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Coccosteidae
Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below: Genera '' Belgiosteus'' A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China. '' Clarkosteus'' '' Coccosteus'' The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America. '' Dickosteus'' '' Jiuchengia'' The earliest known coccosteid from Late Emsian Yunnan province Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The p ...
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Coccosteus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''C ...
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Watsonosteus
''Watsonosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. Phylogeny ''Watsonosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ... of ''Watsonosteus'': References {{Taxonbar, from=Q112327223, from2=Q97371185 Coccosteidae Givetian life Fossils of Scotland Animals described in 1932 ...
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Protitanichthys Rockportensis
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged Delaware ...
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Watsonosteus Fletti
''Watsonosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. Phylogeny ''Watsonosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ... of ''Watsonosteus'': References {{Taxonbar, from=Q112327223, from2=Q97371185 Coccosteidae Givetian life Fossils of Scotland Animals described in 1932 ...
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Dickosteus Threiplandi
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Coccosteus Cuspidatus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Co ...
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Dickosteus
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Millerosteus Minor
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Millerosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological s ... of ''Millerosteus'': References {{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 ...
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Millerosteus
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Millerosteus'' is a member of the family (biology), family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Millerosteus'': References

{{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 ...
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Protitanichthys
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged Delaware Lim ...
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Coccosteoidea
Coccosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References {{Taxonbar, from=Q30971248 Arthrodires ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below: References Arthrodires {{Placoderm-stub ...
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