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Coleoptera
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard e ...
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Protocoleoptera
The Protocoleoptera are a paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern beetle groups during the following Triassic. Protocoleopterans typically possess prognathous (horizontal) heads, distinctive elytra with regular window punctures, culticles with tubercles or scales, as well as a primitive pattern of ventral sclerites, similar to the modern archostematan families Ommatidae and Cupedidae. They are thought to have been xylophagous and wood boring. Kirejtshuk ''et al.'' (2014) argue that the name "Protocoleoptera" should not be used for the group, as Protocoleoptera was originally proposed for the family Protocoleidae , now considered a member of the extinct order Protelytroptera (a stem-group of the modern Dermaptera Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are ...
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Polyphaga
Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the beetle species so far discovered. Key characteristics of Polyphaga are that the hind coxa (base of the leg) does not divide the first and second abdominal/ventral plates which are known as sternites. Also, the notopleural suture (found under the pronotal shield) is not present. Etymology The name of ''polyphaga'' is derived from two Greek words: , meaning 'many', and , meaning 'to eat', so the suborder is called the “eaters of many things”. Classification The five main infraorders are: * Bostrichiformia — including furniture beetles and skin beetles * Cucujiformia — includes lady beetles, longhorn beetles, weevils, checkered beetles and leaf beetles * Elateriformia — includes click beetles and fireflies * Scarabaeiformia — ...
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Archostemata
The Archostemata are the smallest suborder of beetles, consisting 45 living species in five families. They are an ancient lineage with a number of primitive characteristics. Antennae may be thread-shaped (filiform) or like a string of beads (moniliform). This suborder also contains the only beetles where both sexes are paedogenic, ''Micromalthus debilis''. Modern archostematan beetles are considered rare, but were more diverse during the Mesozoic. The term "Archostemata" is used more broadly by some authors to include both modern archostematans as well as stem-group beetles like " protocoleopterans", which some modern archostematans closely resemble to due to their plesiomorphic morphology. Genetic research suggests that modern archostematans are a monophyletic group. Some genetic studies have recovered archostematans as the sister group of Myxophaga. Taxonomy There are five extant families. * Family Crowsoniellidae Iablokoff-Khnzorian, 1983 * Family Cupedidae Laporte, 1838 * F ...
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Dynastinae
Dynastinae or rhinoceros beetles are a subfamily of the scarab beetle family (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include Hercules beetles, unicorn beetles or horn beetles. Over 1500 species and 225 genera of rhinoceros beetles are known. Many rhinoceros beetles are well known for their unique shapes and large sizes. Some famous species are, for example, the Atlas beetle (''Chalcosoma atlas''), common rhinoceros beetle (''Xylotrupes ulysses''), elephant beetle (''Megasoma elephas''), European rhinoceros beetle (''Oryctes nasicornis''), Hercules beetle (''Dynastes hercules''), Japanese rhinoceros beetle or ''kabutomushi'' (''Allomyrina dichotoma''), ox beetle (''Strategus aloeus'') and the Eastern Hercules beetle (''Dynastes tityus''). Description and ecology The Dynastinae are among the largest of beetles, reaching more than in length, but are completely harmless to humans because they cannot bite or sting. So ...
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List Of Subgroups Of The Order Coleoptera
This article scientific classification, classifies the subgroups of the order Coleoptera (beetles) down to the level of family (biology), families, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order Coleoptera **Suborder †Protocoleoptera ***Superfamily †Tshekardocoleoidea Rohdendorf, 1944 ****Family †Tshekardocoleidae Rohdendorf, 1944 ****Family †Labradorocoleidae Ponomarenko, 1969 ****Family †Oborocoleidae Kukalová, 1969 ***Superfamily †Permocupedoidea Martynov, 1933 ****Family †Permocupedidae Martynov, 1933 ****Family †Taldycupedidae Rohdendorf, 1961 ***Superfamily †Permosynoidea Tillyard, 1924 ****Family †Ademosynidae Ponomarenko, 1968 ****Family †Permosynidae Tillyard, 1924 **Suborder Archostemata ***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family Cupedidae Laporte, 1836 ****F ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate subphylum Vertebrata. Familiar examples of invertebrates include arthropods, mollusks, annelids, echinoderms and cnidarians. The majority of animal species are invertebrates; one estimate puts the figure at 97%. Many invertebrate taxa have a greater number and variety of species than the entire subphylum of Vertebrata. Invertebrates vary widely in size, from 50  μm (0.002 in) rotifers to the 9–10 m (30–33 ft) colossal squid. Some so-called invertebrates, such as the Tunicata and Cephalochordata, are more closely related to vertebrates than to other invertebrates. This makes the invertebrates paraphyletic, so the term has little meaning in taxonomy. Etymology The word "invertebrate" comes from the Latin word ''vertebra'', whi ...
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Ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the system through photosynthesis and is incorporated into plant tissue. By feeding on plants and on one another, animals play an important role in the movement of matter and energy through the system. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter, decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and microbes. Ecosystems are controlled by external and internal factors. External factors such as climate, parent material which forms the soil and topography, control the overall structure of an ecosystem but are not themselves influenced by the ecosystem. ...
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Fungus
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''true f ...
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Colorado Potato Beetle
The Colorado potato beetle (''Leptinotarsa decemlineata''), also known as the Colorado beetle, the ten-striped spearman, the ten-lined potato beetle, or the potato bug, is a major pest of potato crops. It is about long, with a bright yellow/orange body and five bold brown stripes along the length of each of its elytra. Native to the Rocky Mountains, it spread rapidly in potato crops across America and then Europe from 1859 onwards. Taxonomy The Colorado potato beetle was first observed in 1811 by Thomas Nuttall and was formally described in 1824 by American entomologist Thomas Say.see pp. 453–454: "Doryphora, Illig.: D. 10-lineata"./ref> The beetles were collected in the Rocky Mountains, where they were feeding on the buffalo bur, '' Solanum rostratum''. The genus ''Leptinotarsa'' is assigned to the chrysolmelid beetle tribe Chrysomelini (in subfamily Chrysomelinae). Description Adult beetles typically are in length and in width. They weigh 50-170 mg. The beetles ar ...
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Elytron
An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alternatively spelled as "hemielytra"), and in most species only the basal half is thickened while the apex is membranous, but when they are entirely thickened the condition is referred to as "coleopteroid". An elytron is sometimes also referred to as a shard. Description The elytra primarily serve as protective wing-cases for the hindwings underneath, which are used for flying. To fly, a beetle typically opens the elytra and then extends the hindwings, flying while still holding the elytra open, though many beetles in the families Scarabaeidae and Buprestidae can fly with the elytra closed (e.g., most Cetoniinae; ). In a number of groups, the elytra are reduced to various degrees, (e.g., the beetle families Staphylinidae and Ripiphoridae), o ...
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Coccinellidae
Coccinellidae () is a widespread family of small beetles ranging in size from . They are commonly known as ladybugs in North America and ladybirds in Great Britain. Some entomologists prefer the names ladybird beetles or lady beetles as they are not true bugs. Many of the species have conspicuous aposematic colours and patterns, such as red with black spots, that warn potential predators that they are distasteful. The majority of the more than 6,000 described species are generally considered beneficial insects, because many prey on herbivorous hemipterans such as aphids or scale insects, which are agricultural pests. Many coccinellids lay their eggs directly in aphid and scale insect colonies, ensuring their larvae have an immediate food source. However, some species such as the herbivorous Mexican bean beetle are agricultural pests. Etymology The name ''coccinellids'', created by Pierre André Latreille, is derived from the Latin word ''coccineus'' meaning "scarlet". The na ...
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