Ulmus × Intermedia 'Coolshade'
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Ulmus × Intermedia 'Coolshade'
The hybrid elm cultivar ''Ulmus × intermedia'' 'Coolshade' is an American hybrid cultivar cloned from a crossing of the Slippery, or Red, Elm ''Ulmus rubra'' (female parent) and the Siberian Elm ''Ulmus pumila'' at the Sarcoxie Nurseries, Sarcoxie, Missouri, in 1946. At Arnold Arboretum, where there was a specimen, herbarium material was labelled ''Ulmus pumila'' 'Coolshade'. Description 'Coolshade' has rapid, stocky growth with a compact crown resistant to breakage under ice and snow. Its foliage is a very dark green.Bean, W. J. (1981). ''Trees and shrubs hardy in Great Britain'', 7th edition. Murray, London. There appears to be little evidence of ''Ulmus rubra'' ingression in the Arnold Arboretum 1960 leaves specimens labelled ''Ulmus pumila'' 'Coolshade', suggesting that its hybridity may have been questioned there. Pests and diseases Reputedly tolerant of Dutch elm disease Dutch elm disease (DED) is caused by a member of the sac fungi (Ascomycota) affecting elm trees, ...
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Ulmus × Intermedia
''Ulmus × intermedia'' Elowsky is a natural hybrid elm occurring across Nebraska and several other Midwestern states, derived from the crossing of ''Ulmus rubra'' and ''Ulmus pumila''. As Red Elm ''U. rubra'' is far less fertile, and highly susceptible to Dutch elm disease (:DED), it could eventually be hybridized out of existence by ''U. × intermedia''. Schlautman, B., Zalapa, J., & Brunet, J. (2012). Genotypic and phenotypic characterization of Ulmus pumila (Siberian elm), Ulmus rubra (Red elm) and their hybrids. ''Proc. Nebraska Acad. Sci. 2012''. 106–107. The hybrid was first reported from the wild in the Chicago region in 1950 and was provisionally named ''U.'' × ''notha'' Wilhelm & Ware in 1994. The horticulture industry made a number of artificial crosses of the two species, such as 'Coolshade' and 'Fremont', in an attempt to create ornamental trees resistant to DED. Description ''Ulmus × intermedia'' is significantly different from both parental species, exc ...
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Ulmus Rubra
''Ulmus rubra'', the slippery elm, is a species of elm native to eastern North America. Other common names include red elm, gray elm, soft elm, moose elm, and Indian elm. Description ''Ulmus rubra'' is a medium-sized deciduous tree with a spreading head of branches,Hillier & Sons. (1990). ''Hillier's Manual of Trees & Shrubs, 5th ed.''. David & Charles, Newton Abbot, UK commonly growing to , very occasionally over in height. Its heartwood is reddish-brown. The broad oblong to obovate leaves are long, rough above but velvety below, with coarse double-serrate margins, acuminate apices and oblique bases; the petioles are long.Bean, W. J. (1970). ''Trees & Shrubs Hardy in the British Isles'', 8th ed., p. 656. (2nd impression 1976) John Murray, London. The leaves are often tinged red on emergence, turning dark green by summer and a dull yellow in autumn. The perfect, apetalous, wind-pollinated flowers are produced before the leaves in early spring, usually in tight, short ...
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Ulmus Pumila
''Ulmus pumila'', the Siberian elm, is a tree native to Asia. It is also known as the Asiatic elm and dwarf elm, but sometimes miscalled the 'Chinese Elm' ('' Ulmus parvifolia''). ''U. pumila'' has been widely cultivated throughout Asia, North America, Argentina, and southern Europe, becoming naturalized in many places, notably across much of the United States. Description The Siberian elm is usually a small to medium-sized, often bushy, deciduous tree growing to tall, the diameter at breast height to . The bark is dark gray, irregularly longitudinally fissured. The branchlets are yellowish gray, glabrous or pubescent, unwinged and without a corky layer, with scattered lenticels. The winter buds dark brown to red-brown, globose to ovoid. The petiole is , pubescent, the leaf blade elliptic-ovate to elliptic-lanceolate, , the colour changing from dark green to yellow in autumn.Fu, L., Xin, Y. & Whittemore, A. (2002). Ulmaceae, in Wu, Z. & Raven, P. (eds) ''Flora of Chin ...
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Hybrid (biology)
In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in blending inheritance), but can show hybrid vigor, sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in the individual parentage. In genetics, attention is focused on the numbers of chromosomes. In taxonomy, a key question is how closely related the parent species are. Species are reproductively isolated by strong barriers to hybridisation, which include genetic and morphological differences, differing times of fertility, mating behaviors and cues, and physiological rejection of sperm cells or the developing embryo. Some act before fertilization and others after it. Similar barriers exist in plants, with differences in flowering t ...
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Cultivar
A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, or carefully controlled seed production. Most cultivars arise from purposeful human manipulation, but some originate from wild plants that have distinctive characteristics. Cultivar names are chosen according to rules of the International Code of Nomenclature for Cultivated Plants (ICNCP), and not all cultivated plants qualify as cultivars. Horticulturists generally believe the word ''cultivar''''Cultivar'' () has two meanings, as explained in '' Formal definition'': it is a classification category and a taxonomic unit within the category. When referring to a taxon, the word does not apply to an individual plant but to all plants that share the unique characteristics that define the cultivar. was coined as a term meaning "cultivated var ...
