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Xenungulata
Xenungulata ("strange ungulates") is an order of extinct and primitive South American hoofed mammals that lived from the Late Paleocene to Early Eocene (Itaboraian to Casamayoran in the SALMA classification). Fossils of the order are known from deposits in Brazil, Argentina, Peru, and Colombia. The best known member of this enigmatic order is the genus ''Carodnia'', a tapir-like and -sized animal with a gait similar to living African elephants. Description Xenungulates are characterized by bilophodont M1–2 and M1–2, similar to pyrotheres, and complex lophate third molars, similar to uintatheres. Though other relationships, to arctocyonids for example, have been suggested, no proofs thereof have been found. The foot bones of xenungulates were short and robust and their digits terminated in broad, flat, and unfissured hoof-like unguals, quite unlike any other meridiungulates. The discovery of ''Etayoa'' in Colombia made it clear that xenungulates are distinct from ot ...
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Carodnia
''Carodnia'' is an extinct genus of South American ungulate known from the Early Eocene of Brazil, Argentina, and Peru. ''Carodnia'' is placed in the order ''Xenungulata'' together with ''Etayoa'' and '' Notoetayoa''. ''Carodnia'' is the largest mammal known from the Eocene of South America. It was heavily built and had large canines and cheek teeth with a crested pattern like the uintatheres to which it can be related. In life, it would have been a tapir-sized animal. It bore strong resemblances to dinoceratans, although without tusks or ossicones. Description Simpson noted that ''Carodnia'' resembles the primitive uintathere '' Probathyopsis''. Although Paula Couto also made the same favorable comparison, he placed ''Carodnia'' in the new order Xenungulata. concluded that ''Probathyopsis'' shares several dental characteristics with ''Carodnia'', but that in the latter the anterior dentition of is more reduced, the second lower and upper premolars are enlarged and pointe ...
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Notoetayoa
''Notoetayoa'' is an extinct genus of mammal, from the order Xenungulata. It contains a single species, ''Notoetayoa gargantuai'', which lived during the Middle Paleocene. Its fossilized remains were discovered in South America. Description This animal is only known from a fragment of its jaw, and it is therefore impossible to precisely reconstruct its appearance. From a comparison with a similar but better known animal, ''Carodnia'', it is possible that ''Notoetayoa'' was an animal with heavy shapes and a powerful body. The size of ''Notoetayoa'' was in any cases smaller than that of ''Carodnia'', and it is possible that it was as large as a small tapir. The lower third molar had a higher trigonid than the thalonid, and the paraconid was well defined, and not fused with the metaconid. Compared to its relative ''Etayoa'', its metachristid was short, and the precingulid was more robust than in ''Etayoa'' and ''Carodnia''. The talonid basin wasn't open lingually as in ''Carodni ...
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Pyrotheria
Pyrotheria is an order of extinct meridiungulate mammals. These mastodon-like ungulates include the genera ''Baguatherium'', ''Carolozittelia'', ''Colombitherium'', ''Griphodon'', ''Propyrotherium'', ''Proticia'', and ''Pyrotherium''. They had the appearance of large, digitigrade, tapir-like mammals with relatively short, slender limbs and five-toed feet with broad, flat phalanges. Their fossils are restricted to Paleocene through Oligocene deposits of Brazil, Peru and Argentina. Some experts place the clade Xenungulata (which contains several genera, including ''Carodnia'') within Pyrotheria, even when dentition, although bilophodont in both orders, is very different. For most scholars, the two orders remain separated. The dentition is complete with strong, procumbent, chisel-shaped incisors, strong sharp-pointed canines, and low-crowned cheek teeth with bilophodont molars. The affinities of the Xenungulata remain uncertain. Affinities with the Dinocerata are strongly supporte ...
