Poposauroid
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Poposauroid
Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. However, it excludes most large predatory quadrupedal "rauisuchians" such as rauisuchids and "prestosuchids". Those reptiles are now allied with crocodylomorphs (crocodile ancestors) in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes ''Poposaurus'' and its close relatives. Poposauroids went extinct at the end of the Triassic period along with other non-crocodylomorph pseudosuchians. However, they were among the most diverse and long-lasting ...
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Paracrocodylomorpha
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing ''Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of "Poposauridae" and Crocodylomorpha and all of its descendants ...
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Loricata
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as ''Postosuchus'' and ''Prestosuchus''. More specifically, Loricata includes Crocodylomorpha (the persistent archosaur subset which crocodilians belong to) and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs. Loricata is one branch of the group Paracrocodylomorpha; the other branch is the clade Poposauroidea, an unusual collection of strange "rauisuchians" including bipedal, herbivorous, and sail-backed forms. The vast majority of typical "rauisuchians", which were usually quadrupedal predators, occupy basal (early-branching) rungs of Loricata leading up to crocodylomorphs. History and Classification Loricata was initially named in a completely different context by German naturalist Blasius Merrem in his 1820 ''Versuch eines Systems der Amphibien''. Merrem consid ...
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Xilousuchus
''Xilousuchus'' is an extinct genus of poposauroid from lower Triassic (Olenekian stage) deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is ''Xilousuchus sapingensis''. Wu (1981) referred ''Xilousuchus'' to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt ''et al.'' (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found ''Xilousuchus'' to be a poposauroid which is most closely related to ''Arizonasaurus''. ''Xilousuchus'' is the oldest archosaur to date, although ''Ctenosauriscus'' ...
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Rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...ns than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large (often ), carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional Taxonomy, taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, Phytosaur, phytosaurs (crocodile-like carnivores), Aetosaur, aetosaurs (armored herbivor ...
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Poposaurus
''Poposaurus'' (" Popo Agie reptile") is an extinct genus of pseudosuchian archosaur from the Late Triassic of the southwestern United States. It belongs to the clade Poposauroidea, an unusual group of Triassic pseudosuchians that includes sail-backed, beaked, and aquatic forms. Fossils have been found in Wyoming, Utah, Arizona, and Texas. Except for the skull, most parts of the skeleton are known. The type species, ''P. gracilis'', was described and named by Maurice Goldsmith Mehl in 1915. A second species, ''P. langstoni'', was originally the type species of the genus ''Lythrosuchus''. Since it was first described, ''Poposaurus'' has been variously classified as a dinosaur, a phytosaur, and a "rauisuchian". Like theropod dinosaurs, ''Poposaurus'' was an obligate biped, meaning that it walked on two legs rather than four. However, as a pseudosuchian, it is more closely related to living crocodilians than to dinosaurs. ''Poposaurus'' is thought to have evolved this form of locomo ...
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Lotosaurus
''Lotosaurus'' is an extinct genus of sail-backed poposauroid Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. Howeve ... known from Hunan Province of central China. Discovery ''Lotosaurus'' is known from the holotype IVPP V 4881 (or possibly V 4880), an articulated and well-preserved skeleton. Other referred specimens include IVPP V 48013 (a skull) as well as many articulated and disarticulated skeletal remains concentrated in a bonebed which is almost completely composed of ''Lotosaurus'' bones. All known specimens of this genus were collected from this bonebed, known as the ''Lotosaurus'' site, which belongs to the Batung Formation (or alternatively Xinlingzhen Formation of the Badong Group). Further excavations in 2018 revealed many more specimens, as well as ...
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Rauisuchid
Rauisuchidae is a group of large (up to or more) predatory Triassic archosaurs. There is some disagreement over which genera should be included in Rauisuchidae and which should be in the related Prestosuchidae and Poposauridae, and indeed whether these should even be thought of as separate valid Family (biology), families. Rauisuchids occurred throughout much of the Triassic, and may have first occurred in the Early Triassic if some archosaurian taxa such as ''Scythosuchus'' and ''Tsylmosuchus'' are considered to be within the family. An early cladistic analysis of crocodylotarsan (pseudosuchian) archosaurs included ''Lotosaurus'', ''Fasolasuchus'', ''Rauisuchus'', and "the Kupferzell rauisuchid" (later called ''Batrachotomus'') within Rauisuchidae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, a later study found that ''Batrachotomus'' was a more Basal (phyl ...
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Effigia
''Effigia'' was an extinct genus of shuvosaurid known from the Late Triassic of New Mexico, south-western USA. With a bipedal stance, long neck, and a toothless beaked skull, ''Effigia'' and other shuvosaurids bore a resemblance to the ornithomimid dinosaurs of the Cretaceous Period. However, shuvosaurids were not dinosaurs, but were instead a specialized family of poposauroid pseudosuchians, meaning that their closest living relatives are crocodilians. Discovery The 2-meter (6 ft 7 in) holotype fossil was collected by Edwin H. Colbert in 1947. At this time Colbert led an excavation to collect blocks of rock from the Whitaker Quarry of Ghost Ranch, near Abiquiu, New Mexico. Colbert's expedition intended to recover abundant fossils of the basal theropod dinosaur ''Coelophysis'', and he believed that no other large vertebrates were present in the quarry. As a result, his crew did not even open the plaster jackets of most of the blocks that were shipped to the American Museum of ...
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Bipedalism
Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' and ''pes'' 'foot'). Types of bipedal movement include walking, running, and hopping. Several groups of modern species are habitual bipeds whose normal method of locomotion is two-legged. In the Triassic period some groups of archosaurs (a group that includes crocodiles and dinosaurs) developed bipedalism; among the dinosaurs, all the early forms and many later groups were habitual or exclusive bipeds; the birds are members of a clade of exclusively bipedal dinosaurs, the theropods. Within mammals, habitual bipedalism has evolved multiple times, with the macropods, kangaroo rats and mice, springhare, hopping mice, pangolins and hominin apes ( australopithecines, including humans) as well as various other extinct groups evolving the tra ...
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Qianosuchus
''Qianosuchus'' is an extinct genus of aquatic poposauroid archosaur from the middle Triassic (Anisian) Guanling Formation of Pan County, China. It is represented by two nearly complete skeletons and a crushed skull preserved in the limestone. ''Qianosuchus'' was at least 3 metres long, and had several skeletal adaptations which indicate a semi-marine lifestyle, similar to modern-day saltwater crocodiles. These adaptations have not been seen in any other archosaur from the Triassic. Description ''Qianosuchus'' had a skull around 33 cm (13 inches) long, with an elongated snout. The rostrum formed by the premaxilla is shallow at the front of the skull but deepens posteriorly. Each premaxilla has nine long teeth, and the maxillae bear eighteen teeth each. All the teeth are laterally compressed, curved backwards and serrated, like those of most other carnivorous archosaurs. The nares are expanded and elongated and almost collide with the antorbital fenestrae, meaning that the ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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