Diversification Rates
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Diversification Rates
Diversification rates are the rates at which new species form (the Speciation rate, λ) and living species go extinct (the extinction rate, μ). Diversification rates can be estimated from fossils, data on the species diversity of clades and their ages, or phylogenetic trees. Diversification rates are typically reported on a per-lineage basis (e.g. speciation rate per lineage per unit of time), and refer to the diversification dynamics expected under a birth–death process. A broad range of studies have demonstrated that diversification rates can vary tremendously both through time and across the tree of life. Current research efforts are focused on predicting diversification rates based on aspects of species or their environment. Diversification rates are also subject to various survivorship biases such as the "Push of the past" Methods for estimating diversification rates Fossil time series Diversification rates can be estimated time-series data on fossil occurrences. Wit ...
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Speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within lineages. Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book ''On the Origin of Species''. He also identified sexual selection as a likely mechanism, but found it problematic. There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric speciation, allopatric, peripatric speciation, peripatric, parapatric speciation, parapatric, and sympatric speciation, sympatric. Speciation may also be induced artificially, through animal husbandry, agriculture, or laboratory experiments of speciation, laboratory experiments. Whether genetic drift is a minor or major contributor to speciation is the subject of much ...
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Extinction
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and recover may have been lost before this point. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively. This difficulty leads to phenomena such as Lazarus taxa, where a species presumed extinct abruptly "reappears" (typically in the fossil record) after a period of apparent absence. More than 99% of all species that ever lived on Earth, amounting to over five billion species, are estimated to have died out. It is estimated that there are currently around 8.7 million species of eukaryote globally, and possibly many times more if microorganisms, like bacteria, are included. Notable extinct animal species include non-avian dinosaurs, saber-toothed cats, dodos, m ...
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Fossils
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Clades
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms t ...
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Phylogenetic Trees
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological species or other entities based upon similarities and differences in their physical or genetic characteristics. All life on Earth is part of a single phylogenetic tree, indicating common ancestry. In a ''rooted'' phylogenetic tree, each node with descendants represents the inferred most recent common ancestor of those descendants, and the edge lengths in some trees may be interpreted as time estimates. Each node is called a taxonomic unit. Internal nodes are generally called hypothetical taxonomic units, as they cannot be directly observed. Trees are useful in fields of biology such as bioinformatics, systematics, and phylogenetics. ''Unrooted'' trees illustrate only the relatedness of the leaf nodes and do not require the ancestral root to be ...
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Birth–death Process
The birth–death process (or birth-and-death process) is a special case of continuous-time Markov process where the state transitions are of only two types: "births", which increase the state variable by one and "deaths", which decrease the state by one. The model's name comes from a common application, the use of such models to represent the current size of a population where the transitions are literal births and deaths. Birth–death processes have many applications in demography, queueing theory, performance engineering, epidemiology, biology and other areas. They may be used, for example, to study the evolution of bacteria, the number of people with a disease within a population, or the number of customers in line at the supermarket. When a birth occurs, the process goes from state ''n'' to ''n'' + 1. When a death occurs, the process goes from state ''n'' to state ''n'' − 1. The process is specified by birth rates \_ and death rates \_. Recu ...
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Push Of The Past
The push of the past is a type of survivorship bias associated with evolutionary diversification when extinction is possible. Groups that survive a long time are likely to have “got off to a flying start”, and this statistical bias creates an illusion of a true slow-down of diversification rate through time. Birth–Death modelling in evolutionary studies The evolutionary processes of speciation and extinction can be modelled with a stochastic “ birth–death model” (BDM), which is an important component in the study of macroevolution. A BDM assigns each species a certain probability of splitting (\lambda) or going extinct (\mu) per interval of time. This gives rise to an exponential distribution, with the number of species in a particular clade ''N'' at any time ''t'' given by N_= N_e^, although this expression only gives the expected value when N and t are large (see below). In the special case of there being no extinction, this simplifies to the so-called "Yule pro ...
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Phanerozoic Biodiversity Blank 01
The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when animals first developed hard shells preserved in the fossil record. The time before the Phanerozoic, called the ''Precambrian'', is now divided into the Hadean, Archaean and Proterozoic eons. The time span of the Phanerozoic starts with the sudden appearance of fossilised evidence of a number of animal phyla; the evolution of those phyla into diverse forms; the emergence and development of complex plants; the evolution of fish; the emergence of insects and tetrapods; and the development of modern fauna. Plant life on land appeared in the early Phanerozoic eon. During this time span, tectonic forces which move the continents had collected them into a single landmass known as Pangaea (the most recent supercontinent), which then separated into ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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