Pampatheres
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Pampatheres
Pampatheriidae ("Pampas beasts") is an extinct family of large plantigrade armored armadillos related to extant armadillos in the order Cingulata. However, pampatheriids have existed as a separate lineage since at least the middle Eocene Mustersan age, . Pampatheres evolved in South America during its long period of Cenozoic isolation. Although widespread, they were less diverse and abundant than the armadillos. ''Holmesina'' spread to North America after the formation of the Isthmus of Panama as part of the Great American Interchange. They finally disappeared on both continents in the end-Pleistocene extinctions, about 12,000 years ago. Description Pampatheres are believed to have attained a weight of up to . Like three-banded armadillos, and unlike glyptodonts, their armored shell was given some flexibility by three movable lateral bands of scutes. The osteoderms (bony plates in the skin comprising the armor) of pampatheres were each covered by a single keratinized scute, u ...
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Cingulata
Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. Dasypodids and chlamyphorids, the armadillos, are the only surviving families in the order. Two groups of cingulates much larger than extant armadillos (maximum body mass of 45 kg (100 lb) in the case of the giant armadillo) existed until recently: pampatheriids, which reached weights of up to 200 kg (440 lb) and chlamyphorid glyptodonts, which attained masses of 2,000 kg (4,400 lb) or more. The cingulate order originated in South America during the Paleocene epoch about 66 to 56 million years ago, and due to the continent's former isolation remained confined to it during most of the Cenozoic. However, the formation of a land bridge allowed members of all three families to migrate to southern North America during the Pliocene or early Pleistocene as part of the Great American Interchange. After surviving for tens of millions of years, all of the pampat ...
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Great American Interchange
The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land and freshwater fauna migrated from North America via Central America to South America and vice versa, as the volcanic Isthmus of Panama rose up from the sea floor and bridged the formerly separated continents. Although earlier dispersals had occurred, probably over water, the migration accelerated dramatically about 2.7 million years ( Ma) ago during the Piacenzian age. It resulted in the joining of the Neotropic (roughly South American) and Nearctic (roughly North American) biogeographic realms definitively to form the Americas. The interchange is visible from observation of both biostratigraphy and nature (neontology). Its most dramatic effect is on the zoogeography of mammals, but it also gave an opportunity for reptiles, amphibia ...
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Pampatherium
''Pampatherium'' is an extinct genus of xenarthran that lived in the Americas during the Pleistocene. Some species went extinct right at the Pleistocene-Holocene border. Distribution ''Pampatherium humboldtii'' and ''P. typum'' lived in South America (mostly Brazil) during the Pleistocene, with ''P. humboldtii'' surviving into the very Late Pleistocene. ''Pampatherium mexicanum'' was the only North American species, reaching as far north as Sonora, Mexico. It lived during the Rancholabrean. Description ''Pampatherium'' resembled a very large armadillo. One species, ''P. humboldtii'', weighed up to . Pampatheres generally resembled armadillos, particularly in the shape of it skull, long snout, and the presence of three areas on the carapace (movable bands, scapular and pelvic shields). Among the features that distinguish them from armadillos are their posterior teeth, which are bilobate rather than peg-like. Their endocranial morphology is also similar to glyptodonts. The osteod ...
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Eocene
The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope Carbon-13, 13C in the atmosphere was exceptionally low in comparison with the more common isotope Carbon-12, 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Popigai impact structure, Siberia and in what is now ...
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Armadillo
Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along with the anteaters and sloths. Nine extinct genera and 21 extant species of armadillo have been described, some of which are distinguished by the number of bands on their armor. All species are native to the Americas, where they inhabit a variety of different environments. Armadillos are characterized by a leathery armor shell and long, sharp claws for digging. They have short legs, but can move quite quickly. The average length of an armadillo is about , including its tail. The giant armadillo grows up to and weighs up to , while the pink fairy armadillo has a length of only . When threatened by a predator, ''Tolypeutes'' species frequently roll up into a ball; they are the only species of armadillo capable of this. Etymology The wor ...
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Holmesina 3 Clean
''Holmesina'' is a genus of pampathere, an extinct group of armadillo-like creatures that were distantly related to extant armadillos. Like armadillos, and unlike the other extinct branch of megafaunal cingulates, the glyptodonts, the shell was made up of flexible plates which allowed the animal to move more easily. ''Holmesina'' species were herbivores that grazed on coarse vegetation; armadillos are mostly insectivorous or omnivorous. ''Holmesina'' individuals were much larger than any modern armadillo: They could reach a length of , and a weight of , while the modern giant armadillo does not attain more than . Distribution They traveled north during the faunal interchange, and adapted well to North America, like the ground sloths, glyptodonts, armadillos, capybaras, and other South American immigrants. Their fossils are found from Brazil to the United States, mostly in Texas and Florida Florida is a state located in the Southeastern region of the United ...
