Apterygota
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Apterygota
The name Apterygota is sometimes applied to a subclass of small, agile insects, distinguished from other insects by their lack of wings in the present and in their evolutionary history; notable examples are the silverfish, the firebrat, and the jumping bristletails. Their first known occurrence in the fossil record is during the Devonian period, 417–354 million years ago. The group Apterygota is not a clade; it is paraphyletic, and not universally recognized. As defined, the group contains two separate clades of wingless insects: Archaeognatha comprises jumping bristletails, while Zygentoma comprises silverfish and firebrats. The Zygentoma are in the clade Dicondylia with winged insects, a clade that includes all other insects, while Archaeognatha is sister to this lineage.A. Blanke, M. Koch, B. Wipfler, F. Wilde, B. Misof (2014) Head morphology of ''Tricholepidion gertschi'' indicates monophyletic Zygentoma. Frontiers in Zoology 11:16 doi:10.1186/1742-9994-11-16 The nymphs ...
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Pterygota
The Pterygota ( grc, πτερυγωτός, pterugōtós, winged) are a subclass of insects that includes the winged insects. It also includes insect orders that are secondarily wingless (that is, insect groups whose ancestors once had wings but that have lost them as a result of subsequent evolution). The pterygotan group comprises almost all insects. The insect orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma (silverfishes and firebrats), two primitively wingless insect orders. Also not included is Entognatha, which consist of three orders no longer considered to be insects: Protura, Collembola, and Diplura. Systematics Traditionally, this group was divided into the infraclasses Paleoptera and Neoptera. The former are nowadays strongly suspected of being paraphyletic, and better treatments (such as dividing or dissolving the group) are presently being discussed. In addition, it is not clear how exactly the neopterans are related among each o ...
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Species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. Other ways of defining species include their karyotype, DNA sequence, morphology, behaviour or ecological niche. In addition, paleontologists use the concept of the chronospecies since fossil reproduction cannot be examined. The most recent rigorous estimate for the total number of species of eukaryotes is between 8 and 8.7 million. However, only about 14% of these had been described by 2011. All species (except viruses) are given a two-part name, a "binomial". The first part of a binomial is the genus to which the species belongs. The second part is called the specific name or the specific epithet (in botanical nomenclature, also sometimes i ...
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Entognatha
The Entognatha are a class (biology), class of wingless and Ametabolism, ametabolous arthropods, which, together with the insects, makes up the subphylum Hexapoda. Their Arthropod mouthparts, mouthparts are entognathous, meaning that they are retracted within the head, unlike the insects. Entognatha are apterous, meaning that they lack insect wing, wings. The class contains three Order (biology), orders: Collembola (springtails, 9000 species), Diplura (“two-tail”, 1000 species) and Protura (“first-tail”, 800 species). These three groups were historically united with the now-obsolete order Thysanura to form the class Apterygota, but it has since been recognized that the hexapodous condition of these animals has evolved independently from that of insects, and independently ''within'' each order. The orders may not be closely related, and Entognatha is now considered to be a Polyphyly, polyphyletic group. Morphology These minute arthropods are apterous, unlike some orders o ...
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Cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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Thysanura
Thysanura is the now deprecated name of what was, for over a century, recognised as an order in the class Insecta. The two constituent groups within the former order, the Archaeognatha (jumping bristletails) and the Zygentoma (silverfish and firebrats), share several characteristics, such as of having three long caudal filaments, the lateral ones being the cerci, while the one between (telson) is a medial cerciform appendage, specifically an epiproct. They are also both wingless, and have bodies covered with fine scales, rather like the scales of the practically unrelated Lepidoptera. In the late 20th century, it was recognized that the two suborders were not sister taxa, therefore Thysanura was paraphyletic, and the two suborders were each raised to the status of an independent monophyletic order, with Archaeognatha sister taxon to the Dicondylia, including the Zygentoma. Although the group Thysanura is no longer recognized, the name still appears in some published material. An ...
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Diplura
The order Diplura ("two-pronged bristletails") is one of three orders of non-insect hexapods within the class Entognatha (alongside Collembola (springtails) and Protura). The name "diplura", or "two tails", refers to the characteristic pair of caudal appendages or filaments at the terminal end of the body. Around 800 species of diplurans have been described, of which around 170 occur in North America and 12 in Great Britain. Anatomy Diplurans are typically long, with most falling between . However, some species of ''Japyx'' may reach . They have no eyes and, apart from the darkened cerci in some species, they are unpigmented. Diplurans have long antennae with 10 or more bead-like segments projecting forward from the head. The abdomens of diplurans bear eversible vesicles, which seem to absorb moisture from the environment and help with the animal's water balance. The body segments themselves may display several types of setae, or scales and setae. Diplurans possess a chara ...
