Orodrominae
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Orodrominae
Orodrominae is a subfamily of thescelosaurid dinosaurs known from the Cretaceous of North America and Asia. Distribution Orodromines were a mostly North American based group with fossils from Canada and United States only. ''Albertadromeus'', as its name suggests, is only from the upper (later) part of the Oldman Formation in the Belly River Group of Alberta, Canada. ''Orodromeus'', the type genus, was widespread through Montana. Its holotype was found at the Egg Mountain in the Two Medicine Formation. ''Oryctodromeus'' fossils were found in the Lima Peaks section of the Blackleaf Formation, also from Montana. ''Zephyrosaurus'', the most widespread genus, lived in southern Montana and northern Wyoming. The locality of its holotype is the Wolf Creek Canyon, which is a sandstone in the Cloverly Formation. Age Orodromines are widespread throughout time, starting in the Aptian and ending in the Campanian. The earliest fossils are of ''Zephyrosaurus'' and are from the Aptian (113 Ma ...
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Thescelosaurid
Thescelosauridae is a clade of neornithischians from the Cretaceous of Asia, North America and possibly South America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was used by Brown and colleagues as the group uniting ''Thescelosaurus'' and ''Orodromeus'', based on their phylogenetic results. During a phylogenetic revision of neornithischians by Clint Boyd in 2015, the authorship of Thescelosauridae was given to Brown and colleagues, which meant that the similar name Parksosauridae, informally defined in 2002 by Buchholz, would have had priority over Thescelosauridae. The two clades had slightly different definitions, with Parksosauridae referring to all animals closer to ''Parksosaurus'' than ''Hypsilophodon'', but they contained the same taxa so Boyd used Parksosauridae under the assumption it had priority. However, in formalizing the clade following the regulations of the PhyloCode, Madzia, Boyd, and c ...
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Thescelosauridae
Thescelosauridae is a clade of neornithischians from the Cretaceous of Asia, North America and possibly South America. The group was originally used as a name by Charles M. Sternberg in 1937, but was not formally defined until 2013, where it was used by Brown and colleagues as the group uniting ''Thescelosaurus'' and ''Orodromeus'', based on their phylogenetic results. During a phylogenetic revision of neornithischians by Clint Boyd in 2015, the authorship of Thescelosauridae was given to Brown and colleagues, which meant that the similar name Parksosauridae, informally defined in 2002 by Buchholz, would have had priority over Thescelosauridae. The two clades had slightly different definitions, with Parksosauridae referring to all animals closer to ''Parksosaurus'' than ''Hypsilophodon'', but they contained the same taxa so Boyd used Parksosauridae under the assumption it had priority. However, in formalizing the clade following the regulations of the PhyloCode, Madzia, Boyd, and c ...
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Albertadromeus
''Albertadromeus'' is an extinct genus of orodromine thescelosaurid dinosaur known from the upper part of the Late Cretaceous Oldman Formation (middle Campanian stage) of Alberta, Canada. It contains a single species, ''Albertadromeus syntarsus''. Etymology The composite term ''Albertadromeus'' is derived from the name of the Canadian province "Alberta", and the Greek word ''dromeus'' (δρομεύς) meaning "runner", a reference to its inferred cursorial nature; thus "runner from Alberta". The specific name, ''syntarsus'' is derived from Greek words "syn" (συν) meaning "together" and "tarsus" (ταρσός) meaning "ankle", hence "together-ankle" a reference to the condition where its distal fibula is fused to its distal tibia. This dinosaur was described and named by Caleb Marshall Brown, David C. Evans, Michael J. Ryan & Anthony P. Russell in 2013 and the type species is ''Albertadromeus syntarsus''. Description The holotype specimen of ''Albertadromeus'' TMP 2009.037.00 ...
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Thescelosaurinae
Thescelosaurinae is a subfamily of thescelosaurid dinosaurs from the Early Cretaceous of Asia and the Late Cretaceous of North America. Distribution The distribution of Thescelosaurinae is quite large. They are widespread through United States and Canada where most of their fossils have been found. They also have a small group in North-Eastern China and Mongolia. Genera Studies disagree about which genera are included in the Thescelosaurinae. The group is defined to include ''Parksosaurus'' and ''Thescelosaurus'' and a cladistic analysis by C.M. Brown in 2013 concluded that the genera '' Changchunsaurus'', '' Haya'', '' Jeholosaurus'' and possibly ''Koreanosaurus'' are also thescelosaurines. Classification All thescelosaurines were originally included in the family Hypsilophodontidae, which is presently considered polyphyletic (unnatural). They are all now considered to be basal neornithischians, though more research is needed to be certain. They are the sister taxa to Oro ...
