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Protitanichthys
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged D ...
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Protitanichthys Fossatus
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged D ...
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Coccosteid
Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below: Genera '' Belgiosteus'' A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China. '' Clarkosteus'' ''Coccosteus'' The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America. ''Dickosteus'' ''Jiuchengia'' The earliest known coccosteid from Late Emsian Yunnan province, China. It is distinguished from other coccosteids by having an elongated occipital. '' Livosteus'' A genus of very large coccosteids known from Middle to Late Devonian strata of Eastern Europe. ''Millerosteus'' A genus of very small ...
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Coccosteidae
Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below: Genera '' Belgiosteus'' A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China. '' Clarkosteus'' ''Coccosteus'' The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America. ''Dickosteus'' ''Jiuchengia'' The earliest known coccosteid from Late Emsian Yunnan province, China. It is distinguished from other coccosteids by having an elongated occipital. '' Livosteus'' A genus of very large coccosteids known from Middle to Late Devonian strata of Eastern Europe. ''Millerosteus'' A genus of very smal ...
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Rockport Quarry Limestone
The Rockport Quarry Limestone is a geologic formation in Michigan. It preserves fossils dating back to the middle Devonian period. Fossil content Vertebrates The Devonian vertebrate fauna of the Rockport Quarry Limestone is the most diverse in Michigan. Acanthodians Cartilaginous fish Placoderms Invertebrates Brachiopods Bryozoans Crustaceans Echinoderms Molluscs Sponges Trilobitomorphs Plants and algae Cyanophytes Foraminifera See also * List of fossiliferous stratigraphic units in Michigan This article contains a list of fossil-bearing stratigraphic units in the state of Michigan, U.S. Sites See also * Paleontology in Michigan References * {{DEFAULTSORT:Fossiliferous stratigraphic units in Michigan Michigan Stratigraphi ... References * Devonian Michigan Middle Devonian Series {{Michigan-geologic-formation-stub ...
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Coccosteus Cuspidatus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Cocc ...
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Millerosteus Minor
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Millerosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Millerosteus'': References {{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 ...
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Coccosteoidea
Coccosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References {{Taxonbar, from=Q30971248 Arthrodires ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References Arthrodires {{Placoderm-stub ...
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Dickosteus Threiplandi
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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