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South American Land Mammal Age
The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Mya (unit), Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as ages, stages, or intervals and were established using geographic place names where fossil materials where obtained.Flynn & Swisher, 1995 The basic unit of measure is the first/last boundary statement. This shows that the first appearance event of one taxon is known to predate the last appearance event of another. If two taxa are found in the same fossil quarry or at the same stratigraphic horizon, then their age-range zones overlap. Background South America was an island continent for much of the Cenozoic, or the "Age of Mammals". As a result, its mammals evolved in their own unique directions, as Australia and Madagascar still have today. Paleogeographic timeline A simplified paleogeography, paleogeographic timeline of South ...
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Riochican
The Riochican ( es, Riochiquense) age is a period of geologic time (57.0–54.0 Ma) within the Paleocene and Eocene epochs of the Paleogene, used more specifically within the South American land mammal ages (SALMA). It follows the Peligran and precedes the Itaboraian age. Etymology This age is named after the Río Chico Group of the Golfo San Jorge Basin The Golfo San Jorge Basin ( es, Cuenca del Golfo San Jorge) is a hydrocarbon-rich sedimentary basin located in eastern Patagonia, Argentina. The basin covers the entire San Jorge Gulf and an inland area west of it, having one half located in San .... Formations Fossils References Bibliography ;Bogotá Formation * * * ;Bororó Formation * ;Chota Formation * * ;Las Flores Formation * ;Koluel Kaike Formation * ;Maíz Gordo Formation * ;Mealla Formation * ;Mogollón Formation * * ;Muñani Formation * ;Peñas Coloradas Formation * * ;Río Loro Formation * ;Salamanca Formation * * * ...
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Etayoa
''Etayoa'' is an ungulate of the family Carodniidae in the order Xenungulata that lived during the Early Eocene (~ 55 Ma) in northern South America. Etymology The genus of the type species ''Etayoa bacatensis'' was named by palaeontologist Carlos Villarroel after Fernando Etayo Serna, who contributed extensively to the paleontology and stratigraphy research in Colombia. The species epithet ''bacatensis'' refers to Bacatá,Villarroel, 1987 the name in Muysccubun for the main settlement of the southern Muisca Confederation; the name of which has been used for the current Colombian capital Bogotá, founded in a different location than the original Bacatá. Description The type species fossil specimen consists of a partial mandible with teeth, found in the Bogotá Formation in the locality Ciudad Bolívar of Bogotá, Colombia.1987 Description of ''Etayoa bacatensis''- Paleobiology Database The estimated size of the ungulate is the size of a dog. 2011 Un xenungulado del Paleoc ...
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Meridiungulata
South American native ungulates, commonly abbreviated as SANUs, are extinct ungulate-like mammals of controversial affinities that were indigenous to South America prior to the Great American Biotic Interchange. They comprise five major groups conventionally ranked as orders—Astrapotheria, Litopterna, Notoungulata, Pyrotheria, and Xenungulata—as well as some other taxa, such as Didolodontidae and Kollpaniidae. It has been proposed that some or all of the members of this group form a clade, named Meridiungulata, though the relationships of South American ungulates remain largely unresolved. The two largest groups of South American ungulates, the notoungulates and the litopterns, were the only groups to persist beyond the mid Miocene. Only a few of the largest species of notoungulates and litopterns survived until the end-Pleistocene extinctions. Though most SANUs lived in South America, astrapotheres and litopterns are known from Eocene aged deposits in the Antarctic Peninsu ...
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Itaboraian
The Itaboraian ( pt, Itaboraiense) age is a period within the Early Eocene geologic time (53.0–50.0 Mya (unit), Ma) epoch of the Paleogene, used more specifically with South American land mammal ages (SALMA). It follows the Riochican and precedes the Casamayoran age. Etymology This age is named after the Itaboraí Formation in the Itaboraí Basin in Itaboraí, a Rio de Janeiro metropolitan area, Greater Rio municipality in the state of Rio de Janeiro (state), Rio de Janeiro in Brazil not very far away from Niterói. Formations Fossils Correlations References Bibliography

;Itaboraí Formation * * * * * * ;Bogotá Formation * * * ;Cerrejón Formation * * * * * * * * * * * * * ;Chota Formation * * ;Las Flores Formation * * * ;Koluel Kaike Formation * ;Maíz Gordo Formation * ;Mogollón Formation * * ;Muñani Formation * ;Peñas Coloradas Formation * * ;Río Loro Formation * {{SALMA Itaboraian, Paleocene Sout ...