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Orodromeus
''Orodromeus'' (meaning "Mountain Runner") is a genus of herbivorous orodromine thescelosaurid dinosaur from the Late Cretaceous of North America. Only one species is known, the type species ''Orodromeus makelai''. Discovery and naming The remains of ''Orodromeus'' were discovered by Robert Makela during the excavation in Teton County, Montana, of the ''Egg Mountain'' brooding colony of a much larger relative, ''Maiasaura''. The type species, ''Orodromeus makelai'', was named and shortly described by Jack Horner and David B. Weishampel in 1988. The generic name is derived from Greek ὄρος, ''oros'', "mountain", in reference to the Egg Mountain site, and δρομεύς, ''dromeus'', "runner", referring to the cursorial habits of the animal. The specific name honoured the late Makela.Horner, J. and Weishampel, D., 1988, "A comparative embryological study of two ornithischian dinosaurs", ''Nature'' (London), 332(No. 6161): 256-257 (1988) The holotype specimen, MOR 294, wa ...
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Dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Koreanosaurus
''Koreanosaurus'' () is a genus of orodromine neornithischian dinosaur. One species has been described, ''Koreanosaurus boseongensis''. Discovery In 2003, three specimens of ''Koreanosaurus'' were found in the Late Cretaceous-age Seonso Conglomerate from the southern coast of the Bibong-ri dinosaur egg site, Boseong, Korean Peninsula. These specimens include the holotype KDRC-BB2, a partial upper skeleton lacking the skull, and two additional specimens which contains portions of the pelvic girdle and lower leg (KDRC-BB1 and KDRC-BB3). The type species was named after its locality (Boseong site 5). This taxon was initially named and described in a master's thesis by Dae-Gil Lee in 2008, and was officially published by Min Huh, Dae-Gil Lee, Jung-Kyun Kim, Jong-Deock Lim and Pascal Godefroit in 2011. Description ''Koreanosaurus'' was a relatively small dinosaur, reaching in body length. Based on its taxonomic position and the existence of small burrows from the Seonso Conglome ...
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Oryctodromeus
''Oryctodromeus'' (meaning "digging runner") was a genus of small orodromine thescelosaurid dinosaur. Fossils are known from the Late Cretaceous Blackleaf Formation of southwestern Montana and the Wayan Formation of southeastern Idaho, USA, both of the Cenomanian stage, roughly 95 million years ago. A member of the small, presumably fast-running herbivorous family Thescelosauridae, ''Oryctodromeus'' is the first non-avian dinosaur published that shows evidence of burrowing behavior. Description ''Oryctodromeus'' was originally described as lacking ossified tendons in the tail. However, specimens from the Wayan Formation demonstrate the presence of a thick tendon lattice in the dorsal, sacral, and caudal columns of some specimens; perhaps indicating more flexibility in ossified tendons than has previously been supposed.Krumenacker, L. J., 2010. Chronostratigraphy and paleontology of the mid-Cretaceous Wayan Formation of eastern Idaho, with a description of the first ''Oryctod ...
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Wolf Creek Canyon
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly understood, comprise wild subspecies. The wolf is the largest extant member of the family Canidae. It is also distinguished from other ''Canis'' species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller ''Canis'' species, such as the coyote and the golden jackal, to produce fertile hybrids with them. The banded fur of a wolf is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white. Of all members of the genus ''Canis'', the wolf is most specialized for cooperative game hunting as demonstrated by its physical adaptations to tackling large prey, its more social nature, and its highly advanced ...
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Blackleaf Formation
The Blackleaf Formation is a geologic formation in Montana. It preserves fossils dating back to the Albian to Cenomanian stages of the Cretaceous period.Blackleaf Formation
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Description

The formation comprises an isolated, sinuous body sitting within and crosscutting a succession of three mudrock units. The lowermost is greater than 35 cm in t ...
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Wyoming
Wyoming () is a U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the south. With a population of 576,851 in the 2020 United States census, Wyoming is the List of U.S. states and territories by population, least populous state despite being the List of U.S. states and territories by area, 10th largest by area, with the List of U.S. states by population density, second-lowest population density after Alaska. The state capital and List of municipalities in Wyoming, most populous city is Cheyenne, Wyoming, Cheyenne, which had an estimated population of 63,957 in 2018. Wyoming's western half is covered mostly by the ranges and rangelands of the Rocky Mountains, while the eastern half of the state is high-elevation prairie called the High Plains (United States), High Plains. It is drier ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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