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Protura
The Protura, or proturans, and sometimes nicknamed coneheads, are very small (0.6-1.5mm long), soil-dwelling animals, so inconspicuous they were not noticed until the 20th century. The Protura constitute an order of hexapods that were previously regarded as insects, and sometimes treated as a class in their own right. Some evidence indicates the Protura are basal to all other hexapods, although not all researchers consider them Hexapoda, rendering the monophyly of Hexapoda unsettled. Uniquely among hexapods, proturans show anamorphic development, whereby body segments are added during moults. There are close to 800 species, described in seven families. Nearly 300 species are contained in a single genus, ''Eosentomon''. Morphology Proturans have no eyes, wings, or antennae, and, lacking pigmentation, are usually white or pale brown. The sensory function of the antennae is fulfilled by the first of three pairs of five-segmented legs, which are held up, pointing forward and have ...
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Collembola
Springtails (Collembola) form the largest of the three lineages of modern hexapods that are no longer considered insects (the other two are the Protura and Diplura). Although the three orders are sometimes grouped together in a class called Entognatha because they have internal mouthparts, they do not appear to be any more closely related to one another than they are to all insects, which have external mouthparts. Collembolans are omnivorous, free-living organisms that prefer moist conditions. They do not directly engage in the decomposition of organic matter, but contribute to it indirectly through the fragmentation of organic matter and the control of soil microbial communities. The word ''Collembola'' is from the ancient Greek "glue" and "peg"; this name was given due to the existence of the collophore, which was previously thought to stick to surfaces to stabilize the creature. Some DNA sequence studies suggest that Collembola represent a separate evolutionary line fro ...
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Cercus
Cerci (singular cercus) are paired appendages on the rear-most segments of many arthropods, including insects and symphylans. Many forms of cerci serve as sensory organs, but some serve as pinching weapons or as organs of copulation. In many insects, they simply may be functionless vestigial structures. In basal arthropods, such as silverfish, the cerci originate from the eleventh abdominal segment. As segment eleven is reduced or absent in the majority of arthropods, in such cases, the cerci emerge from the tenth abdominal segment. It is not clear that other structures so named are homologous. In the Symphyla they are associated with spinnerets. Morphology and functions Most cerci are segmented and jointed, or filiform (threadlike), but some take very different forms. Some Diplura, in particular ''Japyx'' species, have large, stout forcipate (pincer-like) cerci that they use in capturing their prey. The Dermaptera, or earwigs, are well known for the forcipate cerci that most o ...
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Abdomen
The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. The area occupied by the abdomen is called the abdominal cavity. In arthropods it is the posterior (anatomy), posterior tagma (biology), tagma of the body; it follows the thorax or cephalothorax. In humans, the abdomen stretches from the thorax at the thoracic diaphragm to the pelvis at the pelvic brim. The pelvic brim stretches from the lumbosacral joint (the intervertebral disc between Lumbar vertebrae, L5 and Vertebra#Sacrum, S1) to the pubic symphysis and is the edge of the pelvic inlet. The space above this inlet and under the thoracic diaphragm is termed the abdominal cavity. The boundary of the abdominal cavity is the abdominal wall in the front and the peritoneal surface at the rear. In vertebrates, the abdomen is a large body c ...
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Instar
An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form. Differences between instars can often be seen in altered body proportions, colors, patterns, changes in the number of body segments or head width. After shedding their exoskeleton (moulting), the juvenile arthropods continue in their life cycle until they either pupate or moult again. The instar period of growth is fixed; however, in some insects, like the salvinia stem-borer moth, the number of instars depends on early larval nutrition. Some arthropods can continue to moult after sexual maturity, but the stages between these subsequent moults are generally not called instars. For most insect species, an ''instar'' is the developmental stage of the larval forms of holometabolous (complete metamorphism) or nymphal forms o ...
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Metamorphosis
Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some insects, fish, amphibians, mollusks, crustaceans, cnidarians, echinoderms, and tunicates undergo metamorphosis, which is often accompanied by a change of nutrition source or behavior. Animals can be divided into species that undergo complete metamorphosis (" holometaboly"), incomplete metamorphosis ("hemimetaboly"), or no metamorphosis (" ametaboly"). Scientific usage of the term is technically precise, and it is not applied to general aspects of cell growth, including rapid growth spurts. Generally organisms with a larva stage undergo metamorphosis, and during metamorphosis the organism loses larval characteristics. References to "metamorphosis" in mammals are imprecise and only colloquial, but historically idealist ideas of transformation ...
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