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Ziphosuchia
Ziphosuchia is a clade of mesoeucrocodylian crocodyliforms that includes notosuchians and sebecosuchians. Systematics First constructed in 2000, it was considered to include ''Notosuchus'', ''Libycosuchus'', and Sebecosuchia. In a 2004 phylogenetic study, it was defined as the most recent common ancestor of ''Notosuchus'', ''Libycosuchus'', and Baurusuchoidea and all of its descendants. Ziphosuchia is often considered to be the sister group of Neosuchia, a clade that includes modern crocodilians. ''Razanandrongobe ''Razanandrongobe'' (meaning "ancestor f thelarge lizard" in Malagasy) is a genus of carnivorous ziphosuchian crocodyliform from the Middle Jurassic of Madagascar. It contains the type and only species ''Razanandrongobe sakalavae'', named in 2 ...'' is the oldest representative of this clade.Dal Sasso C, Pasini G, Fleury G, Maganuco S. (2017) Razanandrongobe sakalavae, a gigantic mesoeucrocodylian from the Middle Jurassic of Madagascar, is the oldest known notosuc ...
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Notosuchia
Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group (see below); if Sebecosuchia is included within Notosuchia its existence is pushed into the Middle Miocene, about 11 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory ('' Chimaerasuchus''), omnivory (''Simosuchus''), and terrestrial hypercarnivory (''Baurusuchus''). It included many members with highly derived traits unusual for crocodylomorphs, including mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (''Armadillosuchus''), and possibly fleshy cheeks and pig-like snouts (''Notosuchus''). The suborder was first named in 1971 by Zulma Gasparini and has since undergone ...
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Notosuchia
Notosuchia is a suborder of primarily Gondwanan mesoeucrocodylian crocodylomorphs that lived during the Jurassic and Cretaceous. Some phylogenies recover Sebecosuchia as a clade within Notosuchia, others as a sister group (see below); if Sebecosuchia is included within Notosuchia its existence is pushed into the Middle Miocene, about 11 million years ago. Fossils have been found from South America, Africa, Asia, and Europe. Notosuchia was a clade of terrestrial crocodilians that evolved a range of feeding behaviours, including herbivory ('' Chimaerasuchus''), omnivory (''Simosuchus''), and terrestrial hypercarnivory (''Baurusuchus''). It included many members with highly derived traits unusual for crocodylomorphs, including mammal-like teeth, flexible bands of shield-like body armor similar to those of armadillos (''Armadillosuchus''), and possibly fleshy cheeks and pig-like snouts (''Notosuchus''). The suborder was first named in 1971 by Zulma Gasparini and has since undergone ...
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Razanandrongobe
''Razanandrongobe'' (meaning "ancestor f thelarge lizard" in Malagasy) is a genus of carnivorous ziphosuchian crocodyliform from the Middle Jurassic of Madagascar. It contains the type and only species ''Razanandrongobe sakalavae'', named in 2004 by Simone Maganuco and colleagues based on isolated bones found in 2003. The remains, which included a fragment of maxilla and teeth, originated from the Bathonian-aged Sakaraha Formation of Mahajanga, Madagascar. While they clearly belonged to a member of the Archosauria, Maganuco and colleagues refrained from assigning the genus to a specific group because the fragmentary remains resembled lineages among both the theropod dinosaurs and crocodylomorphs. Further remains (including a premaxilla and lower jawbone) had been discovered as early as 1972, but were not described until 2017 by Cristiano Dal Sasso and colleagues. These remains allowed them to confidently assign ''Razanandrongobe'' as the oldest-known member of the Notosuchia, ...
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Doratodon
''Doratodon'' is an extinct genus of Late Cretaceous ziphodont crocodylomorph that was once believed to be a dinosaur,Company, J., Suberbiola, X.P., Ruiz-Omeñaca, J.I. & Buscalioni, A.D. (2005). A new species of ''Doratodon'' (Crocodyliformes: Ziphosuchia) from the Late Cretaceous of Spain. ''Journal of Vertebrate Paleontology'' 25(2): 343–353 that could be a sebecosuchian mesoeucrocodylian. Two species of ''Doratodon'' are known to science: ''D. carcharidens'' from Austria (Grünbach Formation) and Hungary (Csehbánya Formation), the type species; and ''D. ibericus'' from Spain (Sierra Perenchiza Formation). The holotype of ''D. carcharidens'' consists of a denture and a mandible.Zoltan Csiki-Sava, Eric Buffetaut, Attila Ősi, Xabier Pereda-Suberbiola, Stephen L. Brusatte (2015)Island life in the Cretaceous - faunal composition, biogeography, evolution, and extinction of land-living vertebrates on the Late Cretaceous European archipelago ''Zookeys" 469: 1-161 doi: 10.3897/zoo ...
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Sebecosuchia
Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in the Miocene with the last sebecids, but ''Razanandrongobe'' pushes the origin of Sebecosuchia to the Middle Jurassic. Fossils have been found primarily from South America but have also been found in Europe, North Africa, Madagascar, and the Indian subcontinent. History and phylogeny Sebecosuchia was first constructed in 1937 by George Gaylord Simpson.Simpson, G.G. 1937. "New reptiles from the Eocene of South America". ''American Museum Novitates'' 967: 1-20 In 1946 the concept was again used by American paleontologist Edwin Colbert to include ''Sebecus'' and Baurusuchidae. ''Sebecus'', which had been known from South America since 1937, was an unusual crocodyliform with a deep snout and teeth that were ziphodont, or serrated and laterally comp ...
