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Rhamphorhynchoidea
The Rhamphorhynchoidea forms one of the two suborders of pterosaurs and represents an evolutionary grade of primitive members of flying reptiles. This suborder is paraphyletic unlike the Pterodactyloidea, which arose from within the Rhamphorhynchoidea as opposed to a more distant common ancestor. Because it is not a completely natural grouping, Rhamphorhynchoidea is not used as a formal group in most scientific literature, though some pterosaur scientists continue to use it as an informal grouping in popular works, such as ''The Pterosaurs: From Deep Time'' by David Unwin, and in some formal studies. Rhamphorhynchoids were the first pterosaurs to have appeared, in the late Triassic Period (Norian age, about 210 million years agoButler, R.J., Barrett, P.M., and gower, D.J. (2009). "Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs." ''Biology Letters'', 5(4): 557–560. ). Unlike their descendants, the pterodactyloids, most rhamphorhynchoids had teeth and long ...
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Dimorphodontidae
Dimorphodontidae (or dimorphodontids) is a group of early " rhamphorhynchoid" pterosaurs named after ''Dimorphodon'', that lived in the Late Triassic to Early Jurassic. While fossils that can be definitively referred to the group are rare, dimorphodontids may have had a broad distribution, with fossils known from the UK, the southwest United States, and possibly Antarctica. Dimorphodontidae was named in 1870 by Harry Govier Seeley (as "Dimorphodontae") with ''Dimorphodon'' as the only known member. In 2003, David Unwin defined a clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ... Dimorphodontidae, as the group consisting of the last common ancestor of '' Dimorphodon macronyx'' and '' Peteinosaurus zambellii'', and all its descendants.Unwin, D. M. 2003. "On the phylogeny a ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like '' Nyctosaurus'' and '' Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. " Pter ...
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Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. The corresponding series (stratigraphy), series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the Carnian, Norian and Rhaetian Geologic time scale, Ages. Many of the first dinosaurs evolved during the Late Triassic, including ''Plateosaurus'', ''Coelophysis'', and ''Eoraptor''. The Triassic–Jurassic extinction event began during this epoch and is one of the five major mass extinction events of the Earth. Etymology The Triassic was named in 1834 by Friedrich August von Namoh, Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern Germany: the lower Buntsandstein (colourful sandstone'')'', t ...
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Keratin
Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, horns, claws, hooves, and the outer layer of skin among vertebrates. Keratin also protects epithelial cells from damage or stress. Keratin is extremely insoluble in water and organic solvents. Keratin monomers assemble into bundles to form intermediate filaments, which are tough and form strong unmineralized epidermal appendages found in reptiles, birds, amphibians, and mammals. Excessive keratinization participate in fortification of certain tissues such as in horns of cattle and rhinos, and armadillos' osteoderm. The only other biological matter known to approximate the toughness of keratinized tissue is chitin. Keratin comes in two types, the primitive, softer forms found in all vertebrates and harder, derived forms found only ...
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Eudimorphodontia
Eudimorphodontidae is an extinct family of early pterosaurs from the Late Triassic (Late Carnian to Late Rhaetian age) of Europe.Barrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref> It was named by Peter Wellnhofer in 1978 to include '' Eudimorphodon ranzii''. Some phylogenetic analyses suggested that Eudimorphodontidae is a junior synonym of Campylognathoididae,Kellner, A. W. A., (2003): Pterosaur phylogeny and comments on the evolutionary history of the group. pp. 105-137. — ''in'' Buffetaut, E. & Mazin, J.-M., (eds.): ''Evolution and Palaeobiology of Pterosaurs''. Geological Society of London, Special Publications 217, London, 1-347Dalla Vecchia, F.M. (2009). "Anatomy and systematics of the pterosaur ''Carniadactylus'' (gen. n.) ''rosenfeldi'' (Dalla Vecchia, 1995)." ''Rivista Italiana de Paleontologia e Stratigrafia'', 115(2): 159-188. however more comprehensive analyses ...
