Dimorphodontidae
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Dimorphodontidae
Dimorphodontidae (or dimorphodontids) is a group of early "rhamphorhynchoid" pterosaurs named after '' Dimorphodon'', that lived in the Late Triassic to Early Jurassic. While fossils that can be definitively referred to the group are rare, dimorphodontids may have had a broad distribution, with fossils known from the UK, the southwest United States, and possibly Antarctica. Dimorphodontidae was named in 1870 by Harry Govier Seeley (as "Dimorphodontae") with ''Dimorphodon'' as the only known member. In 2003, David Unwin defined a clade Dimorphodontidae, as the group consisting of the last common ancestor of ''Dimorphodon macronyx'' and ''Peteinosaurus zambellii'', and all its descendants.Unwin, D. M. 2003. "On the phylogeny and evolutionary history of pterosaurs". In: Buffetaut, E. & Mazin, J.-M. (eds), ''Evolution and Palaeobiology of Pterosaurs''. Geological Society, London, Special Publications 217: 139-190 However, later studies found that ''Dimorphodon'' may not be closely re ...
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Caelestiventus
''Caelestiventus'' ( , meaning "heavenly wind") is a pterosaur genus from the Late Triassic (Norian or Rhaetian) found in western North America. The type species, ''Caelestiventus hanseni'', honors Robin Hansen, the Bureau of Land Management geologist (Bureau of Land Management, BLM), who facilitated access to the excavation site. ''Caelestiventus'' is important because it is the sole example of a desert-dwelling non-pterodactyloid pterosaur and is 65 million years older than other known desert-dwelling pterosaurs. Additionally, it shows that even the earliest pterosaurs were morphologically and ecologically diverse and that the Dimorphodontidae originated in the Triassic period. Discovery ''Caelestiventus'' was recovered from the Saints & Sinners Quarry of northeastern Utah. The site was discovered in 2007 by Dan Chure and George Engelmann while working on the geology and paleontology of the fossilized dunes of the Nugget Sandstone. In 2015 the discovery of ''Caelestiventus'' wa ...
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Dimorphodon Macronyx
''Dimorphodon'' was a genus of medium-sized pterosaur from the early Jurassic Period. It was named by paleontologist Richard Owen in 1859. ''Dimorphodon'' means "two-form tooth", derived from the Greek (') meaning "two", (') meaning "shape" and (') meaning "tooth", referring to the fact that it had two distinct types of teeth in its jaws – which is comparatively rare among reptiles. ''Dimorphodon'' inhabited Europe. Description The body structure of ''Dimorphodon'' displays many "primitive" characteristics, such as, according to Owen, a very small brain-pan and proportionally short wings. The first phalanx in its flight finger is only slightly longer than its lower arm. The neck was short but strong and flexible and may have had a membranous pouch on the underside. The vertebrae had pneumatic foramina, openings through which the air sacs could reach the hollow interior. ''Dimorphodon'' had an adult body length of long, with a 1.45 metre (4.6 ft) wingspan. The tai ...
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Dimorphodon
''Dimorphodon'' was a genus of medium-sized pterosaur from the early Jurassic Period. It was named by paleontologist Richard Owen in 1859. ''Dimorphodon'' means "two-form tooth", derived from the Greek (') meaning "two", (') meaning "shape" and (') meaning "tooth", referring to the fact that it had two distinct types of teeth in its jaws – which is comparatively rare among reptiles. ''Dimorphodon'' inhabited Europe. Description The body structure of ''Dimorphodon'' displays many "primitive" characteristics, such as, according to Owen, a very small brain-pan and proportionally short wings. The first phalanx in its flight finger is only slightly longer than its lower arm. The neck was short but strong and flexible and may have had a membranous pouch on the underside. The vertebrae had pneumatic foramina, openings through which the air sacs could reach the hollow interior. ''Dimorphodon'' had an adult body length of long, with a 1.45 metre (4.6 ft) wingspan. The tai ...
