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Vegetative Phase Change
Vegetative phase change is the juvenile-to-adult transition in plants. This transition is distinct from the reproductive transition and is most prolonged and pronounced in woody species. Manipulating phase change may be an important avenue for plant improvement. In the model plant ''Arabidopsis thaliana'', vegetative phase change is relatively subtle: leaves become more curled, with an increased number of abaxial trichomes, and increased serration. Studies in ''A. thaliana'' and maize identified microRNA ''MIR156'' genes as master regulators of phase change, through their regulation of SQUAMOSA-PROMOTER-BINDING-LIKE (SBP/SPL) transcription factors. This gene regulatory circuit appears to be conserved (with variations) in all land plants, including mosses. See also * Plant morphology * Heteroblasty Heteroblasty is the significant and abrupt change in form and function, that occurs over the lifespan of certain plants. Characteristics affected include internode length and stem ...
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Juvenile (organism)
A juvenile is an individual organism (especially an animal) that has not yet reached its adult form, sexual maturity or size. Juveniles can look very different from the adult form, particularly in colour, and may not fill the same niche as the adult form. In many organisms the juvenile has a different name from the adult (see List of animal names). Some organisms reach sexual maturity in a short metamorphosis, such as ecdysis in many insects and some other arthropods. For others, the transition from juvenile to fully mature is a more prolonged process—puberty in humans and other species (like higher primates and whales), for example. In such cases, juveniles during this transformation are sometimes called subadults. Many invertebrates cease development upon reaching adulthood. The stages of such invertebrates are larvae or nymphs. In vertebrates and some invertebrates (e.g. spiders), larval forms (e.g. tadpoles) are usually considered a development stage of their own, an ...
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Adult
An adult is an animal that has reached full growth. The biological definition of the word means an animal reaching sexual maturity and thus capable of reproduction. In the human context, the term ''adult'' has meanings associated with social and legal concepts. In contrast to a non-adult or " minor", a legal adult is a person who has attained the age of majority and is therefore regarded as independent, self-sufficient, and responsible. They may also be regarded as "majors". The typical age of attaining legal adulthood is 18 although definition may vary by legal rights, country, and psychological development. Human adulthood encompasses psychological adult development. Definitions of adulthood are often inconsistent and contradictory; a person may be biologically an adult, and have adult behavior, but still be treated as a child if they are under the legal age of majority. Conversely, one may legally be an adult but possess none of the maturity and responsibility that may de ...
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Plants
Plants are the eukaryotes that form the kingdom Plantae; they are predominantly photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with cyanobacteria to produce sugars from carbon dioxide and water, using the green pigment chlorophyll. Exceptions are parasitic plants that have lost the genes for chlorophyll and photosynthesis, and obtain their energy from other plants or fungi. Most plants are multicellular, except for some green algae. Historically, as in Aristotle's biology, the plant kingdom encompassed all living things that were not animals, and included algae and fungi. Definitions have narrowed since then; current definitions exclude fungi and some of the algae. By the definition used in this article, plants form the clade Viridiplantae (green plants), which consists of the green algae and the embryophytes or land plants ( hornworts, liverworts, mosses, lycophytes, ferns, conifers and other ...
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Flower
Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, modified leaves; corolla, the petals; androecium, the male reproductive unit consisting of stamens and pollen; and gynoecium, the female part, containing style and stigma, which receives the pollen at the tip of the style, and ovary, which contains the ovules. When flowers are arranged in groups, they are known collectively as inflorescences. Floral growth originates at stem tips and is controlled by MADS-box genes. In most plant species flowers are heterosporous, and so can produce sex cells of both sexes. Pollination mediates the transport of pollen to the ovules in the ovaries, to facilitate sexual reproduction. It can occur between different plants, as in cross-pollination, or between flowers on the same plant or even the same f ...
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Woody Plant
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to Herbaceous plant, herbaceous plants that die back to the ground until Spring (season), spring. Characteristics Woody plants are usually trees, shrubs, or lianas. These are usually perennial plants whose stems and larger roots are reinforced with wood produced from secondary xylem. The main stem, larger branches, and roots of these plants are usually covered by a layer of Bark (botany), bark. Wood is a structural cell (biology), tissue that allows woody plants to grow from above ground stems year after year, thus making some woody plants the largest and tallest terrestrial plants. Woody plants, like Herbaceous plant, herbaceous perennials, typically have a Dormancy, dormant period of the year when growth does not take place. This occurs in Temperate climate, temperate and Continental clima ...
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Arabidopsis
''Arabidopsis'' (rockcress) is a genus in the family Brassicaceae. They are small flowering plants related to cabbage and mustard. This genus is of great interest since it contains thale cress (''Arabidopsis thaliana''), one of the model organisms used for studying plant biology and the first plant to have its entire genome sequenced. Changes in thale cress are easily observed, making it a very useful model. Status Currently, the genus ''Arabidopsis'' has nine species and a further eight subspecies recognised. This delimitation is quite recent and is based on morphological and molecular phylogenies by O'Kane and Al-Shehbaz and others. Their findings confirm the species formerly included in ''Arabidopsis'' made it polyphyletic. The most recent reclassification moves two species previously placed in ''Cardaminopsis'' and ''Hylandra'' and three species of '' Arabis'' into ''Arabidopsis'', but excludes 50 that have been moved into the new genera ''Beringia, Crucihimalaya, Ianhedg ...