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Libycosuchus
''Libycosuchus'' is an extinct genus of North African crocodylomorph possibly related to ''Notosuchus'';Buffetaut, E. 1982. Radiation évolutive, paléoécologie et biogéographie des Crocodiliens mésosuchienes. ''Mémoires Societé Geologique de France'' 142: 1–88. it is part of the monotypic Libycosuchidae and Libycosuchinae. It was terrestrial, living approximately 95 million years ago in the Cenomanian stage of the Late Cretaceous. Fossil remains have been found in the Bahariya Formation in Egypt, making it contemporaneous with the crocodilian ''Stomatosuchus'', and dinosaurs, including ''Spinosaurus''. It was one of the few fossils discovered by Ernst Stromer that wasn't destroyed by the Royal Air Force during the bombing of Munich in 1944. The type species, ''L. brevirostis'', was named in 1914 Stromer 1914, p. 28 and 29, fn. 1 and described in 1915.Original citation: Stromer, E. 1915. Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Ägyptens. II. Wirb ...
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Sebecosuchia
Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in the Miocene with the last sebecids, but ''Razanandrongobe'' pushes the origin of Sebecosuchia to the Middle Jurassic. Fossils have been found primarily from South America but have also been found in Europe, North Africa, Madagascar, and the Indian subcontinent. History and phylogeny Sebecosuchia was first constructed in 1937 by George Gaylord Simpson.Simpson, G.G. 1937. "New reptiles from the Eocene of South America". ''American Museum Novitates'' 967: 1-20 In 1946 the concept was again used by American paleontologist Edwin Colbert to include ''Sebecus'' and Baurusuchidae. ''Sebecus'', which had been known from South America since 1937, was an unusual crocodyliform with a deep snout and teeth that were ziphodont, or serrated and laterally comp ...
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Baurusuchoidea
Sebecosuchia is an extinct group of mesoeucrocodylian crocodyliforms that includes the families Sebecidae and Baurusuchidae. The group was long thought to have first appeared in the Late Cretaceous with the baurusuchids and become extinct in the Miocene with the last sebecids, but ''Razanandrongobe'' pushes the origin of Sebecosuchia to the Middle Jurassic. Fossils have been found primarily from South America but have also been found in Europe, North Africa, Madagascar, and the Indian subcontinent. History and phylogeny Sebecosuchia was first constructed in 1937 by George Gaylord Simpson.Simpson, G.G. 1937. "New reptiles from the Eocene of South America". ''American Museum Novitates'' 967: 1-20 In 1946 the concept was again used by American paleontologist Edwin Colbert to include '' Sebecus'' and Baurusuchidae. ''Sebecus'', which had been known from South America since 1937, was an unusual crocodyliform with a deep snout and teeth that were ziphodont, or serrated and laterally ...
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Sebecia
Sebecia is an extinct clade of mesoeucrocodylian crocodyliforms that includes peirosaurids and sebecids. It was first constructed in 2007 to include ''Hamadasuchus'', Peirosauridae, and ''Sebecus''. It was initially considered to be the sister taxon of the clade Neosuchia, which includes living crocodilians, although some later studies have placed it within Neosuchia as a basal clade. Sebecians were terrestrial crocodyliforms characterized by their deep snouts and ziphodont dentition. They first appeared in the Late Cretaceous, survived the Cretaceous–Paleogene extinction event, and became extinct in the Miocene epoch. According to paleontologists Hans Larsson and Hans-Dieter Sues, who constructed the clade in 2007, Sebecia also includes genera that were once assigned to Baurusuchidae, namely ''Pabwehshi''. However, other baurusuchids, namely ''Baurusuchus'', were placed outside Sebecia. Therefore, Baurusuchidae was considered polyphyletic and thus not a true clade. Below is ...
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Notosuchus
''Notosuchus'' (; 'southern crocodile') is an extinct genus of South American notosuchian crocodylomorph. It was terrestrial, living approximately 85 million years ago in the Santonian stage of the Late Cretaceous. Description ''Notosuchus'' was relatively small, reaching in length and a weight of . Remains have been found in the Bajo de la Carpa Formation in Patagonia, Argentina. First named in 1896, ''Notosuchus'' was the first known notosuchian. The type species is ''N. terrestris''. A second species, ''N. lepidus'', was named in 1957. A paper published in 2008 by Fiorelli and Calvo described new remains of the type species ''N. terrestris''. In it, the authors suggested that the skull would have supported a short trunk, or "hog's snout" as well as fleshy upper and lower lips. The anteriorly directed nares and the absence of a bony nasal septum (which presumably indicates cartilaginous tissue serving its place) provide evidence for a trunk-like snout, while striations on the ...
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Neogene Crocodylomorphs
The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans ('' Homo habilis'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific ...
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Neosuchia
Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives. It is defined as the most inclusive clade containing all crocodylomorphs more closely related to ''Crocodylus niloticus'' (the Nile Crocodile) than to ''Notosuchus terrestris''. Members of Neosuchia generally share a crocodilian-like bodyform adapted to freshwater aquatic life, as opposed to the terrestrial habits of more basal crocodylomorph groups. The earliest neosuchian is suggested to be the Early Jurassic ''Calsoyasuchus'', which lived during the Sinemurian and Pliensbachian stages in North America. It is often identified as a member of Goniopholididae, though this is disputed, and the taxon may lie outside Neosuchia, which places the earliest records of the group in the Middle Jurassic. Characteristics A tooth notch between the maxilla and premaxilla is a basal characteristic of the Neosuchia, although it is lost in some more derived forms, most nota ...
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