Coccosteoidea
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Coccosteoidea
Coccosteoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References {{Taxonbar, from=Q30971248 Arthrodires ...
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Coccosteomorphi
Coccosteomorphi is an extinct clade of arthrodire placoderms within the Eubrachythoraci (of the suborder Brachythoraci), armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Coccosteomorphi is the sister taxon to Pachyosteomorphi, which together are the two main sub-clades of Eubrachythoraci. Coccosteomorphi can be further sub-divided into Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References Arthrodires {{Placoderm-stub ...
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Panxiosteidae
Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. Phylogeny The family Panxiosteidae was erected by Wang in 1979. Members of the family are noted for showing morphologically intermediate traits between coccosteids and dunkleosteids. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of Panxiosteidae from the 2013 Zhu & Zhu study is shown in the cladogram below: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North Ame ...
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Plourdosteus Canadensis
''Plourdosteus'' is an extinct genus of placoderm arthrodire which was relatively widespread in Euramerica during the Givetian to Frasnian ages of the Devonian. Etymology The name ''Plourdosteus'' commemorates the Plourde family at Miguasha National Park. Taxonomy ''Plourdosteus'' was previously assigned to the family Plourdosteidae within the Coccosteomorphi. However, subsequent studies found the family Plourdosteidae to be polyphyletic and should be dismissed. ''Plourdosteus'' was then proposed to be a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Plourdosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Pa ...
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Panxiosteus Ocullus
''Panxiosteus'' is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Givetian stage of Yunnan province, China. Phylogeny ''Panxiosteus'' is a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Panxiosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clad ...
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Incisoscutoidea
Incisoscutoidea is an extinct superfamily of arthrodire placoderms that lived during the Devonian period. Phylogeny Eubrachythoraci is divided into the sister clades Pachyosteomorphi and Coccosteomorphi, the latter of which can be further sub-divided into the two sister superfamilies Coccosteoidea and Incisoscutoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... below: References {{Taxonbar, from=Q33189516 Arthrodires ...
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Janiosteus Timanicus
''Janiosteus'' is an extinct monospecific genus of placoderm arthrodire from the Middle Devonian: Late Givetian stage found in Timan, Russia. Phylogeny ''Janiosteus'' is a member of Panxiosteidae. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of ''Janiosteus'' within Panxiosteidae is shown in the cladogram below from the 2013 Zhu & Zhu study: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade ...
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Coccosteus Cuspidatus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Cocc ...
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Harrytoombsia Elegans
''Harrytoombsia'' is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found from Kimberley, Australia. It was a fast-swimming predator, and it had sharp blades of enamel-like material around the outer edge of the jaw for catching prey. Phylogeny ''Harrytoombsia'' is a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Harrytoombsia'': References Arthrodires Fossil taxa described in 1979 {{Placoderm-stub ...
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Torosteus Pulchellus
''Torosteus'' is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia. Phylogeny ''Torosteus'' was originally classified as a member of the family Plourdosteidae. However, phylogenetic analysis later found Plourdosteidae to be an invalid grouping, and the family was dismissed. ''Torosteus'' is now considered to be a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Torosteus'': References Arthrodires {{Placoderm-stub ...
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Torosteus Tuberculatus
''Torosteus'' is an extinct genus of arthrodire placoderm from the Early Frasnian stage of the Late Devonian period. Fossils are found in the Kimberley region of Australia. Phylogeny ''Torosteus'' was originally classified as a member of the family Plourdosteidae. However, phylogenetic analysis later found Plourdosteidae to be an invalid grouping, and the family was dismissed. ''Torosteus'' is now considered to be a member of the superfamily Incisoscutoidea, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Torosteus'': References Arthrodires {{Placoderm-stub ...
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Watsonosteus Fletti
''Watsonosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. Phylogeny ''Watsonosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Watsonosteus'': References {{Taxonbar, from=Q112327223, from2=Q97371185 Coccosteidae Givetian life Fossils of Scotland Animals described in 1932 ...
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Mcnamaraspis Kaprios
''Mcnamaraspis'' is an extinct monospecific genus of arthrodire placoderm that inhabited the ancient reef system of north Western Australia during the Frasnian epoch of the Late Devonian period (c. 380-375 million years ago). The type specimen was found and described by John A. Long from the Gogo Formation near Fitzroy Crossing. This fossil fish showed new anatomical features in arthrodires, like the well-preserved annular (ring-shaped) cartilages of the snout, previously inferred to be present by Erik Stensiö of Sweden. It is occasionally referred to as "The Gogo Fish" after the locale the holotype was excavated from. On 5 December 1995, the type species, ''M. kaprios'', was officially proclaimed as the State Fossil emblem of Western Australia by the Governor of Western Australia, thus becoming the first official state fossil emblem for any state of Australia. Description ''M. kaprios'' is an extinct marine fish that grew up to in length. With its large eyes, articulated ja ...
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