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Aculeata
Aculeata is a subclade of Hymenoptera containing ants, bees, and stinging wasps. The name is a reference to the defining feature of the group, which is the modification of the ovipositor into a stinger. However, many members of the group cannot sting, either retaining the ovipositor, or having lost it altogether. A large part of the clade is parasitic. This group includes all of the eusocial Hymenopterans. It is theorized that the possession of a venomous sting was important in the repeated evolution of eusociality within Hymenoptera. The oldest aculeates are known from the Late Jurassic Karabastau Formation of Kazakhstan, represented by the family Bethylonymidae, which may be para or polyphyletic. Classification The use of the name Aculeata has a long history at the rank of infraorder or division. The Aculeata are a monophyletic, or good natural group, containing all the descendants of a single common ancestor. The Aculeata are therefore maintained as a taxon, either at infr ...
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Apocrita
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell provisio ...
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Parasitic Wasp
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders. Parasitoid wasp species differ in which host life-stage they attack: eggs, larvae, pupae, or adults. They mainly follow one of two major strategies within parasitism: either they are endoparasitic, developing inside the host, and koinobiont, allowing the host to continue to feed, develop, and moult; or they are ectoparasitic, developing outside the host, and idiobiont, paralysing the host immediately. Some endoparasitic wasps of the superfamily Ichneumonoidea have a mutualistic relationship with polydnaviruses, the viru ...
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Hymenoptera
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Parasitoid wasp, parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis (biology), metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek language, Ancient Greek wikt:πτερόν, πτερόν (''pteron'') for wing. The Ancient Greek wikt:ὑμήν, ὑμήν (''hymen'') for membrane provides a plausible etymology for the term bec ...
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Chrysidoidea
The superfamily Chrysidoidea is a very large cosmopolitan group (some 6,000 described species, and many more undescribed) , all of which are parasitoids or cleptoparasites of other insects. There are three large, common families (Bethylidae, Chrysididae, and Dryinidae) and four small, rare families (Embolemidae, Plumariidae, Sclerogibbidae, and Scolebythidae). Most species are small (7 mm or less), almost never exceeding 15 mm. This superfamily is traditionally considered to be the basal taxon within the Aculeata, and, as such, some species can sting, though the venom is harmless to humans. Members of the families Dryinidae and Embolemidae are the only parasitoids among the Hymenoptera to have a life cycle in which the wasp larva begins its life inside the body of the host, and then later forms a sac (called a ''thylacium'') that protrudes out of the host's abdomen. The closely related family Sclerogibbidae contains more traditional ectoparasitoids, attacking the nym ...
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Apoidea
The superfamily (zoology), superfamily Apoidea is a major group within the Hymenoptera, which includes two traditionally recognized lineages, the "sphecidae, sphecoid" wasps, and the bees. Molecular phylogeny demonstrates that the bees arose from within the traditional "Crabronidae", so that grouping is paraphyletic, and this has led to a reclassification to produce monophyletic families.Manuela Sann, Oliver Niehuis, Ralph S. Peters, Christoph Mayer, Alexey Kozlov, Lars Podsiadlowski, Sarah Bank, Karen Meusemann, Bernhard Misof, Christoph Bleidorn and Michael Ohl (2018) Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. ''BMC Evolutionary Biology'' 18:71. doi:10.1186/s12862-018-1155-8 Nomenclature Bees appear in recent classifications to be a specialized lineage of "Crabronidae, crabronid" wasps that switched to the use of pollen and nectar as larval food, rather than insect prey; this makes the traditional "Crabronidae" a paraphyletic group. Accordingl ...
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Hymenopteran
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Parasitoid wasp, parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis (biology), metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek language, Ancient Greek wikt:πτερόν, πτερόν (''pteron'') for wing. The Ancient Greek wikt:ὑμήν, ὑμήν (''hymen'') for membrane provides a plausible etymology for the term bec ...
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Ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typically its form is adapted to functions such as preparing a place for the egg, transmitting the egg, and then placing it properly. For most insects, the organ is used merely to attach the egg to some surface, but for many parasitic species (primarily in wasps and other Hymenoptera), it is a piercing organ as well. Some ovipositors only retract partly when not in use, and the basal part that sticks out is known as the scape, or more specifically oviscape, the word ''scape'' deriving from the Latin word '' scāpus'', meaning "stalk" or "shaft". In insects Grasshoppers use their ovipositors to force a burrow into the earth to receive the eggs. Cicadas pierce the wood of twigs with their ovipositors to insert the eggs. Sawflies slit the ...
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Paraphyly
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synapsids, ...
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Vespoidea
Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea (for scoliid wasps) or Formicoidea (for ants). Vespoidea includes wasps with a large variety of lifestyles: eusocial, social, and solitary habits, predators, scavengers, parasitoids, and some herbivores. Description Vespoid wasp females have antennae with 10 flagellomeres, while males have 11 flagellomeres. The edge of the pronotum reaches or passes the tegula. Many species display some level of sexual dimorphism. Most species have fully developed wings, but some have reduced or absent wings in one or both sexes. As in other Aculeata, only the females are ever capable of stinging. Phylogenetics and taxonomy Research based on four nuclear genes (elongation factor-1α F2 copy, long-wavelength rhodopsin, wingless and the D2–D3 regions of 28S ribosomal RNA—2700 bp in total) sugge ...
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Tiphioidea
Tiphioidea is a suggested superfamily of stinging wasps in the order Hymenoptera. There are three families in Tiphioidea, Bradynobaenidae, Tiphiidae, and Sierolomorphidae. Recent research in molecular phylogenetics has resulted in the reorganization of the infraorder Aculeata, which now contains eight superfamilies: Apoidea, Chrysidoidea, Formicoidea, Pompiloidea, Scolioidea, Tiphioidea, Thynnoidea, and Vespoidea Vespoidea is a superfamily of wasps in the order Hymenoptera, although older taxonomic schemes may vary in this categorization, particularly in whether to recognize the superfamilies Scolioidea (for scoliid wasps) or Formicoidea (for ants). V .... References Further reading * * * External links * Hymenoptera Apocrita superfamilies {{hymenoptera-stub ...
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Parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable strategy, evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation. Among parasitoids, strategies range from living inside the host (''endoparasitism''), allowing it to continue growing before emerging as an adult, to Paralysis, paralysing the host and living outside it (''ectoparasitism''). Hosts can include other parasitoids, resulting in hyperparasitism; in the case of oak galls, up to five levels of parasitism are possible. Some parasitoids Behavior-altering parasite, influence their host's behaviour in ways that favour the propagation of the parasitoid. Parasitoids are found in a variety of Taxon, taxa across the insect superorder Endopterygota, whose compl ...
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Infraorder
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow ...
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