''Etayoa'' is an

ungulate Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, cam ...

of the family Carodniidae in the order

Xenungulata Xenungulata ("strange ungulates") is an order of extinct and primitive South American hoofed mammals that lived from the Late Paleocene to Early Eocene (Itaboraian to Casamayoran in the SALMA classification). Fossils of the order are known f ...

that lived during the

Early Eocene In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...

(~ 55 Ma) in northern

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

Etymology

The genus of the type species ''Etayoa bacatensis'' was named by palaeontologist Carlos Villarroel after Fernando Etayo Serna, who contributed extensively to the

paleontology Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

and

stratigraphy Stratigraphy is a branch of geology concerned with the study of rock (geology), rock layers (Stratum, strata) and layering (stratification). It is primarily used in the study of sedimentary rock, sedimentary and layered volcanic rocks. Stratigrap ...

research in Colombia. The species epithet ''bacatensis'' refers to

Bacatá Bacatá is the name given to the main settlement of the Muisca Confederation on the Bogotá savanna. It mostly refers to an area, rather than an individual village, although the name is also found in texts referring to the modern settlement of Fu ...

,Villarroel, 1987 the name in Muysccubun for the main settlement of the southern

Muisca Confederation The Muisca Confederation was a loose confederation of different Muisca rulers (''zaques'', ''zipas'', '' iraca'', and ''tundama'') in the central Andean highlands of present-day Colombia before the Spanish conquest of northern South America. The ...

; the name of which has been used for the current Colombian capital

Bogotá Bogotá (, also , , ), officially Bogotá, Distrito Capital, abbreviated Bogotá, D.C., and formerly known as Santa Fe de Bogotá (; ) during the Spanish period and between 1991 and 2000, is the capital city of Colombia, and one of the larges ...

, founded in a different location than the original Bacatá.

Description

The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

specimen consists of a partial

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...

with teeth, found in the

Bogotá Formation The Bogotá Formation ( es, Formación Bogotá, E1-2b, Tpb, Pgb) is a geological formation of the Eastern Hills and Bogotá savanna on the Altiplano Cundiboyacense, Eastern Ranges of the Colombian Andes. The predominantly shale and siltstone form ...

in the locality

Ciudad Bolívar Ciudad Bolívar (; Spanish for "Bolivar City"), formerly known as Angostura and St. Thomas de Guyana, is the capital of Venezuela's southeastern Bolívar, Venezuela, Bolívar State. It lies at the spot where the Orinoco River narrows to about ...

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

.1987
Description of ''Etayoa bacatensis''
-

Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals, plants, and microorganisms. History The Paleobiology Database (PBDB) originated in the NCEAS-funded Phanerozoic Marine Pale ...

The estimated size of the ungulate is the size of a dog. 2011
Un xenungulado del Paleoceno de la sabana de Bogotá
- Paleontología en Colombia

Paleoclimate and environment

The finds of iguanians, including the fossil record of hoplocercines, and boine, caenophidian, and ungaliophiine snakes in the Bogotá Formation indicate a tropical forest environment, present just before the Early Eocene Climatic Optimum (EECO).Head et al., 2012 The abundant

paleosol In the geosciences, paleosol (''palaeosol'' in Great Britain and Australia) is an ancient soil that formed in the past. The precise definition of the term in geology and paleontology is slightly different from its use in soil science. In geolo ...

s of the Bogotá Formation show an increase in chemical weathering across the Paleocene-Eocene (P-E) transition; the

Paleocene–Eocene Thermal Maximum The Paleocene–Eocene thermal maximum (PETM), alternatively (ETM1), and formerly known as the "Initial Eocene" or "", was a time period with a more than 5–8 °C global average temperature rise across the event. This climate event o ...

.Morón et al., 2013

References

Bibliography

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Etymology

Description

Paleoclimate and environment

References

Bibliography

Further reading