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Hybrid (biology)
In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in blending inheritance), but can show hybrid vigor, sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in the individual parentage. In genetics, attention is focused on the numbers of chromosomes. In taxonomy, a key question is how closely related the parent species are. Species are reproductively isolated by strong barriers to hybridisation, which include genetic and morphological differences, differing times of fertility, mating behaviors and cues, and physiological rejection of sperm cells or the developing embryo. Some act before fertilization and others after it. Similar barriers exist in plants, with differences in flowering t ...
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Sarcoxie
Sarcoxie is a city in Jasper County, Missouri, United States. The population was 1,406 at the 2020 census. It is part of the Joplin, Missouri Metropolitan Statistical Area. History Sarcoxie was platted in the early 1830s, and it was originally called Centerville from its location upon Center Creek. In 1839, the settlement was renamed in honor of Sarcoxie, a chief of the Delaware Indians who had settled near a spring in the present town limits. Sarcoxie was once the strawberry capital of the world and still is the peony capital of the world, and home to Gilbert H. Wild, one of America's largest growers of daylilies, iris, and peonies. Sarcoxie once had its own currency that had a picture of a strawberry on one side. During the start of the Civil War in 1861, Franz Sigel's independent command passed through Sarcoxie on June 28 before attempting to attack the separated commands of Sterling Price and Missouri Governor and General Claiborne Fox Jackson. Price was rumored to be at ...
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Missouri
Missouri is a state in the Midwestern region of the United States. Ranking 21st in land area, it is bordered by eight states (tied for the most with Tennessee): Iowa to the north, Illinois, Kentucky and Tennessee to the east, Arkansas to the south and Oklahoma, Kansas and Nebraska to the west. In the south are the Ozarks, a forested highland, providing timber, minerals, and recreation. The Missouri River, after which the state is named, flows through the center into the Mississippi River, which makes up the eastern border. With more than six million residents, it is the 19th-most populous state of the country. The largest urban areas are St. Louis, Kansas City, Springfield and Columbia; the capital is Jefferson City. Humans have inhabited what is now Missouri for at least 12,000 years. The Mississippian culture, which emerged at least in the ninth century, built cities and mounds before declining in the 14th century. When European explorers arrived in the 17th ...
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Arnold Arboretum
The Arnold Arboretum of Harvard University is a botanical research institution and free public park, located in the Jamaica Plain and Roslindale neighborhoods of Boston, Massachusetts. Established in 1872, it is the oldest public arboretum in North America. The landscape was designed by Charles Sprague Sargent and Frederick Law Olmsted and is the second largest "link" in the Emerald Necklace. The Arnold Arboretum's collection of temperate trees, shrubs, and vines has a particular emphasis on the plants of the eastern United States and eastern Asia, where arboretum staff and colleagues are actively sourcing new material on plant collecting expeditions. The arboretum supports research in its landscape and in its Weld Hill Research Building. History The Arboretum was founded in 1872 when the President and Fellows of Harvard College became trustees of a portion of the estate of James Arnold (1781–1868), a whaling merchant from New Bedford, Massachusetts. Arnold specified th ...
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Harvard University
Harvard University is a private Ivy League research university in Cambridge, Massachusetts. Founded in 1636 as Harvard College and named for its first benefactor, the Puritan clergyman John Harvard, it is the oldest institution of higher learning in the United States and one of the most prestigious and highly ranked universities in the world. The university is composed of ten academic faculties plus Harvard Radcliffe Institute. The Faculty of Arts and Sciences offers study in a wide range of undergraduate and graduate academic disciplines, and other faculties offer only graduate degrees, including professional degrees. Harvard has three main campuses: the Cambridge campus centered on Harvard Yard; an adjoining campus immediately across Charles River in the Allston neighborhood of Boston; and the medical campus in Boston's Longwood Medical Area. Harvard's endowment is valued at $50.9 billion, making it the wealthiest academic institution in the world. Endow ...
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Dutch Elm Disease
Dutch elm disease (DED) is caused by a member of the sac fungi (Ascomycota) affecting elm trees, and is spread by elm bark beetles. Although believed to be originally native to Asia, the disease was accidentally introduced into Americas, America, Europe, and New Zealand. In these regions it has devastated native populations of elms that did not have resistance to the disease. The name "Dutch elm disease" refers to its identification in 1921 and later in the Netherlands by Dutch phytopathologists Marie Beatrice Schol-Schwarz, Bea Schwarz and Christine Buisman, who both worked with professor Johanna Westerdijk. The disease affects species in the genera ''Ulmus'' and ''Zelkova''; therefore it is not specific to the Ulmus × hollandica, Dutch elm hybrid. Overview Dutch elm disease (DED) is caused by ascomycete microfungi.
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Ulmus 'Improved Coolshade'
Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North America and Eurasia, presently ranging southward in the Middle East to Lebanon and Israel,Flora of Israel OnlineUlmus minor Mill. , Flora of Israel Online accessdate: July 28, 2020 and across the Equator in the Far East into Indonesia.Fu, L., Xin, Y. & Whittemore, A. (2002). Ulmaceae, in Wu, Z. & Raven, P. (eds) Flora of China'', Vol. 5 (Ulmaceae through Basellaceae). Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis, US. Elms are components of many kinds of natural forests. Moreover, during the 19th and early 20th centuries, many species and cultivars were also planted as ornamental street, garden, and park trees in Europe, North America, and parts of the Southern Hemisphere, notably Australasia. Some individual ...
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