' 1980: 80-90. References Procolophonomorphs Permian reptiles Prehistoric reptile families {{paleo-reptile-stub ...
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Anapsida
An anapsid is an amniote whose skull lacks one or more skull openings (fenestra, or fossae) near the temples. Traditionally, the Anapsida are the most primitive subclass of amniotes, the ancestral stock from which Synapsida and Diapsida evolved, making anapsids paraphyletic. It is however doubtful that all anapsids lack temporal fenestra as a primitive trait, and that all the groups traditionally seen as anapsids truly lacked fenestra. Anapsids and the turtles While "anapsid reptiles" or "anapsida" were traditionally spoken of as if they were a monophyletic group, it has been suggested that several groups of reptiles that had anapsid skulls might be only distantly related. Scientists still debate the exact relationship between the basal (original) reptiles that first appeared in the late Carboniferous, the various Permian reptiles that had anapsid skulls, and the Testudines (turtles, tortoises, and terrapins). However, it was later suggested that the anapsid-like turtle skull is ...
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Early Permian
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Eureptilia
Eureptilia ("true reptiles") is one of the two major subgroups of the clade Sauropsida, the other one being Parareptilia. Eureptilia includes Diapsida (the clade containing all modern reptiles and birds), as well as a number of primitive Permo- Carboniferous forms previously classified under Anapsida, in the old (no longer recognised) order "Cotylosauria". Eureptilia is characterized by the skull having greatly reduced supraoccipital, tabular, and supratemporal bones that are no longer in contact with the postorbital. Aside from Diapsida, the group notably contains Captorhinidae, a diverse and long lived (Late Carboniferous-Late Permian) clade of initially small carnivores that later evolved into large herbivores. Other primitive eureptiles such as ''Hylonomus'' and "protorothyrids" were all small, superficially lizard-like forms, that were probably insectivorous. One primitive eureptile, the Late Carboniferous "protorothyrid" ''Anthracodromeus,'' is the oldest known climbin ...
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Parareptilia
Parareptilia ("at the side of reptiles") is a subclass or clade of basal sauropsids (reptiles), typically considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as ''Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by the ...
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