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Onocleaceae
Onocleaceae is a small family of terrestrial ferns in the order Polypodiales. It is placed in the suborder Aspleniineae in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). Alternatively, the family, along with Blechnaceae, may be placed in a very broadly defined family Aspleniaceae as the subfamily Blechnoideae. The family may contain from one to four genera, consisting of five species largely in north temperate climes. The four genera, ''Matteuccia'', ''Onoclea'', ''Onocleopsis'' and '' Pentarhizidium'', may be included under the single genus ''Onoclea''. Description Members of the family have the following characteristics, being distinguished by having strongly dimorphic fronds, with the fertile fronds different from the sterile fronds. The rhizomes are long- to short-creeping to ascending, and sometimes stoloniferous (''Matteuccia'' and ''Onocleopsis''). The leaves are strongly dimorphic and the petioles have two vascular bundles uniting distally into a gutter ...
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Matteuccia
''Matteuccia'' is a genus of ferns with one species: ''Matteuccia struthiopteris'' ( common names ostrich fern, fiddlehead fern, or shuttlecock fern). The species epithet ''struthiopteris'' comes from Ancient Greek words () "ostrich" and () "fern". Description The fronds are dimorphic, with the deciduous green sterile fronds being almost vertical, tall and broad, long-tapering to the base but short-tapering to the tip, so that they resemble ostrich plumes, hence the name. The fertile fronds are shorter, long, brown when ripe, with highly modified and constricted leaf tissue curled over the sporangia; they develop in autumn, persist erect over the winter and release the spores in early spring. Along with ''Dryopteris goldieana'', it is one of the largest species of fern in eastern North America. Classification ''Matteuccia struthiopteris'' is the only species in the genus ''Matteuccia''. Some sources include two Asian species, ''M. orientalis'' and ''M. intermedia'', but mol ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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Molecular Phylogenetics
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tree. History The theoretical frame ...
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Pentarhizidium Intermedium
''Pentarhizidium'' is a genus of two Asian fern species. These species have formerly been included in ''Matteuccia'' or ''Onoclea ''Onoclea'' is a genus of plants in the family Onocleaceae, native to moist habitats in eastern Asia and eastern North America. They are deciduous ferns with sterile fronds arising from creeping rhizomes In botany and dendrology, a rhizome ...''. Recent genetic analysis has determined that these two species form a discrete clade that is basal to the rest of this fern group, and so have been located in their own genus. ''P. orientale'' is sometimes grown as a garden plant. Species References Polypodiales Ferns of Asia Fern genera {{Polypodiales-stub ...
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Himalayas
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 peaks exceeding in elevation lie in the Himalayas. By contrast, the highest peak outside Asia (Aconcagua, in the Andes) is tall. The Himalayas abut or cross five countries: Bhutan, India, Nepal, China, and Pakistan. The sovereignty of the range in the Kashmir region is disputed among India, Pakistan, and China. The Himalayan range is bordered on the northwest by the Karakoram and Hindu Kush ranges, on the north by the Tibetan Plateau, and on the south by the Indo-Gangetic Plain. Some of the world's major rivers, the Indus, the Ganges, and the Tsangpo–Brahmaputra, rise in the vicinity of the Himalayas, and their combined drainage basin is home to some 600 million people; 53 million people live in the Himalayas. The Himalayas have ...
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Pentarhizidium Orientale
''Pentarhizidium orientalis'', the Oriental ostrich fern, is a fern native to China, Japan, and the Himalayas. It grows to about 0.6 m (2 ft) in height by 0.6 m (2 ft) wide. It was formerly included in the genus ''Matteuccia'', but phylogenetic studies mandated that it and ''Pentarhizidium intermedium'' be moved to a new genus. References External links * The Plant List entryPlants for a Future entry* Efloras entry* Robert M. LloydFacultative Apomixis and Polyploidy in ''Matteuccia orientalis'' ''American Fern Journal'', Vol. 63, No. 2 (Apr.–Jun. 1973), pp. 43–48. Published by the American Fern Society The American Fern Society was founded in 1893. Today, it has more than 1,000 members around the world, with various local chapters. Among its deceased members, perhaps the most famous is Oliver Sacks, who became a member in 1993. Willard N. Clut .... Polypodiales {{Polypodiales-stub ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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USDA
The United States Department of Agriculture (USDA) is the federal executive department responsible for developing and executing federal laws related to farming, forestry, rural economic development, and food. It aims to meet the needs of commercial farming and livestock food production, promotes agricultural trade and production, works to assure food safety, protects natural resources, fosters rural communities and works to end hunger in the United States and internationally. It is headed by the Secretary of Agriculture, who reports directly to the President of the United States and is a member of the president's Cabinet. The current secretary is Tom Vilsack, who has served since February 24, 2021. Approximately 80% of the USDA's $141 billion budget goes to the Food and Nutrition Service (FNS) program. The largest component of the FNS budget is the Supplemental Nutrition Assistance Program (formerly known as the Food Stamp program), which is the cornerstone of USDA's ...