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Keratin
Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, horns, claws, hooves, and the outer layer of skin among vertebrates. Keratin also protects epithelial cells from damage or stress. Keratin is extremely insoluble in water and organic solvents. Keratin monomers assemble into bundles to form intermediate filaments, which are tough and form strong unmineralized epidermal appendages found in reptiles, birds, amphibians, and mammals. Excessive keratinization participate in fortification of certain tissues such as in horns of cattle and rhinos, and armadillos' osteoderm. The only other biological matter known to approximate the toughness of keratinized tissue is chitin. Keratin comes in two types, the primitive, softer forms found in all vertebrates and harder, derived forms found only amon ...
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Osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amphibians), various groups of dinosaurs (most notably ankylosaurs and stegosaurians), phytosaurs, aetosaurs, placodonts, and hupehsuchians (marine reptiles with possible ichthyosaur affinities). Osteoderms are uncommon in mammals, although they have occurred in many xenarthrans (armadillos and the extinct glyptodonts and mylodontid and scelidotheriid ground sloths). The heavy, bony osteoderms have evolved independently in many different lineages. The armadillo osteoderm is believed to develop in subcutaneous dermal tissues. These varied structures should be thought of as anatomical analogues, not homologues, and do not necessarily indicate monophyly. The structures are however derived from scutes, common to all classes of amniotes and ...
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Scute
A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior portion of the mesonotum in insects as well as some arachnids (e.g., the family Ixodidae, the scale ticks). Properties Scutes are similar to scales and serve the same function. Unlike the scales of lizards and snakes, which are formed from the epidermis, scutes are formed in the lower vascular layer of the skin and the epidermal element is only the top surface . Forming in the living dermis, the scutes produce a horny outer layer that is superficially similar to that of scales. Scutes will usually not overlap as snake scales (but see the pangolin). The outer keratin layer is shed piecemeal, and not in one continuous layer of skin as seen in snakes or lizards. The dermal base may contain bone and produce dermal armour. Scutes with a bony ...
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Glyptodont
Glyptodonts are an extinct subfamily of large, heavily armoured armadillos. They arose in South America around 48 million years ago and spread to southern North America after the continents became connected several million years ago. The best-known genus within the group is ''Glyptodon''. While they were formerly considered to constitute the distinct family Glyptodontidae, in 2016, an analysis of ''Doedicurus'' Mitochondrial DNA (also known as mtDNA / mDNA) found that it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae. For this reason, glyptodonts and all armadillos but '' Dasypus'' were relocated to a new family, Chlamyphoridae, and glyptodonts were demoted from the former family Glyptodontidae to a subfamily. Evolution Glyptodonts first evolved during the Eocene in South America, which remained their center of species diversity. For example, an Early Miocene glyptodont with many primitive feature ...
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Tolypeutes
The genus ''Tolypeutes'' contains the two species of three-banded armadillos. They are restricted to open and semi-open habitats in South America. Of the several armadillo genera, only ''Tolypeutes'' rely heavily on their armor for protection. When threatened by a predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ..., ''Tolypeutes'' species frequently roll up into a ball. Other armadillo species cannot roll up because they have too many plates. This species is endangered due to the deforestation done in their native land of Brazil, including the hunting by the locals. They may have this defense mechanism but they lack the ability to dig burrows such as their competition in that environment but, they will look for abandoned burrows. Deforestation is the main concern due to the ...
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Quaternary Extinction Event
The Quaternary period (from 2.588 ± 0.005 million years ago to the present) has seen the extinctions of numerous predominantly megafaunal species, which have resulted in a collapse in faunal density and diversity and the extinction of key ecological strata across the globe. The most prominent event in the Late Pleistocene is differentiated from previous Quaternary pulse extinctions by the widespread absence of ecological succession to replace these extinct species, and the regime shift of previously established faunal relationships and habitats as a consequence. The earliest casualties were incurred at 130,000 BCE (the start of the Late Pleistocene), in Australia ~ 60,000 years ago, in Americas ~ 15,000 years ago, coinciding in time with the early human migrations. However, the great majority of extinctions in Afro-Eurasia and the Americas occurred during the transition from the Pleistocene to the Holocene epoch (13,000 BCE to 8,000 BCE). This extinction wave di ...
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