Golden Shovel-nosed Salamander
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Golden Shovel-nosed Salamander
The golden shovel-nosed salamander (''Desmognathus aureatus'') is a species of salamander in the family Plethodontidae. It is endemic to the United States. It was described in 1956 but was later reclassified as synonymous with the shovelnose salamander (''D. marmoratus''). However, a 2009 mtDNA study found sufficient genetic divergence to again classify as its own species. It is not recognized by other taxonomic authorities due to it being distinguished by mtDNA studies and problems with mtDNA introgression in the genus '' Desmognathus'', but it is still recognized by Darrel Frost pending publication of counter-evidence. It is found in the Chattahoochee The Chattahoochee River forms the southern half of the Alabama and Georgia (U.S. state), Georgia border, as well as a portion of the Florida - Georgia border. It is a tributary of the Apalachicola River, a relatively short river formed by the con ..., Tallulah, and Chattooga river basins of Georgia and adjacent parts o ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Salamander
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela. Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders rarely have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults. This group of amphibians is capable of regenerating lost lim ...
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Plethodontidae
Plethodontidae, or lungless salamanders, are a family of salamanders. Most species are native to the Western Hemisphere, from British Columbia to Brazil, although a few species are found in Sardinia, Europe south of the Alps, and South Korea. In terms of number of species, they are by far the largest group of salamanders. Biology Adult lungless salamanders have four limbs, with four toes on the fore limbs, and usually with five on the hind limbs. Within many species, mating and reproduction occur solely on land. Accordingly, many species also lack an aquatic larval stage, a phenomenon known as direct development in which the offspring hatch as fully-formed, miniature adults. Direct development is correlated with changes in the developmental characteristics of plethodontids compared to other families of salamanders including increases in egg size and duration of embryonic development. Additionally, the evolutionary loss of the aquatic larval stage is related to a diminishing dep ...
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Endemism
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. For example, the Cape sugarbird is found exclusively in southwestern South Africa and is therefore said to be ''endemic'' to that particular part of the world. An endemic species can be also be referred to as an ''endemism'' or in scientific literature as an ''endemite''. For example '' Cytisus aeolicus'' is an endemite of the Italian flora. '' Adzharia renschi'' was once believed to be an endemite of the Caucasus, but it was later discovered to be a non-indigenous species from South America belonging to a different genus. The extreme opposite of an endemic species is one with a cosmopolitan distribution, having a global or widespread range. A rare alternative term for a species that is endemic is "precinctive", which applies to ...
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United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territories, nine Minor Outlying Islands, and 326 Indian reservations. The United States is also in free association with three Pacific Island sovereign states: the Federated States of Micronesia, the Marshall Islands, and the Republic of Palau. It is the world's third-largest country by both land and total area. It shares land borders with Canada to its north and with Mexico to its south and has maritime borders with the Bahamas, Cuba, Russia, and other nations. With a population of over 333 million, it is the most populous country in the Americas and the third most populous in the world. The national capital of the United States is Washington, D.C. and its most populous city and principal financial center is New York City. Paleo-Americ ...
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Shovelnose Salamander
The shovelnose salamander (''Desmognathus marmoratus'') is a species of salamander in the family Plethodontidae. It is endemic to the United States. Taxonomy The golden shovel-nosed salamander (''D. aureatus'') and black shovel-nosed salamander (''D. melanius'') were both described in 1956, then synonymized with ''D. marmoratus'', but later revived as distinct species in 2009. However, their distinctiveness was quantified using mtDNA; due to heavy mtDNA introgression between ''Desmognathus'' species being a major problem with delineating their taxonomy, both species are not recognized by other taxonomic authorities. However, they are provisionally recognized by Darrel Frost unless proper counter-evidence is released. Description The shovelnose salamander is a robust species, broad with a relatively short tail. It receives its name from the shape of its snout which is more square ended than as seen in other salamanders in its genus. The colour is a dusky brown, grey or black w ...
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Mitochondrial DNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA is only a small portion of the DNA in a eukaryotic cell; most of the DNA can be found in the cell nucleus and, in plants and algae, also in plastids such as chloroplasts. Human mitochondrial DNA was the first significant part of the human genome to be sequenced. This sequencing revealed that the human mtDNA includes 16,569 base pairs and encodes 13 proteins. Since animal mtDNA evolves faster than nuclear genetic markers, it represents a mainstay of phylogenetics and evolutionary biology. It also permits an examination of the relatedness of populations, and so has become important in anthropology and biogeography. Origin Nuclear and mitochondrial DNA are thought to be of separate evolutionary origin, with the mtDNA being derived ...
