Heterokont
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Heterokont
Heterokonts are a group of protists (formally referred to as Heterokonta, Heterokontae or Heterokontophyta). The group is a major line of eukaryotes. Most are algae, ranging from the giant multicellular kelp to the unicellular diatoms, which are a primary component of plankton. Other notable members of the Stramenopiles include the (generally) parasitic oomycetes, including ''Phytophthora'', which caused the Great Famine of Ireland, and ''Pythium'', which causes seed rot and damping off. The name "heterokont" refers to the type of motile life cycle stage, in which the flagellated cells possess two differently arranged flagella (see zoospore). History In 1899, Alexander Luther created the term "Heterokontae" for some algae with unequal flagella, today called Xanthophyceae. Later, some authors (e.g., Copeland, 1956) included other groups in Heterokonta, expanding the name's sense. The term continues to be applied in different ways, leading to Heterokontophyta being applied al ...
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Stramenopile
Stramenopile is a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria. Members of the clade are referred to as 'stramenopiles'. Stramenopiles are eukaryotes; since they are neither fungi, animals, nor plants, they are classified as protists. Most stramenopiles are single-celled, but some are multicellular algae including some brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. T ...
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Diatom
A diatom (New Latin, Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising several Genus, genera of algae, specifically microalgae, found in the oceans, waterways and soils of the world. Living diatoms make up a significant portion of the Earth's Biomass (ecology), biomass: they generate about 20 to 50 percent of the oxygen produced on the planet each year, take in over 6.7 billion metric tons of silicon each year from the waters in which they live, and constitute nearly half of the organic material found in the oceans. The Protist shell, shells of dead diatoms can reach as much as a half-mile (800 m) deep on the ocean floor, and the entire Amazon basin is fertilized annually by 27 million tons of diatom shell dust transported by transatlantic winds from the African Sahara, much of it from the Bodélé Dep ...
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Ochrophytina
The ochrophytes, subphylum Ochrophytina, is a group of mostly photosynthetic heterokonts. Their plastid is of red algal origin. The classification of the group is still being worked out. Originally, the ochrophytes were regarded as a phylum denominated Ochrophyta. Some authors (e.g., Cavalier-Smith) divided it into two subphyla, Phaeista Cavalier-Smith 1995 (comprising Hypogyristea and Chrysista in some classifications, or Limnista and Marista in others) and Khakista Cavalier-Smith, 2000 (comprising '' Bolidomonas'' and diatoms). Others prefer not to use the subphyla, listing only lower taxa (e.g., Reviers, 2002, Guiry & Guiry, 2014). However, it is currently regarded as a subphylum inside of the phylum Gyrista, along with Pseudofungi and Bigyromonada. It contains two infraphyla: Diatomista, containing diatoms and related groups, and Chrysista, containing brown and golden algae and related groups. Phylogeny The cladogram below shows the evolutionary relationships be ...
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Ochrophyta
The ochrophytes, subphylum Ochrophytina, is a group of mostly photosynthesis, photosynthetic heterokonts. Their plastid is of Red algae, red algal origin. The classification of the group is still being worked out. Originally, the ochrophytes were regarded as a phylum denominated Ochrophyta. Some authors (e.g., Thomas Cavalier-Smith, Cavalier-Smith) divided it into two subphyla, Phaeista Cavalier-Smith 1995 (comprising Hypogyristea and Chrysista in some classifications, or Limnista and Marista (alga), Marista in others) and Khakista Cavalier-Smith, 2000 (comprising ''Bolidomonas'' and diatoms). Others prefer not to use the subphyla, listing only lower taxa (e.g., Reviers, 2002, Guiry & Guiry, 2014). However, it is currently regarded as a subphylum inside of the phylum Gyrista, along with Pseudofungi and Bigyromonada. It contains two infraphyla: Diatomista, containing diatoms and related groups, and Chrysista, containing brown and golden algae and related groups. Phylogeny The c ...
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Bigyra
Bigyra is a grouping of heterokont organisms. It includes Bicosoecida, Blastocystis and Labyrinthulida. It has also been described as containing Opalozoa, Bicoecia, and Sagenista. Phylogeny The cladogram below shows the internal relationships within Bigyra: Classification The modern taxonomy of Bigyra is as follows: *Phylum Bigyra **Subphylum Opalozoa ***Infraphylum Placidozoa ****Superclass Wobblata (paraphyletic) *****Class Placididea *****Class Nanomonadea *****Class Opalomonadea ****Superclass Opalinata *****Class Opalinea *****Class Blastocystea ***Infraphylum Bikosia ****Class Bikosea **Subphylum Sagenista ***Class Labyrinthulea ***Class Eogyrea Sagenista is a group of heterokonts containing the labyrinthulids and Eogyrea, a class of yet uncultured protists. Originally, it contained the Labyrinthulids and bicosoecids. However at present the bicosoecids have been removed, and Eogyrea wer ... References External links * * SAR supergroup unranked clades
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Pseudofungi
Pseudofungi is a grouping of heterokonts, also known as the Heterokontimycotina. It consists of the Oomycota and Hyphochytridiomycetes. Although numerous biochemical, ultrastructural, and genetic traits clearly place them in the heterokonts, their growth form (featuring hyphae) and mode of nutrition ( osmotrophy) resemble that of fungi (which are not closely related). Origin and ancestors It is believed that pseudofungi descend from unicellular heterokont chromist algae which lost their plastids. While evidence of these plastids hasn't been found, what has been proven is the existence of endosymbiotic red algae plastids. From this, a unicellular heterotroph proto-pseudofungi (probably a mushroom parasite) got its fungal genes through horizontal gene transfer, which would have led to the development of convergent fungal multicellularity, explaining why the cell wall is sometimes made of both chitin and cellulose Cellulose is an organic compound with the formula , a polys ...