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Maize
Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native Americans planted it alongside beans and squashes in the Three Sisters polyculture. The leafy stalk of the plant gives rise to male inflorescences or tassels which produce pollen, and female inflorescences called ears. The ears yield grain, known as kernels or seeds. In modern commercial varieties, these are usually yellow or white; other varieties can be of many colors. Maize relies on humans for its propagation. Since the Columbian exchange, it has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat and rice. Much maize is used for animal feed, whether as grain or as the whole plant, which can either be baled or made into the more palatable silage. Sugar-rich varieties called sw ...
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MicroRNA
Micro ribonucleic acid (microRNA, miRNA, μRNA) are small, single-stranded, non-coding RNA molecules containing 21–23 nucleotides. Found in plants, animals, and even some viruses, miRNAs are involved in RNA silencing and post-transcriptional regulation of gene expression. miRNAs base-pair to complementary sequences in messenger RNA (mRNA) molecules, then silence said mRNA molecules by one or more of the following processes: * Cleaving the mRNA strand into two pieces. * Destabilizing the mRNA by shortening its poly(A) tail. * Reducing translation of the mRNA into proteins. In cells of humans and other animals, miRNAs primarily act by destabilizing the mRNA. miRNAs resemble the small interfering RNAs (siRNAs) of the RNA interference (RNAi) pathway, except miRNAs derive from regions of RNA transcripts that fold back on themselves to form short stem-loops (hairpins), whereas siRNAs derive from longer regions of double-stranded RNA. The human genome may encode ov ...
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Squamosa Promoter Binding Protein
The SQUAMOSA promoter binding protein-like (SBP or SPL) family of transcription factors are defined by a plant-specific DNA-binding domain. The founding member of the family was identified based on its specific ''in vitro'' binding to the promoter of the snapdragon SQUAMOSA gene. SBP proteins are thought to be transcriptional activators. Function SPB proteins have roles in leaf development, vegetative phase change, flower and fruit development, plant architecture, sporogenesis, Gibberelic acid signaling and toxin response. Structure The domain contains 10 conserved cysteine and histidine residues that probably are zinc ligands. The SBP domain is a highly conserved DNA-binding domain. It is approximately 80 amino acids in length and contains a zinc finger motif with two zinc-binding sites: Cys-Cys-His-Cys and Cys-Cys-Cys-His. It has a three-stranded antiparallel beta-sheet The beta sheet (β-sheet, also β-pleated sheet) is a common structural motif, motif of the re ...
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Transcription Factor
In molecular biology, a transcription factor (TF) (or sequence-specific DNA-binding factor) is a protein that controls the rate of transcription (genetics), transcription of genetics, genetic information from DNA to messenger RNA, by binding to a specific DNA sequence. The function of TFs is to regulate—turn on and off—genes in order to make sure that they are Gene expression, expressed in the desired Cell (biology), cells at the right time and in the right amount throughout the life of the cell and the organism. Groups of TFs function in a coordinated fashion to direct cell division, cell growth, and cell death throughout life; cell migration and organization (body plan) during embryonic development; and intermittently in response to signals from outside the cell, such as a hormone. There are approximately 1600 TFs in the human genome. Transcription factors are members of the proteome as well as regulome. TFs work alone or with other proteins in a complex, by promoting (a ...
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Gene Regulatory Circuit
Genetic regulatory circuits (also referred to as transcriptional regulatory circuits) is a concept that evolved from the Operon Model discovered by François Jacob and Jacques Monod.  They are functional clusters of genes that impact each other's expression through inducible transcription factors and cis-regulatory elements. Genetic regulatory circuits are analogous in many ways to electronic circuits in how they use signal inputs and outputs to determine Regulation of gene expression, gene regulation. Like electronic circuits, their organization determines their efficiency, and this has been demonstrated in circuits working in series to have a greater sensitivity of gene regulation. They also use inputs such as trans and cis sequence regulators of genes, and outputs such as gene expression level. Depending on the type of circuit, they respond constantly to outside signals, such as sugars and hormone levels, that determine how the circuit will return to its fixed point or periodic ...
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Embryophyte
The embryophytes () are a clade of plants, also known as Embryophyta (Plantae ''sensu strictissimo'') () or land plants. They are the most familiar group of photoautotrophs that make up the vegetation on Earth's dry lands and wetlands. Embryophytes have a common ancestor with green algae, having emerged within the Phragmoplastophyta clade of freshwater charophyte green algae as a sister taxon of Charophyceae, Coleochaetophyceae and Zygnematophyceae. Embryophytes consist of the bryophytes and the polysporangiophytes. Living embryophytes include hornworts, liverworts, mosses, lycophytes, ferns, gymnosperms and angiosperms (flowering plants). Embryophytes have diplobiontic life cycles. The embryophytes are informally called "land plants" because they thrive primarily in terrestrial habitats (despite some members having evolved secondarily to live once again in semiaquatic/ aquatic habitats), while the related green algae are primarily aquatic. Embryophytes are complex ...
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