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Austriadactylus
''Austriadactylus'' is a genus of "rhamphorhynchoid" pterosaur. The fossil remains were unearthed in Late Triassic (middle Norian ageBarrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref>) rocks of Austria. The genus was named in 2002 by Fabio Marco Dalla Vecchia e.a.. The type species is ''Austriadactylus cristatus''. The genus name is derived from Latin ''Austria'' and Greek ''daktylos'', "finger", in reference to the wing finger of pterosaurs. The specific epithet means "crested" in Latin, a reference to the skull crest. The genus is based on holotype SMNS 56342, a crushed partial skeleton on a slab, found in an abandoned mine near Ankerschlag in Tyrol, in the Norian Seefelder Beds. The counterslab has been lost and with it some of the bone. The fossil consists of the skull, lower jaws, some vertebrae, parts of the limbs and pelvic girdle, and the first part of the tail. The elo ...
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Preondactylus
''Preondactylus'' is a genus of long-tailed pterosaurs from the Late Triassic (late Norian)Barrett, P. M., Butler, R. J., Edwards, N. P., & Milner, A. R. (2008). Pterosaur distribution in time and space: an atlas. ''Zitteliana'', 61-107/ref> that inhabited what is now Italy. It contains a single known species, ''Preondactylus buffarinii'', which was discovered by Nando Buffarini in 1982 at the Forni Dolostone near Udine in the Preone valley of the Italian Alps."Preondactylus." In: Cranfield, Ingrid (ed.). ''The Illustrated Directory of Dinosaurs and Other Prehistoric Creatures''. London: Salamander Books, Ltd. Pp. 284-285. Discovery When Buffarini first discovered ''Preondactylus'', the thin slab of bituminous, dolomitic limestone containing the fossil was accidentally broken into pieces while being extracted. After reassembly the rock was cleaned with water by him and his wife and the marl and in it the bone was washed away and lost. All that was left was a negative imprint ...
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Candeleros Formation
The Candeleros Formation (formerly known as the Candeleros Member of the "Río Limay Formation") is a geologic formation that crops out in the Río Negro, Neuquén, and Mendoza provinces of northern Patagonia, Argentina. It is the oldest formation in the Neuquén Group and belongs to the Rio Limay Subgroup. Formerly that subgroup was treated as a formation, and the Candeleros Formation was known as the Candeleros Member.Sánchez ''et al.'', 2006 Description The type locality of the Candeleros Formation is Candeleros Hill in Neuquén Province, after which the formation was named by Wichmann in 1929.Wichmann, 1929 This formation unconformably overlies the Lohan Cura Formation, and it is in turn overlain by the Huincul Formation, also a unit of the Neuquén Group. The sediments of the latter are of lighter greenish and yellow colors and the boundary between the Candeleros and Huincul formations is easily recognizable.Leanza ''et al.'', 2004 The Candeleros Formation is almost ...
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Daohugou Beds
The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates, sandstone, mudstone, and thin coal layers deposited in deltaic and lacustrine environments. The formation dates to the Callovian of the Middle Jurassic to the Oxfordian of the Late Jurassic. The most prominent locality of the Haifanggou Formation are the Daohugou Beds, located near the village of Daohugou in southeastern Inner Mongolia. Other localities include Wuhuaxigou, Chentaizi, Jiangzhangzi, Wubaiding, Guancaishan, Haifenggou, Fanzhangzi, and Zhuanshanzi. Dating Daohugou bed The age of the Daohugou bed has been debated, and a number of studies, using different methodologies, have reached conflicting conclusions. Various papers have placed the fossils here as being anywhere from the Middle Jurassic period (169 million y ...
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Wukongopteridae
Wukongopteridae is a group of basal pterosaurs, found in China and the UK. It contains eight species in five genera, all dated to the Middle to Late Jurassic period, The Wukongopteridae were first named by Wang ''et al.'' in 2009, not yet giving an exact definition.* The clade Wukongopteridae was first defined by Wang ''et al.'' in 2010 as "the most recent common ancestor of '' Wukongopterus lii'' and '' Kunpengopterus sinensis'', and all of its descendants". Description Wukongopterids are characterized by a unique combination of "primitive" and advanced pterosaurian features. While they had long tails and other features characteristic of other "rhamphorhynchoids", they also had distinct pterodactyloid features, such as long vertebrae in the neck and a single skull opening in front of the eyes, the nasoantorbital fenestra (in most "rhamphorhynchoids", the antorbital fenestra and the nasal opening are separate). This feature lead to ''Darwinopterus modularis'' being placed by L ...
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