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Rhamphorhynchoidea
The Rhamphorhynchoidea forms one of the two suborders of pterosaurs and represents an evolutionary grade of primitive members of flying reptiles. This suborder is paraphyletic unlike the Pterodactyloidea, which arose from within the Rhamphorhynchoidea as opposed to a more distant common ancestor. Because it is not a completely natural grouping, Rhamphorhynchoidea is not used as a formal group in most scientific literature, though some pterosaur scientists continue to use it as an informal grouping in popular works, such as ''The Pterosaurs: From Deep Time'' by David Unwin, and in some formal studies. Rhamphorhynchoids were the first pterosaurs to have appeared, in the late Triassic Period (Norian age, about 210 million years agoButler, R.J., Barrett, P.M., and gower, D.J. (2009). "Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs." ''Biology Letters'', 5(4): 557–560. ). Unlike their descendants, the pterodactyloids, most rhamphorhynchoids had teeth and long ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Peteinosaurus Zambellii
''Peteinosaurus'' ( ; meaning "winged lizard") was a prehistoric genus of pterosaur. It lived in the late Triassic period in the late Norian age (about 221 to 210 million years ago), and at a wingspan of around , was one of the smallest and earliest pterosaurs, although other estimates suggest a wingspan of up to . Discovery Three fossils have been found near Cene, Italy. The first fossil, the holotype MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype MCSNB 3359, lacks any diagnostic features of ''Peteinosaurus'' and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third example is MCSNB 3496, another fragmentary skeleton. All specimens are those of subadults and of none has the skull been preserved. Like most pterosaurs, ''Peteinosaurus'' had bones that were strong but very light. ''Peteinosaurus'' is t ...
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Peteinosaurus
''Peteinosaurus'' ( ; meaning "winged lizard") was a prehistoric genus of pterosaur. It lived in the late Triassic period in the late Norian age (about 221 to 210 million years ago), and at a wingspan of around , was one of the smallest and earliest pterosaurs, although other estimates suggest a wingspan of up to . Discovery Three fossils have been found near Cene, Italy. The first fossil, the holotype MCSNB 2886, is fragmentary and disarticulated. The second, the articulated paratype MCSNB 3359, lacks any diagnostic features of ''Peteinosaurus'' and thus might be a different species. This paratype has a long tail (20 cm) made more stiff by long extensions of the vertebrae; this feature is common among pterosaurs of the Triassic. The third example is MCSNB 3496, another fragmentary skeleton. All specimens are those of subadults and of none has the skull been preserved. Like most pterosaurs, ''Peteinosaurus'' had bones that were strong but very light. ''Peteinosaurus'' is t ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Nugget Sandstone
The Nugget Sandstone is a Late Triassic to Early Jurassic geologic formation that outcrops in Colorado, Idaho and Utah, western United States. Fossil theropod tracks have been reported from the formation.Weishampel, et al. (2004). "Dinosaur distribution." Pp. 517-607. Fossil content Intermediate theropod, sphenosuchian, drepanosaurid and sphenodontian remains are known.B. B. Britt, D. J. Chure, G. F. Engelmann and J. D. Shumway. 2016. Rise of the erg—paleontology and paleoenvironments of the Triassic-Jurassic transition in northeastern Utah. Geology of the Intermountain West 3:1-32 ;Other fossils * '' Rhynchosauroides sp.'' * Lepidosauria indet. Ichnofossils * '' Batrachopus sp.'' * '' Brachychirotherium sp.'' * '' Brasilichnium sp.'' * '' Cochlichnus sp.'' * '' Diplichnites sp.'' * '' Grallator (Eubrontes)'' * '' Gwyneddichnium sp.'' * '' Octopodichnus sp.'' * '' Otozoum sp.'' * '' cf. Paleohelcura sp.'' * '' Pterichnus sp.'' * '' Scoyenia sp.'' * '' Treptichnus sp.'' ...
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Pterodactylus
''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of ''Pterodactylus'' have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of ''Pterodactylus'' have tentatively been identified from elsewhere in Europe and in Africa. ''Pterodactylus'' was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, ''Pterodactylus'' had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. ''Pterodactylus'' was a small pterosaur compared to other famo ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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