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Bogotá Formation
The Bogotá Formation ( es, Formación Bogotá, E1-2b, Tpb, Pgb) is a geological formation of the Eastern Hills and Bogotá savanna on the Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The predominantly shale and siltstone formation, with sandstone beds intercalated, dates to the Paleogene period; Upper Paleocene to Lower Eocene epochs, with an age range of 61.66 to 52.5 Ma, spanning the Paleocene–Eocene Thermal Maximum. The thickness of the Bogotá Formation ranges from near Tunja to near Bogotá. Fossils of the ungulate '' Etayoa bacatensis'' have been found in the Bogotá Formation, as well as numerous reptiles, unnamed as of 2017. Etymology The formation was first described by Hettner in 1892,Acosta & Ulloa, 2002, p.59 then by Hubach in 1931, 1945 and 1957, and named in 1963 by Julivert after the Colombian capital Bogotá and its savanna.Montoya & Reyes, 2005, p.57 Description Lithologies The Bogotá Formation consists mainly of grayish-red ...
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Rodcania
''Rodcania'' is an extinct genus of mammal, belonging to the order Xenungulata. It contains a single species, ''Rodcania kakan'', which lived during the Paleocene. Its remains were found in South America. The genus name is an anagram of '' Carodnia''. Description This animal is only known from scarce fossil remains, and its appearance is therefore only conjectural. From a comparison with its relative '' Carodnia'', it is supposed that ''Rodcania'' was a rather heavy animal with a massive build and strong legs. An adult ''Rodcania'' could weigh around 165 kilograms. ''Rodcania'' is only known from a fragmentary left mandible, bearing the second and third molars. Both teeth are characterized by a thick layer of enamel, vertically oriented Hunter-Schreger bands, and distally inclined dental wear marks. ''Rodcania'' differs from its similar relative ''Carodnia'' and other xenungulates by the simplified and mesiodistally short trigonid of the third molar, the absence of paraconi ...
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Uintatheriidae
Uintatheriidae is a family of extinct ungulate mammals that includes ''Uintatherium''. Uintatheres belong to the order Dinocerata, one of several extinct orders of primitive hoofed mammals that are sometimes united in the Condylarthra. Uintatheres were the largest land animals of their time, surviving from the late Paleocene into the Uintan Epoch of the Middle Eocene. They were heavy animals, with thick legs, massive bones, broad feet, and tiny brains. The most distinctive feature of the great majority of species, however, was the presence of multiple blunt "horns", perhaps similar to the ossicones of modern giraffes, and the presence of large, sabre-like canine teeth. They were eventually replaced as large browsing animals by the even larger brontotheres. Genera Family Uintatheriidae * Subfamily Uintatheriinae ** Genus ''Bathyopsis'' ** Genus ''Eobasileus'' ** Genus ''Prodinoceras'' ** Genus ''Tetheopsis'' ** Genus ''Uintatherium'' * Subfamily Gobiatheriinae ** Genus ''Gobiathe ...
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Arctocyonidae
Arctocyonidae (from Greek ''arktos'' ''kyôn'', "bear/dog-like") has been defined as an extinct family of unspecialized, primitive mammals with more than 20 genera. Animals assigned to this family were most abundant during the Paleocene, but extant from the late Cretaceous to the early Eocene (). Like most early mammals, their actual relationships are very difficult to resolve. No Paleocene fossil has been unambiguously assigned to any living order of placental mammals, and many genera resemble each other: generalized robust, not very agile animals with long tails and all-purpose chewing teeth, living in warm closed-canopy forests with many niches left vacant by the K-T extinction. Arctocyonids were early defined as a family of creodonts (early predators), then reassigned to the condylarths (primitive plant-eaters, now understood as a wastebasket taxon). More recently, these animals have been thought to be the ancestors of the orders Mesonychia and Cetartiodactyla, although some ...
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