Chromalveolates
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Chromalveolates
Chromista is a biological kingdom consisting of single-celled and multicellular eukaryotic species that share similar features in their photosynthetic organelles (plastids). It includes all protists whose plastids contain chlorophyll ''c'', such as some algae, diatoms, oomycetes, and protozoans. It is probably a polyphyletic group whose members independently arose as a separate evolutionary group from the common ancestor of all eukaryotes. As it is assumed the last common ancestor already possessed chloroplasts of red algal origin, the non-photosynthetic forms evolved from ancestors able to perform photosynthesis. Their plastids are surrounded by four membranes, and are believed to have been acquired from some red algae. Chromista as a biological kingdom was created by British biologist Thomas Cavalier-Smith in 1981 to differentiate some protists from typical protozoans and plants. According to Cavalier-Smith, the kingdom originally included only algae, but his later analysis in ...
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Chloroplast
A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant and algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, and stores it in the energy-storage molecules ATP and NADPH while freeing oxygen from water in the cells. The ATP and NADPH is then used to make organic molecules from carbon dioxide in a process known as the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from one, in unicellular algae, up to 100 in plants like ''Arabidopsis'' and wheat. A chloroplast is characterized by its two membranes and a high concentration of chlorophyll. Other plastid types, such as the leucoplast and the chromoplast, contain little chlorophyll and do not carry out photosynthesis. Chloroplasts are highly dynamic—they circulat ...
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Eukaryotic
Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacteria and Archaea (both prokaryotes) make up the other two domains. The eukaryotes are usually now regarded as having emerged in the Archaea or as a sister of the Asgard archaea. This implies that there are only two domains of life, Bacteria and Archaea, with eukaryotes incorporated among archaea. Eukaryotes represent a small minority of the number of organisms, but, due to their generally much larger size, their collective global biomass is estimated to be about equal to that of prokaryotes. Eukaryotes emerged approximately 2.3–1.8 billion years ago, during the Proterozoic eon, likely as flagellated phagotrophs. Their name comes from the Greek εὖ (''eu'', "well" or "good") and κάρυον (''karyon'', "nut" or "kernel"). Euka ...
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Diatoms
A diatom (New Latin, Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising several Genus, genera of algae, specifically microalgae, found in the oceans, waterways and soils of the world. Living diatoms make up a significant portion of the Earth's Biomass (ecology), biomass: they generate about 20 to 50 percent of the oxygen produced on the planet each year, take in over 6.7 billion metric tons of silicon each year from the waters in which they live, and constitute nearly half of the organic material found in the oceans. The Protist shell, shells of dead diatoms can reach as much as a half-mile (800 m) deep on the ocean floor, and the entire Amazon basin is fertilized annually by 27 million tons of diatom shell dust transported by transatlantic winds from the African Sahara, much of it from the Bodélé Dep ...
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Kingdom (biology)
In biology, a kingdom is the second highest taxonomic rank, just below domain. Kingdoms are divided into smaller groups called phyla. Traditionally, some textbooks from the United States and Canada used a system of six kingdoms (Animalia, Plantae, Fungi, Protista, Archaea/Archaebacteria, and Bacteria/Eubacteria) while textbooks in Great Britain, India, Greece, Brazil and other countries use five kingdoms only (Animalia, Plantae, Fungi, Protista and Monera). Some recent classifications based on modern cladistics have explicitly abandoned the term ''kingdom'', noting that some traditional kingdoms are not monophyletic, meaning that they do not consist of all the descendants of a common ancestor. The terms ''flora'' (for plants), ''fauna'' (for animals), and, in the 21st century, ''funga'' (for fungi) are also used for life present in a particular region or time. Definition and associated terms When Carl Linnaeus introduced the rank-based system of nomenclature into biology i ...
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Centroheliozoa
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth. Characteristics Individuals are unicellular and spherical, usually around 30–80 μm in diameter, and covered with long radial axopods, narrow cellular projections that capture food and allow mobile forms to move about. A few genera have no cell covering, but most have a gelatinous coat holding scales and spines, produced in special deposition vesicles. These may be organic or siliceous and come in various shapes and sizes. For instance, in '' Raphidiophrys'' the coat extends along the bases of the axopods, covering them with curved spicules that give them a pine-treeish look, and in ''Raphidiocystis'' there are both short cup-shaped spicules and long tubular spicules that are only a little shorter than the axopods. Some other common genera include '' Heterophrys'', ''Actinocystis'', and ...
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Haptophyta
The haptophytes, classified either as the Haptophyta, Haptophytina or Prymnesiophyta (named for ''Prymnesium''), are a clade of algae. The names Haptophyceae or Prymnesiophyceae are sometimes used instead. This ending implies classification at the class rank rather than as a division. Although the phylogenetics of this group has become much better understood in recent years, there remains some dispute over which rank is most appropriate. Characteristics The chloroplasts are pigmented similarly to those of the heterokonts, but the structure of the rest of the cell is different, so it may be that they are a separate line whose chloroplasts are derived from similar red algal endosymbionts. The cells typically have two slightly unequal flagella, both of which are smooth, and a unique organelle called a ''haptonema'', which is superficially similar to a flagellum but differs in the arrangement of microtubules and in its use. The name comes from the Greek ''hapsis'', touch, and ''n ...
