Hydroid (botany)
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Hydroid (botany)
A hydroid is a type of vascular cell that occurs in certain bryophytes. In some mosses such as members of the Polytrichaceae family, hydroids form the innermost layer of cells in the stem. At maturity they are long, colourless, thin walled cells of small diameter, containing water but no living protoplasm. Collectively, hydroids function as a conducting tissue, known as the hydrome, transporting water and minerals drawn from the soil. They are surrounded by bundles of living cells known as leptoids which carry sugars and other nutrients in solution. The hydroids are analogous to the tracheids of vascular plants but there is no lignin present in the cell walls to provide structural support. Hydroids have been found in some fossilised plants from the Rhynie chert, including ''Aglaophyton'', where they were initially mistaken for xylem tracheids. See also *Leptoid, a related sucrose-transporting vessel analogous to the phloem Phloem (, ) is the living biological tissue, tissue in ...
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Cell (biology)
The cell is the basic structural and functional unit of life forms. Every cell consists of a cytoplasm enclosed within a membrane, and contains many biomolecules such as proteins, DNA and RNA, as well as many small molecules of nutrients and metabolites.Cell Movements and the Shaping of the Vertebrate Body
in Chapter 21 of
Molecular Biology of the Cell
'' fourth edition, edited by Bruce Alberts (2002) published by Garland Science. The Alberts text discusses how the "cellular building blocks" move to shape developing embryos. It is also common to describe small molecules such as ...
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Bryophyte
The Bryophyta s.l. are a proposed taxonomic division containing three groups of non-vascular land plants (embryophytes): the liverworts, hornworts and mosses. Bryophyta s.s. consists of the mosses only. They are characteristically limited in size and prefer moist habitats although they can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetic evidence supports the monophyly of this group, as originally classified by Wilhelm Schimper in 1879. The term ''bryophyte'' comes . Terminology The term "Bryophyta" was first suggested by Braun in 1864. G.M. Smith placed this group between Algae and Pteridophyta. Features The d ...
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Moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically tall, though some species are much larger. ''Dawsonia'', the tallest moss in the world, can grow to in height. There are a ...
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Polytrichaceae
Polytrichaceae is a common family of mosses. Members of this family tend to be larger than other mosses with a thickened central stem and a rhizome. The leaves have a midrib that bears photosynthetic lamellae on the upper surface. Species in this group are dioicous. Another characteristic that identifies them is that they have from 32 to 64 peristome teeth in their sporangium. Classification Genera Extinct genera * '' Eopolytrichum'' Konopka, Herendeen, Merrill & Crane (1997), Gaillard Formation, Georgia, USA, Upper Cretaceous (Campanian) * '' Meantoinea'' Bippus, Stockey, Rothwell & Tomescu (2017) Apple Bay locality, Vancouver Island, Canada, Lower Cretaceous (Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 139.8 ± 3.0 Ma and 132.9 ± 2.0 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretace ...) References Moss families
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Protoplasm
Protoplasm (; ) is the living part of a cell that is surrounded by a plasma membrane. It is a mixture of small molecules such as ions, monosaccharides, amino acid, and macromolecules such as proteins, polysaccharides, lipids, etc. In some definitions, it is a general term for the cytoplasm (e.g., Mohl, 1846), but for others, it also includes the nucleoplasm (e.g., Strasburger, 1882). For Sharp (1921), "According to the older usage the extra-nuclear portion of the protoplast 'the entire cell, excluding the cell wall''was called "protoplasm," but the nucleus also is composed of protoplasm, or living substance in its broader sense. The current consensus is to avoid this ambiguity by employing Strasburger's '(1882)''terms cytoplasm Albert_von_Kölliker.html"_;"title="'coined_by_Albert_von_Kölliker">Kölliker_(1863),_originally_as_synonym_for_protoplasm''and_nucleoplasm_([''term_coined_by_Edouard_Van_Beneden.html" ;"title="Albert von Kölliker">Kölliker (1863), originally as syn ...
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Leptoid
A leptoid is a type of elongated food-conducting cell like phloem in the stems of some mosses, such as the family Polytrichaceae. They surround strands of water-conducting hydroids. They have some structural and developmental similarities to the sieve elements of seedless vascular plants. At maturity they have inclined end cell walls with small pores and degenerate nuclei. The conduction cells of mosses, leptoids and hydroids, appear similar to those of fossil protracheophytes. However they're not thought to represent an intermediate stage in the evolution of plant vascular tissues but to have had an independent evolutionary origin.Renzaglia, K.S., Schette, S. and Duff, R.J. (2007) Bryophyte phylogeny: advancing the molecular and morphological frontiers. ''Bryologist'', 110'', 179-213. See also * Hydroid, a related water-transporting cell analogous the xylem Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xyle ...