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Chrysophyceae
The Chrysophyceae, usually called chrysophytes, chrysomonads, golden-brown algae or golden algae are a large group of algae, found mostly in freshwater. Golden algae is also commonly used to refer to a single species, '' Prymnesium parvum'', which causes fish kills. The Chrysophyceae should not be confused with the Chrysophyta, which is a more ambiguous taxon. Although "chrysophytes" is the anglicization of "Chrysophyta", it generally refers to the Chrysophyceae. Members Originally they were taken to include all such forms of the diatoms and multicellular brown algae, but since then they have been divided into several different groups (e.g., Haptophyceae, Synurophyceae) based on pigmentation and cell structure. Some heterotrophic flagellates as the bicosoecids and choanoflagellates were sometimes seen as related to golden algae too. They are now usually restricted to a core group of closely related forms, distinguished primarily by the structure of the flagella in motile cel ...
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Algae
Algae ( , ; : alga ) are any of a large and diverse group of photosynthetic, eukaryotic organisms. The name is an informal term for a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular microalgae, such as '' Chlorella'', '' Prototheca'' and the diatoms, to multicellular forms, such as the giant kelp, a large brown alga which may grow up to in length. Most are aquatic and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds, while the most complex freshwater forms are the '' Charophyta'', a division of green algae which includes, for example, '' Spirogyra'' and stoneworts. Algae that are carried by water are plankton, specifically phytoplankton. Algae constitute a polyphyletic group since they do not include a common ancestor, and although their plastids seem to have a single ori ...
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Protists
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and ''protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, b ...
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Xanthophyceae
Yellow-green algae or the Xanthophyceae (xanthophytes) are an important group of heterokont algae. Most live in fresh water, but some are found in marine and soil habitats. They vary from single-celled flagellates to simple colonial and filamentous forms. Xanthophyte chloroplasts contain the photosynthetic pigments chlorophyll ''a'', chlorophyll ''c'', β-carotene, and the carotenoid diadinoxanthin. Unlike other heterokonts, their chloroplasts do not contain fucoxanthin, which accounts for their lighter colour. Their storage polysaccharide is chrysolaminarin. Xanthophyte cell walls are produced of cellulose and hemicellulose. They appear to be the closest relatives of the brown algae. Classifications The species now placed in the Xanthophyceae were formerly included in the Chlorophyceae. In 1899, Lüther created the group Heterokontae for green algae with unequal flagella. Pascher (1914) included the Heterokontae in the Chrysophyta. In 1930, Allorge renamed the group ...
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Sagenista
Sagenista is a group of heterokonts containing the labyrinthulids and Eogyrea, a class of yet uncultured protists. Originally, it contained the Labyrinthulids and bicosoecids. However at present the bicosoecids have been removed, and Eogyrea were added, in order to make the group monophyletic. Some have a special organelle called a bothrosome (or sagenogenetosome). It is usually found in a marine environments rich in algae and sea grass. It is capable of movement by use of this organelle. They are generally decomposers. They are cultivated for their active production of Omega-3 fatty acids. These acids are used as an approved additive for animal feed. There is a debate about whether some species of Sagenista contains a photosynthetic pigment labeled chlorophyll C. Bothrosome They are capable of excreting an extoplasmic net of filaments for cells to glide upon. These tiny filaments provide a network for cells to travel upon to soak up nutrients from the surrounding env ...
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Phytophthora
''Phytophthora'' (from Greek (''phytón''), "plant" and (), "destruction"; "the plant-destroyer") is a genus of plant-damaging oomycetes (water molds), whose member species are capable of causing enormous economic losses on crops worldwide, as well as environmental damage in natural ecosystems. As well as impacting large scale agriculture, ''Phytophthora'' is a nuisance to garden and indoor plant hobbyists as well as bonsai artists. The cell wall of ''Phytophthora'' is made up of cellulose. The genus was first described by Heinrich Anton de Bary in 1875. Approximately 170 species have been described, although 100–500 undiscovered ''Phytophthora'' species are estimated to exist. Pathogenicity ''Phytophthora'' spp. are mostly pathogens of dicotyledons, and many are relatively host-specific parasites. ''Phytophthora cinnamomi'', though, infects thousands of species ranging from club mosses, ferns, cycads, conifers, grasses, lilies, to members of many dicotyledonous fam ...
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