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Desmognathus
''Desmognathus'' is a genus of lungless salamanders in the family Plethodontidae known as dusky salamanders. They range from Texas to the eastern United States and to south-eastern Canada. Characteristics Species of the genus ''Desmognathus'' have a unique jaw-opening mechanism where the lower jaw is stationary and the skull swings open. There are additional stalked condyles, an atlanto-mandibular ligament, along with other skeletal and musculature features that have evolved to accompany this type of jaw-opening mechanism. Additionally, they are known to exhibit maternal care by brooding over their eggs. Species This genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ... includes the following 30-31 species: References External links Tree of Life: Desmognathus* Further ...
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Darrel Frost
Darrel Richmond Frost (born 1951) is an American herpetologist and systematist. He was previously head curator of herpetology at the American Museum of Natural History, as well as president of both the Society for the Study of Amphibians and Reptiles (1998) and the American Society of Ichthyologists and Herpetologists (2006). Four taxa are named in his honor: the toad genus '' Frostius'' (which includes Frost's toad), the tree frog '' Dendropsophus frosti'', Darrel's chorus frog ''Microhyla darreli'', and Frost's arboreal alligator lizard '' Abronia frosti''. Life Frost became interested in animals after witnessing his father kill a rattlesnake at the age of four. He earned a B.S. in Biology from the University of Arizona in 1973, an M.S. in Zoology from Louisiana State University in 1978, and a Ph.D. in Ecology and Systematics from the University of Kansas in 1988. He became an adjunct professor at Columbia University in 2000. In 1990, Frost was appointed Assistant Curator of H ...
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Chattahoochee River
The Chattahoochee River forms the southern half of the Alabama and Georgia border, as well as a portion of the Florida - Georgia border. It is a tributary of the Apalachicola River, a relatively short river formed by the confluence of the Chattahoochee and Flint Flint, occasionally flintstone, is a sedimentary cryptocrystalline form of the mineral quartz, categorized as the variety of chert that occurs in chalk or marly limestone. Flint was widely used historically to make stone tools and start fir ... rivers and emptying from Florida into Apalachicola Bay in the Gulf of Mexico. The Chattahoochee River is about long. The Chattahoochee, Flint, and Apalachicola rivers together make up the Apalachicola–Chattahoochee–Flint River Basin (ACF River Basin). The Chattahoochee makes up the largest part of the ACF's drainage basin. Course The River source, source of the Chattahoochee River is located in Jacks Gap at the southeastern foot of Jacks Knob, in the very southeaste ...
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Tallulah River
:''This article refers to the river. For the town, see Tallulah Falls, Georgia, for the lake, see Tallulah Falls Lake and for the waterfalls and gorge, see Tallulah Gorge.'' The Tallulah River () is a U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed April 26, 2011 river in Georgia and North Carolina. It begins in Clay County, North Carolina, near Standing Indian Mountain in the Southern Nantahala Wilderness and flows south into Georgia, crossing the state line into Towns County.Mast, M.A., and Turk, J.T., 1999U.S. Geological Survey Circular 1173-A, p.158USGS.gov (accessed October 19, 2006) The river travels through Rabun County and ends in Habersham County. It cuts through the Tallulah Dome rock formation to form the Tallulah Gorge and its several waterfalls (collectively known as Tallulah Falls). The Tallulah River intersects with the Chattooga River to form the Tugaloo River at Lake Tugalo in Habersham County. It joi ...
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Chattooga River
The Chattooga River (also spelled Chatooga, Chatuga, and Chautaga, variant name Guinekelokee River) is the main tributary of the Tugaloo River. Water course The headwaters of the Chattooga River are located southwest of Cashiers, North Carolina, and it stretches U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed April 26, 2011 to its confluence with the Tallulah River within Lake Tugalo, which was created by the Tugalo Dam. The Chattooga begins in southern Jackson County, North Carolina, and flows southwestward between northwestern Oconee County, South Carolina, and eastern Rabun County, Georgia. The "Chattooga" spelling was approved by the US Board on Geographic Names in 1897. The Chattooga and the Tallulah rivers combine to make the Tugaloo River, which is considered to start at the outlet of Lake Tugalo. Downriver from the Tugaloo's confluence with the Seneca River, it is known as the Savannah River below Lake Hart ...
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