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Tracheid
A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms use another type of tracheary element, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns. The tracheid was first named after the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''. Evolution Tracheids were the main conductive cells found in early vascular plants. In the first 140-150 million years of vascular plant evolution, tr ...
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Lignin
Lignin is a class of complex organic polymers that form key structural materials in the support tissues of most plants. Lignins are particularly important in the formation of cell walls, especially in wood and bark, because they lend rigidity and do not rot easily. Chemically, lignins are polymers made by cross-linking phenolic precursors. History Lignin was first mentioned in 1813 by the Swiss botanist A. P. de Candolle, who described it as a fibrous, tasteless material, insoluble in water and alcohol but soluble in weak alkaline solutions, and which can be precipitated from solution using acid. He named the substance “lignine”, which is derived from the Latin word '' lignum'', meaning wood. It is one of the most abundant organic polymers on Earth, exceeded only by cellulose. Lignin constitutes 30% of non-fossil organic carbon on Earth, and 20 to 35% of the dry mass of wood. Lignin is present in red algae, which suggest that the common ancestor of plants and red algae ...
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Rhynie Chert
The Rhynie chert is a Lower Devonian sedimentary deposit exhibiting extraordinary fossil detail or completeness (a Lagerstätte). It is exposed near the village of Rhynie, Aberdeenshire, Scotland; a second unit, the Windyfield chert, is located some 700 m away. The Rhynie chert contains exceptionally preserved plant, fungus, lichen and animal material preserved in place by an overlying volcanic deposit. The bulk of the Devonian fossil bed consists of primitive plants (which had water-conducting cells and sporangia, but no true leaves), along with arthropods, lichens, algae and fungi. This fossil bed is remarkable for two reasons. First, the age of the site ( Pragian, Early Devonian, formed about ) places it at an early stage in the colonisation of land. Second, these cherts are famous for their exceptional state of ultrastructural preservation, with individual cell walls easily visible in polished specimens. Stomata have been counted and lignin remnants detected in the ...
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Aglaophyton
''Aglaophyton major'' (or more correctly ''Aglaophyton majus'') was the sporophyte generation of a diplohaplontic, pre-vascular, axial, free-sporing land plant of the Lower Devonian (Pragian stage, around ). It had anatomical features intermediate between those of the bryophytes and vascular plants or tracheophytes. ''A. major'' was first described by Kidston and Lang in 1920 as the new species ''Rhynia major''. The species is known only from the Rhynie chert in Aberdeenshire, Scotland, where it grew in the vicinity of a silica-rich hot spring, together with a number of associated vascular plants such as a smaller species ''Rhynia gwynne-vaughanii'' which may be interpreted as a representative of the ancestors of modern vascular plants and '' Asteroxylon mackei'', which was an ancestor of modern clubmosses (Lycopsida). Description The stems of ''Aglaophyton'' were round in cross-section, smooth, unornamented, and up to about 6mm in diameter. Kidston and Lang interpreted the pl ...
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Xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from the Ancient Greek word (''xylon''), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Xylem also contains two other type of cells: parenchyma and fibers. Xylem can be found: * in vascular bundles, present in non-woody plants and non-woody parts of woody plants * in secondary xylem, laid down by a meristem called the vascular cambium in woody plants * as part of a stelar arrangement n ...
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Leptoid
A leptoid is a type of elongated food-conducting cell like phloem in the stems of some mosses, such as the family Polytrichaceae. They surround strands of water-conducting hydroids. They have some structural and developmental similarities to the sieve elements of seedless vascular plants. At maturity they have inclined end cell walls with small pores and degenerate nuclei. The conduction cells of mosses, leptoids and hydroids, appear similar to those of fossil protracheophytes. However they're not thought to represent an intermediate stage in the evolution of plant vascular tissues but to have had an independent evolutionary origin.Renzaglia, K.S., Schette, S. and Duff, R.J. (2007) Bryophyte phylogeny: advancing the molecular and morphological frontiers. ''Bryologist'', 110'', 179-213. See also * Hydroid, a related water-transporting cell analogous the xylem Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xyle ...
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