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Dryopteridaceae
The Dryopteridaceae are a Family (biology), family of leptosporangiate ferns in the Order (biology), order Polypodiales. They are known Common name, colloquially as the wood ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family is placed in the suborder Polypodiineae. Alternatively, it may be treated as the subfamily Dryopteridoideae of a very broadly defined family Polypodiaceae ''Sensu, sensu lato''. The family contains about 1700 species and has a cosmopolitan distribution. Species may be terrestrial plant, terrestrial, epipetric, hemiepiphyte, hemiepiphytic, or epiphytic. Many are Gardening, cultivated as ornamental plants. The largest genera are ''Elaphoglossum'' (600+), ''Polystichum'' (260), ''Dryopteris'' (225), and ''Ctenitis'' (150). These four genera contain about 70% of the species. Dryopteridaceae Evolutionary radiation, diverged from the other families in eupolypods I about 100 million years ago. Description The rhizomes are ...
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Matteuccia Struthiopteris
''Matteuccia'' is a genus of ferns with one species: ''Matteuccia struthiopteris'' (common names ostrich fern, fiddlehead fern, or shuttlecock fern). The species epithet ''struthiopteris'' comes from Ancient Greek words () "ostrich" and () "fern". Description The fronds are dimorphic, with the deciduous green sterile fronds being almost vertical, tall and broad, long-tapering to the base but short-tapering to the tip, so that they resemble ostrich plumes, hence the name. The fertile fronds are shorter, long, brown when ripe, with highly modified and constricted leaf tissue curled over the sporangia; they develop in autumn, persist erect over the winter and release the spores in early spring. Along with ''Dryopteris goldieana'', it is one of the largest species of fern in eastern North America. Classification ''Matteuccia struthiopteris'' is the only species in the genus ''Matteuccia''. Some sources include two Asian species, ''M. orientalis'' and ''M. intermedia'', but molec ...
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Blechnales
Aspleniineae is a suborder of ferns in the order Polypodiales. It is equivalent to the clade eupolypods II in earlier systems; it is also treated as a single very broadly defined family Aspleniaceae. The suborder generally corresponds with the order Blechnales as described by J. L. Reveal in 1993. Aspleniineae includes some important ferns, including ''Onoclea sensibilis'', the sensitive fern, which grows as a virtual weed throughout much of its temperate North American range, and ferns of the genus ''Thelypteris'', a genus that has shown remarkable speciation. It also includes one of the more common horticultural ferns, ''Matteuccia struthiopteris'', the ostrich fern. Taxonomy In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the group is treated as the suborder Aspleniinae, and divided into 11 families. Alternatively, it may be treated as a single, very broadly circumscribed family Aspleniaceae ''sensu lato ''Sensu'' is a Latin word meaning "in the sense o ...
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Sorus
A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient Greek σωρός (''sōrós'' 'stack, pile, heap'). Structure In lichens and other fungi, the sorus is surrounded by an external layer. In some red algae, it may take the form of depression into the thallus. In ferns, the sori form a yellowish or brownish mass on the edge or underside of a fertile frond. In some species, they are protected during development by a scale or film of tissue called the indusium, which forms an umbrella-like cover. Lifecycle significance Sori occur on the sporophyte generation, the sporangia within producing haploid meiospores. As the sporangia mature, the indusium shrivels so that spore release is unimpeded. The sporangia then burst and release the spores. As an aid to identification The shape, arrangemen ...
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