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Photosynthesis
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in carbohydrate molecules, such as sugars and starches, which are synthesized from carbon dioxide and water – hence the name ''photosynthesis'', from the Greek ''phōs'' (), "light", and ''synthesis'' (), "putting together". Most plants, algae, and cyanobacteria perform photosynthesis; such organisms are called photoautotrophs. Photosynthesis is largely responsible for producing and maintaining the oxygen content of the Earth's atmosphere, and supplies most of the energy necessary for life on Earth. Although photosynthesis is performed differently by different species, the process always begins when energy from light is absorbed by proteins called reaction centers that contain green chlorophyll (and other colored) pigments/chromoph ...
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Plastid
The plastid (Greek: πλαστός; plastós: formed, molded – plural plastids) is a membrane-bound organelle found in the Cell (biology), cells of plants, algae, and some other eukaryotic organisms. They are considered to be intracellular endosymbiotic cyanobacteria. Examples include chloroplasts (used for photosynthesis), chromoplasts (used for pigment synthesis and storage), and leucoplasts (non-pigmented plastids that can sometimes differentiate). The event which led to permanent endosymbiosis in the Archaeplastida clade (of Embryophyte, land plants, red algae, and green algae) probably occurred with a cyanobiont (a symbiotic cyanobacteria) related to the genus ''Gloeomargarita lithophora, Gloeomargarita'', around 1.5 billion years ago. A later primary endosymbiosis event occurred in photosynthetic ''Paulinella'' amoeboids about 90–140 million years ago. This plastid belongs to the "PS-clade" (of the cyanobacteria genera ''Prochlorococcus'' and ''Synechococcus''). Chloroplas ...
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Protists
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exclusion of other eukaryotes means that protists do not form a natural group, or clade. Therefore, some protists may be more closely related to animals, plants, or fungi than they are to other protists. However, like the groups ''algae'', ''invertebrates'', and ''protozoans'', the biological category ''protist'' is used for convenience. Others classify any unicellular eukaryotic microorganism as a protist. The study of protists is termed protistology. History The classification of a third kingdom separate from animals and plants was first proposed by John Hogg in 1860 as the kingdom Protoctista; in 1866 Ernst Haeckel also proposed a third kingdom Protista as "the kingdom of primitive forms". Originally these also included prokaryotes, but ...
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Chlorophyll C
Chlorophyll ''c'' is a form of chlorophyll found in certain marine algae, including the photosynthetic Chromista (e.g. diatoms and brown algae) and dinoflagellates. It has a blue-green color and is an accessory pigment, particularly significant in its absorption of light in the 447–52 nm wavelength region. Like chlorophyll ''a'' and chlorophyll ''b'', it helps the organism gather light and passes a quanta of excitation energy through the light harvesting antennae to the photosynthetic reaction centre. Chlorophyll ''c'' is unusual because it has a porphyrin ring structure and does not have an isoprenoid tail or a reduced ring D, features typical of the other chlorophylls commonly found in algae and plants. Chlorophyll ''c'' can be further divided into chlorophyll ''c1'', chlorophyll ''c2'', and chlorophyll ''c3'', plus at least eight other more recently found subtypes. Chlorophyll ''c1'' Chlorophyll ''c1'' is a common form of chlorophyll ''c''. It differs from chlo ...
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Cavalier-Smith
Thomas (Tom) Cavalier-Smith, FRS, FRSC, NERC Professorial Fellow (21 October 1942 – 19 March 2021), was a professor of evolutionary biology in the Department of Zoology, at the University of Oxford. His research has led to discovery of a number of unicellular organisms (protists) and advocated for a variety of major taxonomic groups, such as the Chromista, Chromalveolata, Opisthokonta, Rhizaria, and Excavata. He was known for his systems of classification of all organisms. Life and career Cavalier-Smith was born on 21 October 1942 in London. His parents were Mary Maude (née Bratt) and Alan Hailes Spencer Cavalier Smith. He was educated at Norwich School, Gonville and Caius College, Cambridge (MA) and King's College London (PhD). He was under the supervision of Sir John Randall for his PhD thesis between 1964 and 1967; his thesis was entitled "''Organelle Development in'' Chlamydomonas reinhardii". From 1967 to 1969, Cavalier-Smith was a guest investigato ...
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Algae
Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular microalgae, such as ''Chlorella,'' ''Prototheca'' and the diatoms, to multicellular forms, such as the giant kelp, a large brown alga which may grow up to in length. Most are aquatic and autotrophic (they generate food internally) and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds, while the most complex freshwater forms are the ''Charophyta'', a division of green algae which includes, for example, ''Spirogyra'' and stoneworts. No definition of algae is generally accepted. One definition is that algae "have chlorophyll ''a'' as their primary photosynthetic pigment and lack a sterile covering of cells around thei ...
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